2212 lines
168 KiB
XML
2212 lines
168 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1075.67459" ID-GBIF-Dataset="e83d47ed-a6b8-4706-9df8-e1686d1be39f" ID-PMC="PMC8671714" ID-Pensoft-Pub="1313-2970-1075-33" ID-Pensoft-UUID="DBA502BCE85F5F3C922FDD3123C6365B" ID-PubMed="35046751" ID-ZooBank="BB034E5C10CB49179FCDEF6A72816F6C" ModsDocID="1313-2970-1075-33" checkinTime="1638937131209" checkinUser="pensoft" docAuthor="Moreno-Gonzalez 1, Jairo A., Pinto-da-Rocha 1, Ricardo & Gallao 2, Jonas E." docDate="2021" docId="05B3683E83175DC39567AA5195E1784D" docLanguage="en" docName="ZooKeys 1075: 33-75" docOrigin="ZooKeys 1075" docPubDate="2021-12-07" docSource="http://dx.doi.org/10.3897/zookeys.1075.67459" docTitle="Tityus spelaeus Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2 2021, sp. nov." docType="treatment" docUuid="3AE5D4E6-C2F1-47A7-9768-046B09B2FF48" docUuidSource="ZooBank" docVersion="5" id="DBA502BCE85F5F3C922FDD3123C6365B" lastPageNumber="33" masterDocId="DBA502BCE85F5F3C922FDD3123C6365B" masterDocTitle="Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species" masterLastPageNumber="75" masterPageNumber="33" pageNumber="33" updateTime="1668151205184" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
|
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<mods:title>Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Moreno-Gonzalez 1, Jairo A.</mods:namePart>
|
||
<mods:affiliation>Instituto de Biociencias - Universidade de Sao Paulo, Departamento de Zoologia. Rua do Matao, travessa 14, 321, 005508 - 900, Sao Paulo, Brazil</mods:affiliation>
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||
</mods:name>
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||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Pinto-da-Rocha 1, Ricardo</mods:namePart>
|
||
<mods:affiliation>Instituto de Biociencias - Universidade de Sao Paulo, Departamento de Zoologia. Rua do Matao, travessa 14, 321, 005508 - 900, Sao Paulo, Brazil</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Gallao 2, Jonas E.</mods:namePart>
|
||
<mods:affiliation>Instituto de Biociencias - Universidade de Sao Paulo, Departamento de Zoologia. Rua do Matao, travessa 14, 321, 005508 - 900, Sao Paulo, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
|
||
<mods:date>2021</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2021-12-07</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>1075</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>33</mods:start>
|
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<mods:end>75</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1075.67459</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1075.67459</mods:identifier>
|
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<mods:identifier type="Pensoft-Pub">1313-2970-1075-33</mods:identifier>
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<mods:identifier type="ZooBank">BB034E5C10CB49179FCDEF6A72816F6C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">DBA502BCE85F5F3C922FDD3123C6365B</mods:identifier>
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</mods:mods>
|
||
<subSection pageId="0" pageNumber="33" type="results">
|
||
<treatment ID-GBIF-Taxon="191254655" LSID="urn:lsid:zoobank.org:act:3AE5D4E6-C2F1-47A7-9768-046B09B2FF48" httpUri="http://treatment.plazi.org/id/05B3683E83175DC39567AA5195E1784D" lastPageNumber="33" pageId="0" pageNumber="33">
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<subSubSection pageId="0" pageNumber="33" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<taxonomicName LSID="http://zoobank.org/3AE5D4E6-C2F1-47A7-9768-046B09B2FF48" authority="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2, 2021" authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus" status="sp. nov.">Tityus spelaeus</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="33">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="description">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, and Cytochrome c Oxidase I- COI). Maximum likelihood tree (Log-likelihood = - 12896.086), showing species-groups and subgenera. Values on nodes correspond to ultrafast-bootstrap (Ubst) values." figureDoi="10.3897/zookeys.1075.67459.figure1" httpUri="https://binary.pensoft.net/fig/619807" pageId="0" pageNumber="33">Figures 1</figureCitation>
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, and Cytochrome c Oxidase I- COI), showing the distribution of the characters states of the ventral setae of telotarsi I-IV (orange: an irregularly distributed tuft of setae (type I); turquoise: two ventro-submedian rows of setae (type II )) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold." figureDoi="10.3897/zookeys.1075.67459.figure2" httpUri="https://binary.pensoft.net/fig/619808" pageId="0" pageNumber="33">, 2</figureCitation>
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, Cytochrome c Oxidase I), showing distribution of the characters states exhibited by the female basal pectinal piece (orange: GR absent; grey: medium-sized GR; turquoise: relatively large GR; blue: very large GR) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold. Abbreviations = GR, glandular region." figureDoi="10.3897/zookeys.1075.67459.figure3" httpUri="https://binary.pensoft.net/fig/619809" pageId="0" pageNumber="33">, 3</figureCitation>
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tityus C. L. Koch, 1836, telotarsi IV, showing ventral macrosetae A, C, E, G, I, K white light B, D, F, H, J, L UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (type II) (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (type I) (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (type II) (MZSP) G, H Tityus (Atreus) obscurus Gervais, 1843 (type I) (MNRJ 07610) I, J Tityus (Tityus) serrulatus Lutz & Mello, 1922 (type II) (MZSP 28205) K, L Tityus (Tityus) spelaeus sp. nov. (MZSP 74633) (type II). Observations = telotarsi I-IV ventral setae distribution: Type I = tuft of irregularly distributed setae. Type II = two discrete ventrosubmedian rows of setae. Scale bars: 500 μm." figureDoi="10.3897/zookeys.1075.67459.figure4" httpUri="https://binary.pensoft.net/fig/619810" pageId="0" pageNumber="33">, 4</figureCitation>
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (MZSP). Abbreviations: BML, basal middle lamellae; GR, glandular region. Scale bars: 500 μm." figureDoi="10.3897/zookeys.1075.67459.figure5" httpUri="https://binary.pensoft.net/fig/619811" pageId="0" pageNumber="33">, 5</figureCitation>
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Atreus) obscurus Gervais, 1843 (MNRJ 07610) C, D Tityus (Tityus) serrulatus Lutz & Mello, 1922 (MZSP 28205) E, F Tityus (Tityus) spelaeus sp. nov. (MZSP 74633). Abbreviations: BML, basal middle lamellae; GR, glandular region of the pectinal basal piece. Scale bars: 500 μm." figureDoi="10.3897/zookeys.1075.67459.figure6" httpUri="https://binary.pensoft.net/fig/619812" pageId="0" pageNumber="33">, 6</figureCitation>
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33">, 7</figureCitation>
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), carapace, dorsal view A white light B UV light. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1075.67459.figure8" httpUri="https://binary.pensoft.net/fig/619814" pageId="0" pageNumber="33">, 8</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="type material">
|
||
<paragraph pageId="0" pageNumber="33">Type material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
Brazil: State of
|
||
<normalizedToken originalValue="Goiás">Goias</normalizedToken>
|
||
:
|
||
<emphasis bold="true" pageId="0" pageNumber="33">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Holotype</emphasis>
|
||
.
|
||
</emphasis>
|
||
Adult female from Posse,
|
||
<normalizedToken originalValue="Russão">Russao</normalizedToken>
|
||
II cave,
|
||
<geoCoordinate degrees="14" direction="south" minutes="05" orientation="latitude" precision="1" seconds="05.3" value="-14.0848055">14°05'05.3"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="46" direction="west" minutes="23" orientation="longitude" precision="1" seconds="07.1" value="-46.385307">46°23'07.1"W</geoCoordinate>
|
||
, 01.iv.2007, R. Pinto-da-Rocha leg. (MZSP 74633).
|
||
<emphasis bold="true" pageId="0" pageNumber="33">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Paratypes</emphasis>
|
||
.
|
||
</emphasis>
|
||
Four adult female paratypes, same data as the holotype (MZSP 74634); eight adult females, same locality as the holotype, 23.iv.2015, J. E.
|
||
<normalizedToken originalValue="Gallão">Gallao</normalizedToken>
|
||
& C. C. de Paula leg. (LES/UFSCar 14668; LES/UFSCar 14669; LES/UFSCar 14670; LES/UFSCar 14671; LES/UFSCar 14672; LES/UFSCar 14673); four adult females, same locality as the holotype, 01.iv.2007, R. Pinto-da-Rocha et al. (MZSP 52228, 52229, 52230, 52231).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="etymology">
|
||
<paragraph pageId="0" pageNumber="33">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
The species epithet is a derivative form of the Greek noun,
|
||
<normalizedToken originalValue="σπήλαιον">σπήλαιον</normalizedToken>
|
||
(Latin: caverna), which means cave, in reference to the subterranean habitat where
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus spelaeus</emphasis>
|
||
</taxonomicName>
|
||
has an established population. It is a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="33">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
(Based on female). This species belongs to the
|
||
<taxonomicName authorityName="Kraepelin" authorityYear="1898" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus trivittatus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="trivittatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus trivittatus</emphasis>
|
||
</taxonomicName>
|
||
species-group (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, and Cytochrome c Oxidase I- COI), showing the distribution of the characters states of the ventral setae of telotarsi I-IV (orange: an irregularly distributed tuft of setae (type I); turquoise: two ventro-submedian rows of setae (type II )) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold." figureDoi="10.3897/zookeys.1075.67459.figure2" httpUri="https://binary.pensoft.net/fig/619808" pageId="0" pageNumber="33">2</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, Cytochrome c Oxidase I), showing distribution of the characters states exhibited by the female basal pectinal piece (orange: GR absent; grey: medium-sized GR; turquoise: relatively large GR; blue: very large GR) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold. Abbreviations = GR, glandular region." figureDoi="10.3897/zookeys.1075.67459.figure3" httpUri="https://binary.pensoft.net/fig/619809" pageId="0" pageNumber="33">3</figureCitation>
|
||
). Among members of the group distributed in Brazil (
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. carvalhoi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="carvalhoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. carvalhoi</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
|
||
, 1945;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. charreyroni" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="charreyroni">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. charreyroni</emphasis>
|
||
</taxonomicName>
|
||
Vellard, 1932;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. confluens" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="confluens">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. confluens</emphasis>
|
||
</taxonomicName>
|
||
Borelli, 1899;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. fasciolatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
Pessoa, 1935;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. jeanvellardi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="jeanvellardi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. jeanvellardi</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
|
||
, 2001;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. karaja" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="karaja">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. karaja</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
|
||
, 2016;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. rupestre" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="rupestre">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. rupestre</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
|
||
, 2019;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. sylviae" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="sylviae">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. sylviae</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
|
||
, 2005, and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. trivittatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="trivittatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. trivittatus</emphasis>
|
||
</taxonomicName>
|
||
Kraepelin, 1898),
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus spelaeus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be readily recognized.
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus spelaeus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. carvalhoi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="carvalhoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. carvalhoi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. charreyroni" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="charreyroni">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. charreyroni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. confluens" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="confluens">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. confluens</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. fasciolatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. rupestre" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="rupestre">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. rupestre</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. trivittatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="trivittatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. trivittatus</emphasis>
|
||
</taxonomicName>
|
||
share a subaculear tubercle small, and acute, pointing towards the tip of the aculeus (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33">12A</figureCitation>
|
||
). In contrast,
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. jeanvellardi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="jeanvellardi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. jeanvellardi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. karaja" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="karaja">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. karaja</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. sylviae" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="sylviae">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. sylviae</emphasis>
|
||
</taxonomicName>
|
||
exhibit a small and coarse subaculear tubercle that points either towards the tip of the aculeus (
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. sylviae" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="sylviae">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. sylviae</emphasis>
|
||
</taxonomicName>
|
||
) or towards the middle of the aculeus (
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. jeanvellardi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="jeanvellardi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. jeanvellardi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. karaja" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="karaja">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. karaja</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
On the other hand,
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus spelaeus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. sylviae" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="sylviae">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. sylviae</emphasis>
|
||
</taxonomicName>
|
||
share a very large glandular region occupying beyond the anterior two thirds of the medial region of the pectinal basal piece of female pectines (Figs
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Atreus) obscurus Gervais, 1843 (MNRJ 07610) C, D Tityus (Tityus) serrulatus Lutz & Mello, 1922 (MZSP 28205) E, F Tityus (Tityus) spelaeus sp. nov. (MZSP 74633). Abbreviations: BML, basal middle lamellae; GR, glandular region of the pectinal basal piece. Scale bars: 500 μm." figureDoi="10.3897/zookeys.1075.67459.figure6" httpUri="https://binary.pensoft.net/fig/619812" pageId="0" pageNumber="33">6E, F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tityus (Tityus) spelaeus sp. nov., female paratype (MZSP 74633) genital area and pectines A genital operculum and pectinal basal piece B closeup of the pectinal basal piece, showing cuticular pores on glandular region C right pectine D peg sensillae, distribution E closeup of a peg sensilla. Scale bars: 800 µm (A); 60 µm (B); 1000 µm (C); 20 µm (D); 3 µm (E)." figureDoi="10.3897/zookeys.1075.67459.figure11" httpUri="https://binary.pensoft.net/fig/619817" pageId="0" pageNumber="33">11</figureCitation>
|
||
). In
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. carvalhoi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="carvalhoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. carvalhoi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. charreyroni" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="charreyroni">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. charreyroni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. confluens" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="confluens">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. confluens</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. fasciolatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. trivittatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="trivittatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. trivittatus</emphasis>
|
||
</taxonomicName>
|
||
(females of
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. jeanvellardi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="jeanvellardi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. jeanvellardi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. karaja" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="karaja">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. karaja</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. rupestre" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="rupestre">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. rupestre</emphasis>
|
||
</taxonomicName>
|
||
are unknow) the glandular region occupies a large area of anterior two thirds of the anteriomedian region of the pectinal basal piece of female pectines (e.g., Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (MZSP). Abbreviations: BML, basal middle lamellae; GR, glandular region. Scale bars: 500 μm." figureDoi="10.3897/zookeys.1075.67459.figure5" httpUri="https://binary.pensoft.net/fig/619811" pageId="0" pageNumber="33">5A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
Finally,
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus spelaeus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. karaja" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="karaja">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. karaja</emphasis>
|
||
</taxonomicName>
|
||
can be readily distinguished from
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. carvalhoi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="carvalhoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. carvalhoi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. charreyroni" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="charreyroni">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. charreyroni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. fasciolatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. jeanvellardi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="jeanvellardi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. jeanvellardi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. rupestre" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="rupestre">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. rupestre</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. sylviae" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="sylviae">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. sylviae</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. trivittatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="trivittatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. trivittatus</emphasis>
|
||
</taxonomicName>
|
||
, based on the presence of residual spots on tergites (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33">7</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tityus (Tityus) spelaeus sp. nov., female paratypes under laboratory conditions A, B female paratype with scorpionlings A feeding upon scorpionlings B litter on female's back C specimens feeding on a cricket." figureDoi="10.3897/zookeys.1075.67459.figure13" httpUri="https://binary.pensoft.net/fig/619819" pageId="0" pageNumber="33">13</figureCitation>
|
||
), and having the carapace (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33">7</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), carapace, dorsal view A white light B UV light. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1075.67459.figure8" httpUri="https://binary.pensoft.net/fig/619814" pageId="0" pageNumber="33">8A</figureCitation>
|
||
) and chericeral manus immaculate (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), carapace, dorsal view A white light B UV light. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1075.67459.figure8" httpUri="https://binary.pensoft.net/fig/619814" pageId="0" pageNumber="33">8A</figureCitation>
|
||
). In contrast,
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. charreyroni" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="charreyroni">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. charreyroni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. confluens" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="confluens">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. confluens</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. fasciolatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. jeanvellardi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="jeanvellardi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. jeanvellardi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. rupestre" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="rupestre">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. rupestre</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. sylviae" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="sylviae">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. sylviae</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. trivittatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="trivittatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. trivittatus</emphasis>
|
||
</taxonomicName>
|
||
have the carapace and tergites moderately covered with brownish spots and the cheliceral manus with reticulations (except
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. jeanvellardi" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="jeanvellardi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. jeanvellardi</emphasis>
|
||
</taxonomicName>
|
||
that exhibit a cheliceral manus immaculate).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 7.</emphasis>
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus) spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus" subGenus="Tityus">Tityus (Tityus) spelaeus</taxonomicName>
|
||
sp. nov., female holotype (MZSP 74633)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
ventral view. Scale bars: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="remarks">
|
||
<paragraph pageId="0" pageNumber="33">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
In an unpublished comprehensive phylogenetic analysis of
|
||
<taxonomicName authorityName="C. L. Koch" authorityYear="1836" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Moreno-Gonzalez, JA" journalOrPublisher="Universidade de Sao Paulo, Sao Paulo" pageId="0" pageNumber="33" refId="B50" refString="Moreno-Gonzalez, JA, 2021. Phylogenetic analysis of the Tityus clathratus species-group and other species-groups and subgenera of Tityus (Scorpiones: Buthidae) based on molecular and morphological characters [In portuguese: Analise filogenetica do grupo de especies Tityus clathratus e outros grupos e subgeneros de Tityus (Scorpiones: Buthidae) baseada em caracteres moleculares e morfologicos]. Ph.D. dissertation. Universidade de Sao Paulo, Sao Paulo" title="Phylogenetic analysis of the Tityus clathratus species-group and other species-groups and subgenera of Tityus (Scorpiones: Buthidae) based on molecular and morphological characters [In portuguese: Analise filogenetica do grupo de especies Tityus clathratus e outros grupos e subgeneros de Tityus (Scorpiones: Buthidae) baseada em caracteres moleculares e morfologicos]. Ph. D. dissertation." year="2021">
|
||
<normalizedToken originalValue="Moreno-González">Moreno-Gonzalez</normalizedToken>
|
||
2021
|
||
</bibRefCitation>
|
||
), the
|
||
<taxonomicName authorityName="Kraepelin" authorityYear="1898" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus trivittatus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="trivittatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus trivittatus</emphasis>
|
||
</taxonomicName>
|
||
species-group was one of the most morphologically homogeneous species-groups of the genus. In fact, no somatic character of the morphological matrix (~164 chars) was optimized as a synapomorphy in the nodes within the clade representing the
|
||
<taxonomicName authorityName="Kraepelin" authorityYear="1898" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus trivittatus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="trivittatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus trivittatus</emphasis>
|
||
</taxonomicName>
|
||
species-group. Instead, those nodes were solely supported by unambiguous molecular synapomorphies. It is worth mentioning that, although coloration patterns presented high levels of homoplasy, they also showed significant differences at the species level, and the diagnosis of
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus spelaeus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is based on this background knowledge. Nonetheless, additional studies including molecular and phenotypical evidence of poorly described species from the Central region of Brazil are required to untangle the phylogeny of this cryptic species complex.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1075.67459.figure8" httpUri="https://binary.pensoft.net/fig/619814" pageId="0" pageNumber="33" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 8.</emphasis>
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus) spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus" subGenus="Tityus">Tityus (Tityus) spelaeus</taxonomicName>
|
||
sp. nov., female holotype (MZSP 74633), carapace, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
white light
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
UV light. Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
On the other hand,
|
||
<taxonomicName authorityName="Lourenco" authorityYear="2016" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus karaja" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="karaja">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus karaja</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
|
||
, 2016 was described based on a single male collected in 1929 in the region that corresponds to the northern portion of the state of
|
||
<normalizedToken originalValue="Goiás">Goias</normalizedToken>
|
||
, Brazil. According to the brief description of
|
||
<bibRefCitation author="Lourenco, WR" journalOrPublisher="Journal of Venomous Animals and Toxins Including Tropical Diseases" pageId="0" pageNumber="33" refId="B37" refString="Lourenco, WR, 2016. Une nouvelle espece de Tityus CL Koch, 1836 (Scorpiones: Buthidae), collectee par Jean A. Vellard dans l'ancien Etat de Goias, aujourd'hui Tocantins, Bresil. Revista Iberica de Aracnologia: 75-78." title="Une nouvelle espece de Tityus CL Koch, 1836 (Scorpiones: Buthidae), collectee par Jean A. Vellard dans l'ancien Etat de Goias, aujourd'hui Tocantins, Bresil. Revista Iberica de Aracnologia: 75 - 78." year="2016">
|
||
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
|
||
(2016)
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName authorityName="Lourenco" authorityYear="2016" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus karaja" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="karaja">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus karaja</emphasis>
|
||
</taxonomicName>
|
||
could share a similar body coloration pattern to that of
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. spelaeus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. spelaeus</emphasis>
|
||
</taxonomicName>
|
||
. However, given that the male of
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. karaja" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="karaja">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. karaja</emphasis>
|
||
</taxonomicName>
|
||
is almost a hundred years old, the coloration needs to be corroborated with fresh specimens. Despite this, according to
|
||
<bibRefCitation author="Lourenco, WR" journalOrPublisher="Journal of Venomous Animals and Toxins Including Tropical Diseases" pageId="0" pageNumber="33" refId="B37" refString="Lourenco, WR, 2016. Une nouvelle espece de Tityus CL Koch, 1836 (Scorpiones: Buthidae), collectee par Jean A. Vellard dans l'ancien Etat de Goias, aujourd'hui Tocantins, Bresil. Revista Iberica de Aracnologia: 75-78." title="Une nouvelle espece de Tityus CL Koch, 1836 (Scorpiones: Buthidae), collectee par Jean A. Vellard dans l'ancien Etat de Goias, aujourd'hui Tocantins, Bresil. Revista Iberica de Aracnologia: 75 - 78." year="2016">
|
||
<normalizedToken originalValue="Lourenço’s">Lourenco's</normalizedToken>
|
||
(2016
|
||
</bibRefCitation>
|
||
: fig. 5) illustration, the subaculear tubercle of
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. karaja" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="karaja">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. karaja</emphasis>
|
||
</taxonomicName>
|
||
is conical, small, and coarse, pointing towards the middle of the aculeus, whereas in
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. spelaeus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">T. spelaeus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. it points towards the tip of the aculeus (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33">12A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="description">
|
||
<paragraph pageId="0" pageNumber="33">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">Based on the female holotype (MZSP 74633). Male unknown.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
Total length. Female: 53.52 mm (measurements in Table
|
||
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Measurements (mm) of Tityus spelaeus sp. nov." httpUri="http://table.plazi.org/id/6304231CAF3B5B7548B43B657862B777" pageId="0" pageNumber="33" tableUuid="6304231CAF3B5B7548B43B657862B777">4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-Table-UUID="6304231CAF3B5B7548B43B657862B777" httpUri="http://table.plazi.org/id/6304231CAF3B5B7548B43B657862B777" pageId="0" pageNumber="33" start="Table 4" startId="T4">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Table 4.</emphasis>
|
||
Measurements (mm) of
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus spelaeus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<table pageId="0" pageNumber="33">
|
||
<tr pageId="0" pageNumber="33">
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="2">Structure</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="2">Measure</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female holotype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype # 1</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype # 2</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype #4</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype #5</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Female paratype</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">MZSP 74633</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">MZSP 74633</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">MZSP 74633</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">MZSP 74633</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">MZSP 74633</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">MZSP 52228</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">MZSP 52229</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">MZSP 52230</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">MZSP 52231</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">LES014668</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">LES014669</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">LES014670</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">LES014671</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">LES014672</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">LES014673</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">LES014673</th>
|
||
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">LES014673</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Total length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">53.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">51.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">57.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">51.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">51.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">48.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">49.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">51.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">49.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">57.98</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">53.69</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">53.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">54.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">54.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">50.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">52.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">54.29</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Carapace</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.83</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.98</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.73</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.07</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Carapace</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">anterior width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.10</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Carapace</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">posterior width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.92</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.21</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Carapace</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">eye diameter</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.43</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Carapace</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">interocular distance</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.59</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Carapace</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">ocular diada width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.19</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Tergite I</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">0.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.17</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Tergite II</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.41</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Tergite III</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.73</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.76</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Tergite IV</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.94</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.06</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Tergite V</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.38</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Tergite VI</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.56</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Tergite VII</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.96</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.83</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.28</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Mesosoma</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">total length (tergites)</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">15.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">14.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">16.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">14.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">14.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">13.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">14.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">14.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">14.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">16.83</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">15.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">15.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">15.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">15.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">14.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">14.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">15.62</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma I</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.63</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.66</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.84</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma I</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.95</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.99</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.83</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.79</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma I</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">height</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.65</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.65</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.83</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.66</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.63</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma II</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.97</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.69</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.64</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma II</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.69</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.77</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma II</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">height</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.95</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.73</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.59</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma III</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.96</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.05</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma III</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.79</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.83</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.79</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.95</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma III</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">height</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.63</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.58</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma IV</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.79</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.82</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma IV</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.86</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma IV</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">height</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.59</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.58</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma V</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.83</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.92</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.84</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma V</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.91</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.59</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.63</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.59</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma V</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">height</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.73</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.69</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.61</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">25.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">25.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">28.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">25.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">25.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">18.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">19.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">20.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">19.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">27.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">26.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">25.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">26.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">26.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">24.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">25.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">26.19</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Telson</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">vesicle length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.37</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.43</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Telson</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">vesicle width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.95</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.95</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.95</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.96</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.03</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Telson</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">vesicle height</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.97</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.11</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Telson</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">aculeus length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.99</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.63</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.57</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Telson</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">total length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.92</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.98</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.41</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Metasoma+ Telson</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">total length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">32.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">30.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">35.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">30.98</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">31.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">29.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">29.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">31.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">30.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">34.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">32.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">32.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">33.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">33.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">30.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">31.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">32.60</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Femur</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.98</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.96</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.61</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Femur</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.69</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.60</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Patella</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">5.78</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.21</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Patella</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.94</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Chela</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">13.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">12.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10.79</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.63</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Chela</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">width</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.96</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.02</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Chela</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">height</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.73</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.92</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.95</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">1.93</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Chela</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">movable finger length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8.97</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.51</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.99</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Chela</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">fixed finger length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.91</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6.92</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">Chela</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">palm length</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">3.92</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">4.03</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="coloration">
|
||
<paragraph pageId="0" pageNumber="33">Coloration.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
General pattern (in ethanol 70%) (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33">7</figureCitation>
|
||
): light yellow, without variegated pigmentation.
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Carapace</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33">7A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), carapace, dorsal view A white light B UV light. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1075.67459.figure8" httpUri="https://binary.pensoft.net/fig/619814" pageId="0" pageNumber="33">8A</figureCitation>
|
||
): light yellow; lateral and median eyes, surrounded by black variegated pigments.
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Chelicerae</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33">7A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), carapace, dorsal view A white light B UV light. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1075.67459.figure8" httpUri="https://binary.pensoft.net/fig/619814" pageId="0" pageNumber="33">8A</figureCitation>
|
||
): coxa and hand light yellow, without pigments; fingers, dark reddish-brown.
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Mesosoma, coxosternal region, pedipalps, legs</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33">7A, B</figureCitation>
|
||
): all light yellow.
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Metasoma</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33">7A, B</figureCitation>
|
||
): segments light yellow, progressively becoming darker towards the telson.
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Telson</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33">7A, B</figureCitation>
|
||
): dark yellow; aculeus dark reddish-brown. Live coloration pattern (Fig.
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tityus (Tityus) spelaeus sp. nov., female paratypes under laboratory conditions A, B female paratype with scorpionlings A feeding upon scorpionlings B litter on female's back C specimens feeding on a cricket." figureDoi="10.3897/zookeys.1075.67459.figure13" httpUri="https://binary.pensoft.net/fig/619819" pageId="0" pageNumber="33">13A-C</figureCitation>
|
||
) similar to that of preserved specimens, except for mesosoma with a faint brown median stripe crossing all tergites, telson light reddish-brown, pedipalp chela fingers and metasomal segments IV-V dark reddish-brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="morphology">
|
||
<paragraph pageId="0" pageNumber="33">Morphology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Carapace</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogeny of Tityus representatives from South America obtained by analysis of DNA sequences (12 S rDNA, 16 S rDNA, 28 S rDNA, and Cytochrome c Oxidase I- COI), showing the distribution of the characters states of the ventral setae of telotarsi I-IV (orange: an irregularly distributed tuft of setae (type I); turquoise: two ventro-submedian rows of setae (type II )) across different Tityus subgenera and species-groups. Boxes on branches and associated values correspond to ultrafast-bootstrap (Ubst) values. Observations = Tityus (Tityus) spelaeus sp. nov. is marked in bold." figureDoi="10.3897/zookeys.1075.67459.figure2" httpUri="https://binary.pensoft.net/fig/619808" pageId="0" pageNumber="33">2B</figureCitation>
|
||
): densely covered with fine granulation and few coarse granules; anterior margin with deep median notch; anterior median carinae only feebly marked over anterior 1/3; central lateral, central median, lateral ocular, posterior, posterior median and superciliary carinae, all well-marked; and furrows (anterior median, anterior marginal, central transverse, lateral ocular, supercialiary, posterior transverse, posterior lateral and posterior marginal), all well-marked; ocular tubercle well-marked, located on the anterior half of carapace; median eyes separated by about 0.53 ocular diameters; with three pairs of lateral eyes and two pairs of lateral micro-ocelli.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Chelicerae</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), carapace, dorsal view A white light B UV light. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1075.67459.figure8" httpUri="https://binary.pensoft.net/fig/619814" pageId="0" pageNumber="33">8B</figureCitation>
|
||
): dentition characteristic of the family
|
||
<taxonomicName authorityName="C. L. Koch" authorityYear="1837" class="Arachnida" family="Buthidae" kingdom="Animalia" lsidName="" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Vachon, M" journalOrPublisher="Subterranean Biology" pageId="0" pageNumber="33" refId="B87" refString="Vachon, M, 1963. De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris: 161-166." title="De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris: 161 - 166." year="1963">Vachon 1963</bibRefCitation>
|
||
), densely covered with setae over the internal and ventral surfaces.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Pedipalps</emphasis>
|
||
: Chela, short and slender (female, L/W= 5.5). Orthobothriotaxic pattern Type A, femur with alfa configuration (hand: Eb3:Eb2:Eb1:Esb:Est:Et, fixed finger: eb:esb:est:et:db:dt:it).
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Femur</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), right pedipalp segments A femur, dorsal view B, C patella B dorsal view C external view. Scale bars: 1.5 mm." figureDoi="10.3897/zookeys.1075.67459.figure9" httpUri="https://binary.pensoft.net/fig/619815" pageId="0" pageNumber="33">9A</figureCitation>
|
||
) with five carinae: VI, DI, DE, and VE crenulate, EM serratocrenulate, complete and pronounced, with intercarinal areas densely covered with fine granulation and few coarse granules.
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Patella</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), right pedipalp segments A femur, dorsal view B, C patella B dorsal view C external view. Scale bars: 1.5 mm." figureDoi="10.3897/zookeys.1075.67459.figure9" httpUri="https://binary.pensoft.net/fig/619815" pageId="0" pageNumber="33">9B, C</figureCitation>
|
||
) with seven carinae: VI, VE, DI, DE, and EM complete and crenulate; DM incomplete and crenulate; IM complete and serratocrenulate, with a short spiniform granule near the segment base; with intercarinal areas densely covered with fine granulation.
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Chela</emphasis>
|
||
(tibia) (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), chela A external view B dorsal view C ventral view. Scale bar: 3 mm." figureDoi="10.3897/zookeys.1075.67459.figure10" httpUri="https://binary.pensoft.net/fig/619816" pageId="0" pageNumber="33">10A-C</figureCitation>
|
||
) with eight carinae: VI, VE, D, DS, DMA, IM, and ES, complete and crenulate; SA, incomplete and crenulate, only present on the anterior half of the hand. Pedipalp movable and fixed fingers without basal lobe (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), chela A external view B dorsal view C ventral view. Scale bar: 3 mm." figureDoi="10.3897/zookeys.1075.67459.figure10" httpUri="https://binary.pensoft.net/fig/619816" pageId="0" pageNumber="33">10A</figureCitation>
|
||
). Movable finger with 17-17 rows.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1075.67459.figure9" httpUri="https://binary.pensoft.net/fig/619815" pageId="0" pageNumber="33" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 9.</emphasis>
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus) spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus" subGenus="Tityus">Tityus (Tityus) spelaeus</taxonomicName>
|
||
sp. nov., female holotype (MZSP 74633), right pedipalp segments
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
femur, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B, C</emphasis>
|
||
patella
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
external view. Scale bars: 1.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1075.67459.figure10" httpUri="https://binary.pensoft.net/fig/619816" pageId="0" pageNumber="33" start="Figure 10" startId="F10">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 10.</emphasis>
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus) spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus" subGenus="Tityus">Tityus (Tityus) spelaeus</taxonomicName>
|
||
sp. nov., female holotype (MZSP 74633), chela
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
external view
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
ventral view. Scale bar: 3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Coxosternal region</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633) A dorsal view B ventral view. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1075.67459.figure7" httpUri="https://binary.pensoft.net/fig/619813" pageId="0" pageNumber="33">7B</figureCitation>
|
||
): Sternum with posterior depression, outer ridge, and apical button, well-marked; sclerite covered with fine granulation, and few setae, except for the coxapophyses I-II, which are smooth; genital operculum longitudinally divided, composed of two sub-triangular plates.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Pectines</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tityus (Tityus) spelaeus sp. nov., female paratype (MZSP 74633) genital area and pectines A genital operculum and pectinal basal piece B closeup of the pectinal basal piece, showing cuticular pores on glandular region C right pectine D peg sensillae, distribution E closeup of a peg sensilla. Scale bars: 800 µm (A); 60 µm (B); 1000 µm (C); 20 µm (D); 3 µm (E)." figureDoi="10.3897/zookeys.1075.67459.figure11" httpUri="https://binary.pensoft.net/fig/619817" pageId="0" pageNumber="33">11</figureCitation>
|
||
). Pectinal basal piece sub-rectangular and covered with a large and raised glandular region occupying beyond the anterior two thirds of the anteromedian region (Figs
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Atreus) obscurus Gervais, 1843 (MNRJ 07610) C, D Tityus (Tityus) serrulatus Lutz & Mello, 1922 (MZSP 28205) E, F Tityus (Tityus) spelaeus sp. nov. (MZSP 74633). Abbreviations: BML, basal middle lamellae; GR, glandular region of the pectinal basal piece. Scale bars: 500 μm." figureDoi="10.3897/zookeys.1075.67459.figure6" httpUri="https://binary.pensoft.net/fig/619812" pageId="0" pageNumber="33">6E, F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tityus (Tityus) spelaeus sp. nov., female paratype (MZSP 74633) genital area and pectines A genital operculum and pectinal basal piece B closeup of the pectinal basal piece, showing cuticular pores on glandular region C right pectine D peg sensillae, distribution E closeup of a peg sensilla. Scale bars: 800 µm (A); 60 µm (B); 1000 µm (C); 20 µm (D); 3 µm (E)." figureDoi="10.3897/zookeys.1075.67459.figure11" httpUri="https://binary.pensoft.net/fig/619817" pageId="0" pageNumber="33">11A, B</figureCitation>
|
||
; Table
|
||
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Phenotypic characters useful for the taxonomy of Tityus. (***) Species here transferred to the indicated subgenus; ventral macrosetae distribution on telotarsi I-IV: Type I = tuft of irregularly distributed macrosetae. Type II = two discrete ventrosubmedian rows of macrosetae. Abbreviations: BML, basal middle lamellae; D, dilated; NA, not applicable; ND, not dilated; PBP, pectinal basal piece." httpUri="http://table.plazi.org/id/ED09BE3D09BD0AD5F4D3DFBDE0DEAF3E" pageId="0" pageNumber="33" tableUuid="ED09BE3D09BD0AD5F4D3DFBDE0DEAF3E">3</tableCitation>
|
||
); pectinal tooth count of 19-22. Marginal lamellae, median lamellae, and fulcra moderately covered with setae (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (MZSP). Abbreviations: BML, basal middle lamellae; GR, glandular region. Scale bars: 500 μm." figureDoi="10.3897/zookeys.1075.67459.figure5" httpUri="https://binary.pensoft.net/fig/619811" pageId="0" pageNumber="33">5C</figureCitation>
|
||
). Basal middle lamellae, not dilated (Figs
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tityus C. L. Koch, 1836, female pectinal basal piece and basal middle lamellae of the pectines, showing glandular regions A, C, E White light B, D, F UV light A, B Tityus (Atreus) obscurus Gervais, 1843 (MNRJ 07610) C, D Tityus (Tityus) serrulatus Lutz & Mello, 1922 (MZSP 28205) E, F Tityus (Tityus) spelaeus sp. nov. (MZSP 74633). Abbreviations: BML, basal middle lamellae; GR, glandular region of the pectinal basal piece. Scale bars: 500 μm." figureDoi="10.3897/zookeys.1075.67459.figure6" httpUri="https://binary.pensoft.net/fig/619812" pageId="0" pageNumber="33">6E, F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tityus (Tityus) spelaeus sp. nov., female paratype (MZSP 74633) genital area and pectines A genital operculum and pectinal basal piece B closeup of the pectinal basal piece, showing cuticular pores on glandular region C right pectine D peg sensillae, distribution E closeup of a peg sensilla. Scale bars: 800 µm (A); 60 µm (B); 1000 µm (C); 20 µm (D); 3 µm (E)." figureDoi="10.3897/zookeys.1075.67459.figure11" httpUri="https://binary.pensoft.net/fig/619817" pageId="0" pageNumber="33">11C</figureCitation>
|
||
). Pectinal tooth peg sensillae rectangular in cross-section, with a narrow distal openning (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Tityus (Tityus) spelaeus sp. nov., female paratype (MZSP 74633) genital area and pectines A genital operculum and pectinal basal piece B closeup of the pectinal basal piece, showing cuticular pores on glandular region C right pectine D peg sensillae, distribution E closeup of a peg sensilla. Scale bars: 800 µm (A); 60 µm (B); 1000 µm (C); 20 µm (D); 3 µm (E)." figureDoi="10.3897/zookeys.1075.67459.figure11" httpUri="https://binary.pensoft.net/fig/619817" pageId="0" pageNumber="33">11D, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1075.67459.figure11" httpUri="https://binary.pensoft.net/fig/619817" pageId="0" pageNumber="33" start="Figure 11" startId="F11">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 11.</emphasis>
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus) spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus" subGenus="Tityus">Tityus (Tityus) spelaeus</taxonomicName>
|
||
sp. nov., female paratype (MZSP 74633) genital area and pectines
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
genital operculum and pectinal basal piece
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
closeup of the pectinal basal piece, showing cuticular pores on glandular region
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
right pectine
|
||
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
peg sensillae, distribution
|
||
<emphasis bold="true" pageId="0" pageNumber="33">E</emphasis>
|
||
closeup of a peg sensilla. Scale bars: 800
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
); 60
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
); 1000
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
); 20
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
); 3
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="33">E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Legs</emphasis>
|
||
: Carinae present; intercarinal areas with sparse fine granulation; ventral telotarsal macrosetae acute and fine, arranged in two ventrosubmedian rows (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tityus C. L. Koch, 1836, telotarsi IV, showing ventral macrosetae A, C, E, G, I, K white light B, D, F, H, J, L UV light A, B Tityus (Tityus) brazilae Lourenco & Eickstedt, 1984 (type II) (MZSP 75619) C, D Tityus (Archaeotityus) clathratus C. L. Koch, 1844 (type I) (MZSP 31468) E, F Tityus (Atreus) forcipula (Gervais, 1843) (type II) (MZSP) G, H Tityus (Atreus) obscurus Gervais, 1843 (type I) (MNRJ 07610) I, J Tityus (Tityus) serrulatus Lutz & Mello, 1922 (type II) (MZSP 28205) K, L Tityus (Tityus) spelaeus sp. nov. (MZSP 74633) (type II). Observations = telotarsi I-IV ventral setae distribution: Type I = tuft of irregularly distributed setae. Type II = two discrete ventrosubmedian rows of setae. Scale bars: 500 μm." figureDoi="10.3897/zookeys.1075.67459.figure4" httpUri="https://binary.pensoft.net/fig/619810" pageId="0" pageNumber="33">4K, L</figureCitation>
|
||
); telotarsi, counts of ventral macrosetae in the left (L) and right (R) legs on prolateral (pro) and retrolateral (retro) rows of legs I to IV (L (pro/retro) R (pro/retro)): 7/6 7/7: 7/7 7/7: 9/8 9/10: 10/10 9/11. Claws short and symmetrical.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Mesosoma</emphasis>
|
||
: Tergites I-VI, moderately covered with fine granulation and few coarse granules; pre-tergites well defined, with median carina visible on the posterior margin of the post-tergites; tergite VII with DSM and DL carinae complete and crenulate, and median carina composed of a crenulate anteromedian eminence present on the anterior half of the post-tergite. Sternites densely covered with fine granulation; sternites III-VI with a pair of elliptic spiracles on the posterior half, which are progressively larger; sternite V with a hyaline subtriangular area on the posterior margin; sternite VI with VSM carinae crenulate, present on posterior half; sternite VII with VSM and VL carinae crenulate, present on posterior two thirds.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Metasoma</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33">12C, D</figureCitation>
|
||
): Segments II-V short and robust (L/W ratio: II= 1.9; III= 1.9; IV= 2.0; V= 2.5); segment V not incrassate (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33">12C</figureCitation>
|
||
). Segments I-II (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33">12C, D</figureCitation>
|
||
) with 10 complete carinae, parallel to one another and crenulate (paired DSM, DL, ML, VL, and VSM), ML of segment II represented by coarse granules on posterior two thirds, intercarinal areas densely covered with fine granulation; segments III-IV (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33">12C, D</figureCitation>
|
||
) with eight complete carinae, parallel to one another and crenulate (paired DSM, DL, VL, and VSM), intercarinal areas densely covered with fine granulation; segment V (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33">12C, D</figureCitation>
|
||
) with five complete carinae, crenulate (VM, paired DSM, and VL: DSM carinae feebly marked), intercarinal areas moderately covered with fine granulation and few coarse granules. Segments II-IV (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33">12C</figureCitation>
|
||
) with DSM carinae feebly marked, composed of evenly sized granules, without enlarged distoterminal granule.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33" start="Figure 12" startId="F12">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 12.</emphasis>
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus) spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus" subGenus="Tityus">Tityus (Tityus) spelaeus</taxonomicName>
|
||
sp. nov., female holotype (MZSP 74633), metasoma and telson
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A, B</emphasis>
|
||
telson
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C, D</emphasis>
|
||
metasoma
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
ventral view. Scale bars: 2 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A, B</emphasis>
|
||
); 5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C, D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Metasomal macrosetae</emphasis>
|
||
: Segments I-IV each with two pairs of VSM macrosetae (2/2): pair of VSM1 located on the anterior third, and pair of VSM2 located near posterior margin of segment; and with two pairs of VL macrosetae (2/2): pair of VL1 located near anterior margin of segment, and pair of VL2 located on posterior two thirds of segment. Segment V with two pairs of VSM macrosetae (2/2), two pairs of VL macrosetae (2/2), and a single pair of ML macrosetae (1/1); pairs of VSM1 and VL1 located near anterior margin of segment; pair of VL2 located on posterior two thirds of segment, and pair of ML1 located dorsolaterally behind the DSM carinae near posterior margin of segment; anal arch with two pairs of setae on the intercrestal area: one pair of VSM macrosetae (1/1) and one pair of VL macrosetae (1/1).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Telson</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33">12A, B</figureCitation>
|
||
): Vesicle suboval, not elongated (L/H= 1.8), dorsal surface smooth, lateral surfaces with shallow longitudinal furrow; with VM, paired VSM, VL, and DL carinae, vestigial. Subaculear tubercle large, conical, with spiniform apex directed towards the distal region of the aculeus (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), metasoma and telson A, B telson A lateral view B ventral view C, D metasoma C lateral view D ventral view. Scale bars: 2 mm (A, B); 5 mm (C, D)." figureDoi="10.3897/zookeys.1075.67459.figure12" httpUri="https://binary.pensoft.net/fig/619818" pageId="0" pageNumber="33">12A</figureCitation>
|
||
); subaculear tubercle with ventral pair of small, rounded granules, pointing towards the basal portion of the aculeus; aculeus strongly curved, shorter than vesicle and with ventral groove.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="variability">
|
||
<paragraph pageId="0" pageNumber="33">Variability (females).</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Morphometrics.</emphasis>
|
||
Total length (including telson): 48.30-57.98 mm (n= 17, mean= 52.70, standard deviation (SD)= 2.66). Chela L/W ratio: 4.42-5.78 (n= 17, mean= 5.44, SD= 0.37). Metasomal segment I L/W ratio: 1.24-1.51 (n= 17, mean= 1.30, SD= 0.06). Metasomal segment V L/W ratio: 2.31-2.68 (n= 17, mean 2.52, SD= 0.12). Telson vesicle L/H: 1.63-1.89 (n= 17, mean= 1.76, SD= 0.09).
|
||
<emphasis bold="true" pageId="0" pageNumber="33">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Meristics</emphasis>
|
||
.
|
||
</emphasis>
|
||
Pectinal tooth count: 19-22 (n= 34, mode= 20). Number of movable finger oblique granular rows: 16-18 (n= 34, mode= 18). Metasomal macrosetae count: (n= 17): 2/2 VSM and 2/2 VL macrosetae on segments I-IV, 3/3 VSM and 2/2 VL macrosetae on segment V. However, one specimen (LES/UFSCar 14668) lost VSM1 on segment II, a second specimen (LES/UFSCAR 14669) lost VL1 on segment II, and a third specimen (LES/UFSCAR 014673) lost one VSM1 on segment I. Variation in the count of telotarsal ventrosubmedian setae is presented in Table
|
||
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Variation in the number of macrosetae of the ventrosubmedian setal rows on telotarsi I-IV across paratypes of Tityus spelaeus sp. nov. Abbreviations: L, left leg; Pl, prolateral row; Rl, retrolateral row; R, right leg." httpUri="http://table.plazi.org/id/85F82781ADEFF51DC133AD43BE48DE9A" pageId="0" pageNumber="33" tableUuid="85F82781ADEFF51DC133AD43BE48DE9A">5</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="85F82781ADEFF51DC133AD43BE48DE9A" httpUri="http://table.plazi.org/id/85F82781ADEFF51DC133AD43BE48DE9A" pageId="0" pageNumber="33" start="Table 5" startId="T5">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Table 5.</emphasis>
|
||
Variation in the number of macrosetae of the ventrosubmedian setal rows on telotarsi I-IV across paratypes of
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus spelaeus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Abbreviations: L, left leg; Pl, prolateral row; Rl, retrolateral row; R, right leg.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<table pageId="0" pageNumber="33">
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="2">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Telotarsus</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">MZSP 74634 (1)</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">MZSP 74634 (2)</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">MZSP 74634 (3)</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">MZSP 74634 (4)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">I</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/6</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6/7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">II</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">III</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">IV</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/11</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="2">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Telotarsus</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">MZSP 52228</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">MZSP 52230</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">MZSP 52229</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">MZSP 52231</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">I</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6/6</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/6</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/6</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6/8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">II</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/6</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">III</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">7/6</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6/6</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">IV</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/11</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/10</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="2">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Telotarsus</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">LES 14668</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">LES 14669</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">LES 14670</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">LES 14671</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">I</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">6/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">II</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/7</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">III</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">IV</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11/11</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/11</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/11</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">12/12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/11</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="2">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Telotarsus</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">LES 14672</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">LES 14673-1</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">LES 14673-2</emphasis>
|
||
</td>
|
||
<td colspan="2" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">LES 14673-3</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">L(Pl/Rl)</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">R(Pl/Rl)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">I</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">II</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">III</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8/8</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/9</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9/8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="33">
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">IV</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10/11</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11/12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11/10</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">12/12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11/12</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">12/11</td>
|
||
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11/12</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="natural history">
|
||
<paragraph pageId="0" pageNumber="33">Natural history.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<normalizedToken originalValue="Russão">Russao</normalizedToken>
|
||
II cave is formed by limestone (a karstified type of rock), located in Posse municipality, the northeastern state of
|
||
<normalizedToken originalValue="Goiás">Goias</normalizedToken>
|
||
, Central Brazil. This karst region is part of the
|
||
<normalizedToken originalValue="Bambuí">Bambui</normalizedToken>
|
||
geomorphological group, the large geomorphological group in Brazil, occurring in states of Bahia,
|
||
<normalizedToken originalValue="Goiás">Goias</normalizedToken>
|
||
, Minas Gerais, and Tocantins.
|
||
<normalizedToken originalValue="Russão">Russao</normalizedToken>
|
||
II cave is inserted on the Cerrado morphoclimatic domain (
|
||
<bibRefCitation author="Ab'Saber, NA" journalOrPublisher="Geomorfologia" pageId="0" pageNumber="33" pagination="1 - 21" refId="B1" refString="Ab'Saber, NA, 1977. Os dominios morfoclimaticos na America do Sul. Geomorfologia 52: 1 - 21" title="Os dominios morfoclimaticos na America do Sul." volume="52" year="1977">
|
||
<normalizedToken originalValue="Ab’Saber">Ab'Saber</normalizedToken>
|
||
1977
|
||
</bibRefCitation>
|
||
), and the climate is tropical semi-humid (
|
||
<bibRefCitation author="Nimer, E" journalOrPublisher="Universidade de Sao Paulo, Sao Paulo" pageId="0" pageNumber="33" refId="B51" refString="Nimer, E, 1979. Climatologia do Brasil. Vol. 4. Rio de Janeiro: SUPREN." title="Climatologia do Brasil. Vol. 4. Rio de Janeiro: SUPREN." year="1979">Nimer 1979</bibRefCitation>
|
||
). There is a stream crossing the cave although there are no surface drainages nearby (
|
||
<bibRefCitation DOI="https://doi.org/10.1371/journal.pone.0171309" author="Tencatt, LFC" journalOrPublisher="Entomological News" pageId="0" pageNumber="33" refId="B80" refString="Tencatt, LFC, Bichuette, ME, 2017. Aspidoras mephisto, new species: the first troglobitic Callichthyidae (Teleostei: Siluriformes) from South America. PLoS ONE 12: e0171309. https://doi.org/10.1371/journal.pone.0171309" title="Aspidoras mephisto, new species: the first troglobitic Callichthyidae (Teleostei: Siluriformes) from South America. PLoS ONE 12: e 0171309." url="https://doi.org/10.1371/journal.pone.0171309" year="2017">Tencatt and Bichuette 2017</bibRefCitation>
|
||
).
|
||
<normalizedToken originalValue="Russão">Russao</normalizedToken>
|
||
II cave is located on private property, and in addition surface habitats are under impact from pollution through the discharge of domestic sewage, deforestation of surroundings for cattle pasture, and small mining projects (
|
||
<bibRefCitation DOI="https://doi.org/10.1371/journal.pone.0171309" author="Tencatt, LFC" journalOrPublisher="Entomological News" pageId="0" pageNumber="33" refId="B80" refString="Tencatt, LFC, Bichuette, ME, 2017. Aspidoras mephisto, new species: the first troglobitic Callichthyidae (Teleostei: Siluriformes) from South America. PLoS ONE 12: e0171309. https://doi.org/10.1371/journal.pone.0171309" title="Aspidoras mephisto, new species: the first troglobitic Callichthyidae (Teleostei: Siluriformes) from South America. PLoS ONE 12: e 0171309." url="https://doi.org/10.1371/journal.pone.0171309" year="2017">Tencatt and Bichuette 2017</bibRefCitation>
|
||
).
|
||
<normalizedToken originalValue="Russão">Russao</normalizedToken>
|
||
II cave, like other caves in the region, has no legal protection under Brazilian environmental laws. The cave has a significant amount of bat guano piles and a large cricket population that is preyed upon by scorpions. In the aphotic zone of
|
||
<normalizedToken originalValue="Russão">Russao</normalizedToken>
|
||
II cave, the temperature was 30.04 °C, the relative humidity of the air was 72.02%.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
This species was studied in the past by
|
||
<bibRefCitation DOI="https://doi.org/10.1590/s1984-46702009000100008" author="Outeda-Jorge, S" journalOrPublisher="Zoologia (Curitiba)" pageId="0" pageNumber="33" pagination="43 - 53" refId="B55" refString="Outeda-Jorge, S, Mello, T, Pinto-da-Rocha, R, 2009. Litter size, effects of maternal body size, and date of birth in South American scorpions (Arachnida: Scorpiones). Zoologia (Curitiba) 26: 43 - 53, DOI: https://doi.org/10.1590/s1984-46702009000100008" title="Litter size, effects of maternal body size, and date of birth in South American scorpions (Arachnida: Scorpiones)." url="https://doi.org/10.1590/s1984-46702009000100008" volume="26" year="2009">Outeda-Jorge et al. (2009)</bibRefCitation>
|
||
who reported a litter size of two scorpionlings, but under laboratory conditions (Fig.
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tityus (Tityus) spelaeus sp. nov., female paratypes under laboratory conditions A, B female paratype with scorpionlings A feeding upon scorpionlings B litter on female's back C specimens feeding on a cricket." figureDoi="10.3897/zookeys.1075.67459.figure13" httpUri="https://binary.pensoft.net/fig/619819" pageId="0" pageNumber="33">13A-C</figureCitation>
|
||
), another two females had a litter of four scorpionlings, and both females were fed upon their litter (Fig.
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Tityus (Tityus) spelaeus sp. nov., female paratypes under laboratory conditions A, B female paratype with scorpionlings A feeding upon scorpionlings B litter on female's back C specimens feeding on a cricket." figureDoi="10.3897/zookeys.1075.67459.figure13" httpUri="https://binary.pensoft.net/fig/619819" pageId="0" pageNumber="33">13A, B</figureCitation>
|
||
). The population of
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Tityus spelaeus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. at the
|
||
<normalizedToken originalValue="Russão">Russao</normalizedToken>
|
||
cave is well-established (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Tityus (Tityus) spelaeus sp. nov., female holotype (MZSP 74633), carapace, dorsal view A white light B UV light. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1075.67459.figure8" httpUri="https://binary.pensoft.net/fig/619814" pageId="0" pageNumber="33">8A, B</figureCitation>
|
||
). During a one-hour-long visit to the cave in 2007, more than 20 live scorpions were observed on the ground and walls (Fig.
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Habitat of Tityus (Tityus) spelaeus sp. nov. in the Russao cave A inside landscape of the cave B females on the cave walls." figureDoi="10.3897/zookeys.1075.67459.figure14" httpUri="https://binary.pensoft.net/fig/619820" pageId="0" pageNumber="33">14A, B</figureCitation>
|
||
). In another two-hour visit in 2015, 32 individuals were counted, both adults and juveniles.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1075.67459.figure13" httpUri="https://binary.pensoft.net/fig/619819" pageId="0" pageNumber="33" start="Figure 13" startId="F13">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 13.</emphasis>
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus) spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus" subGenus="Tityus">Tityus (Tityus) spelaeus</taxonomicName>
|
||
sp. nov., female paratypes under laboratory conditions
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A, B</emphasis>
|
||
female paratype with scorpionlings
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
feeding upon scorpionlings
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
litter on
|
||
<normalizedToken originalValue="female’s">female's</normalizedToken>
|
||
back
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
specimens feeding on a cricket.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1075.67459.figure14" httpUri="https://binary.pensoft.net/fig/619820" pageId="0" pageNumber="33" start="Figure 14" startId="F14">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 14.</emphasis>
|
||
Habitat of
|
||
<taxonomicName authorityName="Moreno-González 1 & Pinto-da-Rocha 1 & Gallão 2" authorityYear="2021" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus) spelaeus" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="spelaeus" subGenus="Tityus">Tityus (Tityus) spelaeus</taxonomicName>
|
||
sp. nov. in the
|
||
<normalizedToken originalValue="Russão">Russao</normalizedToken>
|
||
cave
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
inside landscape of the cave
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
females on the cave walls.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |