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<document id="BA4252BEBAA697446F5D466953A8BA21" ID-DOI="10.11646/zootaxa.2186.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5311592" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629629438740" checkinUser="felipe" docAuthor="Jones, Joshua R. &amp; Deitz, Lewis L." docDate="2009" docId="CF7A87E4FFF689737D9DA46ABADEFEF8" docLanguage="en" docName="zt02186p120.pdf" docOrigin="Zootaxa 2186 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2186.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Arenoledra Kuoh 1992" docType="treatment" docVersion="4" lastPageNumber="34" masterDocId="3343FF9CFFD689527D0AA225BA23FC37" masterDocTitle="Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae) 2186" masterLastPageNumber="120" masterPageNumber="1" pageNumber="33" updateTime="1699086546085" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CE1A85F9A4562BED8B70370BEEB99B15">Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae) 2186</mods:title>
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<mods:namePart id="F8AEAC6909069F87CB6E1E0EC01C2A86">Deitz, Lewis L.</mods:namePart>
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<mods:date id="FC8DAC232FCD5CFFA96D4F7D012B88F2">2009</mods:date>
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<mods:number id="5FF953A796F553460C5ED84F06B46463">2186</mods:number>
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<treatment id="CF7A87E4FFF689737D9DA46ABADEFEF8" ID-DOI="http://doi.org/10.5281/zenodo.5777744" ID-Zenodo-Dep="5777744" LSID="urn:lsid:plazi:treatment:CF7A87E4FFF689737D9DA46ABADEFEF8" httpUri="http://treatment.plazi.org/id/CF7A87E4FFF689737D9DA46ABADEFEF8" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<subSubSection id="0FC96579FFF689727D9DA46ABBE7FA5E" box="[151,452,1615,1641]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="476C36F2FFF689727D9DA46ABBE7FA5E" blockId="32.[151,452,1615,1641]" box="[151,452,1615,1641]" pageId="32" pageNumber="33">
<heading id="1C24819EFFF689727D9DA46ABBE7FA5E" bold="true" box="[151,452,1615,1641]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="75A7EAE0FFF689727D9DA46ABBE7FA5E" bold="true" box="[151,452,1615,1641]" pageId="32" pageNumber="33">
Genus
<taxonomicName id="80D34D71FFF689727DE4A46ABBE7FA5E" ID-CoL="8HD7S" authority="Kuoh" authorityName="Kuoh" authorityYear="1992" box="[238,452,1615,1641]" class="Insecta" family="Cicadellidae" genus="Arenoledra" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF689727DE4A46ABB54FA5E" bold="true" box="[238,375,1615,1641]" italics="true" pageId="32" pageNumber="33">Arenoledra</emphasis>
Kuoh
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0FC96579FFF689727D9DA4BCBBDCFA87" box="[151,511,1689,1712]" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="476C36F2FFF689727D9DA4BCBBDCFA87" blockId="32.[151,1067,1689,1776]" box="[151,511,1689,1712]" pageId="32" pageNumber="33">
<treatmentCitationGroup id="67C311DCFFF689727D9DA4BCBBDCFA87" box="[151,511,1689,1712]" pageId="32" pageNumber="33">
<treatmentCitation id="C67210E3FFF689727D9DA4BCBBE5FA87" author="Kuoh, C. &amp; Cai, P." box="[151,454,1689,1712]" page="254" pageId="32" pageNumber="33" year="1992">
<taxonomicName id="80D34D71FFF689727D9DA4BCBBE5FA87" ID-CoL="8HD7S" authority="Kuoh, 1992: 254" authorityName="Kuoh" authorityPageNumber="254" authorityYear="1992" box="[151,454,1689,1712]" class="Insecta" family="Cicadellidae" genus="Arenoledra" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF689727D9DA4BCBB33FA87" box="[151,272,1689,1712]" italics="true" pageId="32" pageNumber="33">Arenoledra</emphasis>
<bibRefCitation id="23424B03FFF689727C1FA4BCBBE5FA87" author="Kuoh, C. &amp; Cai, P." box="[277,454,1689,1712]" pageId="32" pageNumber="33" pagination="136 - 138" refId="ref57812" refString="Kuoh, C. - L and Cai, P. (1992) A new species of the genus Epiclinata from China (Homoptera: Cicadelloidea: Ledridae). Acta Zoologica Sinica, 38, 136 - 138." type="journal article" year="1992">Kuoh, 1992:254</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 307.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0FC96579FFF689727D9DA49CBB73FAC7" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="476C36F2FFF689727D9DA49CBE08FAE7" blockId="32.[151,1067,1689,1776]" box="[151,1067,1721,1744]" pageId="32" pageNumber="33">
<emphasis id="75A7EAE0FFF689727D9DA49CBB3FFAE7" box="[151,284,1721,1744]" italics="true" pageId="32" pageNumber="33">
<typeStatus id="98688850FFF689727D9DA49CBAEAFAE7" box="[151,201,1721,1744]" pageId="32" pageNumber="33">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="80D34D71FFF689727C20A49CB8B9FAE7" authority="Kuoh, 1992: 255" authorityName="Kuoh" authorityPageNumber="255" authorityYear="1992" box="[298,666,1721,1744]" class="Insecta" family="Cicadellidae" genus="Arenoledra" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nigrimaculata">
<emphasis id="75A7EAE0FFF689727C20A49CBBFDFAE7" box="[298,478,1721,1744]" italics="true" pageId="32" pageNumber="33">A. nigrimaculata</emphasis>
<bibRefCitation id="23424B03FFF689727CEEA49CB8B9FAE7" author="Kuoh, C. &amp; Cai, P." box="[484,666,1721,1744]" pageId="32" pageNumber="33" pagination="136 - 138" refId="ref57812" refString="Kuoh, C. - L and Cai, P. (1992) A new species of the genus Epiclinata from China (Homoptera: Cicadelloidea: Ledridae). Acta Zoologica Sinica, 38, 136 - 138." type="journal article" year="1992">Kuoh, 1992: 255</bibRefCitation>
</taxonomicName>
, 307, fig. 13, by original designation.
</paragraph>
<paragraph id="476C36F2FFF689727D9DA4FCBB73FAC7" blockId="32.[151,1067,1689,1776]" box="[151,336,1753,1776]" pageId="32" pageNumber="33">
<emphasis id="75A7EAE0FFF689727D9DA4FCBB2AFAC7" box="[151,265,1753,1776]" italics="true" pageId="32" pageNumber="33">Synonymy.</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="0FC96579FFF689737D9DA53AB8B8FC85" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="476C36F2FFF689727D9DA53ABE63FB99" blockId="32.[151,1436,1823,2004]" pageId="32" pageNumber="33">
<emphasis id="75A7EAE0FFF689727D9DA53ABB0CFB0E" bold="true" box="[151,303,1823,1849]" pageId="32" pageNumber="33">Description.</emphasis>
Kuoh (Ge 1992): “The vertex of the head as long as breadth between eyes or shorter, nail shape, the body above densely covered [with] granules as rough as sandpaper, the veins of tegmina radialized towards the apex, especially in the process of pygofer which differs from
<taxonomicName id="80D34D71FFF689727EF7A548BE85FBB1" box="[1021,1190,1901,1926]" class="Insecta" family="Cicadellidae" genus="Petalocephala" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF689727EF7A548BE85FBB1" box="[1021,1190,1901,1926]" italics="true" pageId="32" pageNumber="33">Petalocephala</emphasis>
</taxonomicName>
and the shape of the aedeagus and styli differs from both of them [
<taxonomicName id="80D34D71FFF689727FA5A5B1B97BFB9A" box="[687,856,1940,1965]" class="Insecta" family="Cicadellidae" genus="Petalocephala" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF689727FA5A5B1B97BFB9A" box="[687,856,1940,1965]" italics="true" pageId="32" pageNumber="33">Petalocephala</emphasis>
</taxonomicName>
and
<taxonomicName id="80D34D71FFF689727E98A5B1BE00FB9A" authorityName="McKamey" authorityYear="2006" box="[914,1059,1940,1965]" class="Insecta" family="Cicadellidae" genus="Latycephala" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF689727E98A5B1BE00FB9A" box="[914,1059,1940,1965]" italics="true" pageId="32" pageNumber="33">Latycephala</emphasis>
</taxonomicName>
].”
</paragraph>
<paragraph id="476C36F2FFF689727DCDA59FBEC5FBE3" blockId="32.[151,1436,1823,2004]" box="[199,1254,1978,2004]" pageId="32" pageNumber="33">
<emphasis id="75A7EAE0FFF689727DCDA59FBB0AFBE3" bold="true" box="[199,297,1978,2004]" pageId="32" pageNumber="33">Species.</emphasis>
[4]:
<taxonomicName id="80D34D71FFF689727C68A59EB800FBE3" authority="Kuoh" authorityName="Kuoh" authorityYear="1992" box="[354,547,1978,2004]" class="Insecta" family="Cicadellidae" genus="Arenoledra" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nigrifrons">
<emphasis id="75A7EAE0FFF689727C68A59EBBF5FBE3" box="[354,470,1979,2004]" italics="true" pageId="32" pageNumber="33">nigrifrons</emphasis>
Kuoh
</taxonomicName>
;
<taxonomicName id="80D34D71FFF689727F24A59EB902FBE3" authority="Kuoh" authorityName="Kuoh" authorityYear="1992" box="[558,801,1978,2004]" class="Insecta" family="Cicadellidae" genus="Arenoledra" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nigrimaculata">
<emphasis id="75A7EAE0FFF689727F24A59EB8F7FBE3" box="[558,724,1979,2004]" italics="true" pageId="32" pageNumber="33">nigrimaculata</emphasis>
Kuoh
</taxonomicName>
;
<taxonomicName id="80D34D71FFF689727E26A59EB9C5FBE3" authority="Kuoh" authorityName="Cai &amp; Kuoh" authorityYear="1991" box="[812,998,1978,2004]" class="Insecta" family="Cicadellidae" genus="Petalocephala" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="75A7EAE0FFF689727E26A59EB9BAFBE3" box="[812,921,1979,2004]" italics="true" pageId="32" pageNumber="33">ochracea</emphasis>
Kuoh
</taxonomicName>
;
<taxonomicName id="80D34D71FFF689727EFBA59EBEC5FBE3" authority="Cai &amp; Sun." box="[1009,1254,1978,2004]" class="Insecta" family="Cicadellidae" genus="Arenoledra" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="tubularis">
<emphasis id="75A7EAE0FFF689727EFBA59EBE78FBE3" box="[1009,1115,1979,2004]" italics="true" pageId="32" pageNumber="33">tubularis</emphasis>
Cai &amp; Sun.
</taxonomicName>
</paragraph>
<paragraph id="476C36F2FFF789737DCDA2B2B8B8FC85" blockId="33.[151,1436,151,719]" box="[199,667,151,178]" pageId="33" pageNumber="34">
<emphasis id="75A7EAE0FFF789737DCDA2B2BB3DFC86" bold="true" box="[199,286,151,177]" pageId="33" pageNumber="34">Range.</emphasis>
<collectingCountry id="3FC47662FFF789737C2FA2BDBB48FC85" box="[293,363,152,178]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
(Lushui,
<collectingRegion id="8517F810FFF789737CD6A2BDB81AFC85" box="[476,569,152,178]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Weixi).
</paragraph>
</subSubSection>
<subSubSection id="0FC96579FFF789737DCDA29BBBFCFCEF" box="[199,479,190,216]" pageId="33" pageNumber="34" type="biology_ecology">
<paragraph id="476C36F2FFF789737DCDA29BBBFCFCEF" blockId="33.[151,1436,151,719]" box="[199,479,190,216]" pageId="33" pageNumber="34">
<emphasis id="75A7EAE0FFF789737DCDA29BBB7FFCEF" bold="true" box="[199,348,190,216]" pageId="33" pageNumber="34">Host plants.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0FC96579FFF789737DCDA2C0B82AFCC8" box="[199,521,229,255]" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="476C36F2FFF789737DCDA2C0B82AFCC8" blockId="33.[151,1436,151,719]" box="[199,521,229,255]" pageId="33" pageNumber="34">
<emphasis id="75A7EAE0FFF789737DCDA2C0BB98FCC8" bold="true" box="[199,443,229,255]" pageId="33" pageNumber="34">Material examined.</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="0FC96579FFF789737DCDA32EBADEFEF8" pageId="33" pageNumber="34" type="discussion">
<paragraph id="476C36F2FFF789737DCDA32EBADEFEF8" blockId="33.[151,1436,151,719]" pageId="33" pageNumber="34">
<emphasis id="75A7EAE0FFF789737DCDA32EBB63FD12" bold="true" box="[199,320,267,293]" pageId="33" pageNumber="34">Remarks.</emphasis>
The
<typeStatus id="98688850FFF789737C77A329BB8CFD11" box="[381,431,268,294]" pageId="33" pageNumber="34">type</typeStatus>
specimen of
<taxonomicName id="80D34D71FFF789737F44A329B9A2FD12" authorityName="Kuoh" authorityYear="1992" box="[590,897,268,293]" class="Insecta" family="Cicadellidae" genus="Arenoledra" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="nigrimaculata">
<emphasis id="75A7EAE0FFF789737F44A329B9A2FD12" box="[590,897,268,293]" italics="true" pageId="33" pageNumber="34">Arenoledra nigrimaculata</emphasis>
</taxonomicName>
, which was not made available for this study, appears to be located in either the Institute of Zoology, Chinese Academy of Sciences (Academica Sinica),
<collectingRegion id="8517F810FFF789737D9DA37CBAD3FD44" box="[151,240,345,371]" country="China" name="Beijing" pageId="33" pageNumber="34">Beijing</collectingRegion>
, or in
<collectingRegion id="8517F810FFF789737C33A37CBBA1FD44" box="[313,386,345,371]" country="China" name="Anhui" pageId="33" pageNumber="34">Anhui</collectingRegion>
Agricultural College,
<collectingRegion id="8517F810FFF789737F80A37CB8F5FD44" box="[650,726,345,371]" country="China" name="Anhui" pageId="33" pageNumber="34">Anhui</collectingRegion>
,
<collectingCountry id="3FC47662FFF789737FE8A37CB90BFD44" box="[738,808,345,371]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
(Ge 1992). Based on Kuoh’s description of the genus and drawings of
<taxonomicName id="80D34D71FFF789737C57A3A5B807FDAE" authorityName="Kuoh" authorityYear="1992" box="[349,548,384,410]" class="Insecta" family="Cicadellidae" genus="Arenoledra" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="nigrimaculata">
<emphasis id="75A7EAE0FFF789737C57A3A5BB4CFDAE" box="[349,367,384,409]" italics="true" pageId="33" pageNumber="34">A</emphasis>
.
<emphasis id="75A7EAE0FFF789737C74A3A5B807FDAE" box="[382,548,384,409]" italics="true" pageId="33" pageNumber="34">nigrimaculata</emphasis>
</taxonomicName>
, the exact phylogenetic placement of this genus is uncertain, but it appears to be very similar to
<taxonomicName id="80D34D71FFF789737CBAA382B833FDF7" box="[432,528,423,448]" class="Insecta" family="Cicadellidae" genus="Midoria" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF789737CBAA382B833FDF7" box="[432,528,423,448]" italics="true" pageId="33" pageNumber="34">Midoria</emphasis>
</taxonomicName>
and
<taxonomicName id="80D34D71FFF789737F52A382B8CAFDF7" authorityName="McKamey" authorityYear="2006" box="[600,745,423,448]" class="Insecta" family="Cicadellidae" genus="Latycephala" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF789737F52A382B8CAFDF7" box="[600,745,423,448]" italics="true" pageId="33" pageNumber="34">Latycephala</emphasis>
</taxonomicName>
. All three genera may be proximate to
<taxonomicName id="80D34D71FFF7897379FBA382BF56FDF7" authorityName="Linnavuori" authorityYear="1972" box="[1265,1397,423,448]" class="Insecta" family="Cicadellidae" genus="Hangklipia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF7897379FBA382BF56FDF7" box="[1265,1397,423,448]" italics="true" pageId="33" pageNumber="34">Hangklipia</emphasis>
</taxonomicName>
or
<taxonomicName id="80D34D71FFF789737D9DA3E8BB54FDD1" authorityName="Kato" authorityYear="1931" box="[151,375,461,486]" class="Insecta" family="Cicadellidae" genus="Parapetalocephala" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF789737D9DA3E8BB54FDD1" box="[151,375,461,486]" italics="true" pageId="33" pageNumber="34">Parapetalocephala</emphasis>
</taxonomicName>
(Kuoh stated that
<taxonomicName id="80D34D71FFF789737F93A3E8B93EFDD1" authorityName="Kuoh" authorityYear="1992" box="[665,797,461,486]" class="Insecta" family="Cicadellidae" genus="Arenoledra" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF789737F93A3E8B93EFDD1" box="[665,797,461,486]" italics="true" pageId="33" pageNumber="34">Arenoledra</emphasis>
</taxonomicName>
was between
<taxonomicName id="80D34D71FFF789737EF0A3E8BE80FDD1" box="[1018,1187,461,486]" class="Insecta" family="Cicadellidae" genus="Petalocephala" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF789737EF0A3E8BE80FDD1" box="[1018,1187,461,486]" italics="true" pageId="33" pageNumber="34">Petalocephala</emphasis>
</taxonomicName>
and
<taxonomicName id="80D34D71FFF78973780BA3E8BFB8FDD1" authorityName="Kuoh" authorityYear="1992" box="[1281,1435,461,486]" class="Insecta" family="Chloropidae" genus="Platycephala" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF78973780BA3E8BFB8FDD1" box="[1281,1435,461,486]" italics="true" pageId="33" pageNumber="34">Platycephala</emphasis>
</taxonomicName>
[=
<taxonomicName id="80D34D71FFF789737DBBA3D1BBF2FE39" authority="McKamey" authorityName="McKamey" authorityYear="2006" box="[177,465,500,526]" class="Insecta" family="Cicadellidae" genus="Latycephala" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF789737DBBA3D1BB61FE3A" box="[177,322,500,525]" italics="true" pageId="33" pageNumber="34">Latycephala</emphasis>
McKamey
</taxonomicName>
]). A 2004 collecting expedition in
<collectingCountry id="3FC47662FFF789737E77A3D1B9F0FE39" box="[893,979,500,526]" name="Taiwan" pageId="33" pageNumber="34">Taiwan</collectingCountry>
in which the first author participated (organized by C. H. Dietrich, INHS, and host M. M. Yang, Chung Hsing University,
<collectingRegion id="8517F810FFF789737975A03FBECEFE03" box="[1151,1261,538,564]" country="Taiwan" name="Taichung" pageId="33" pageNumber="34">Taichung</collectingRegion>
) yielded a few specimens resembling
<taxonomicName id="80D34D71FFF789737CB8A064B815FE6D" authorityName="Kuoh" authorityYear="1992" box="[434,566,577,602]" class="Insecta" family="Cicadellidae" genus="Arenoledra" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="75A7EAE0FFF789737CB8A064B815FE6D" box="[434,566,577,602]" italics="true" pageId="33" pageNumber="34">Arenoledra</emphasis>
</taxonomicName>
, and a number of specimens borrowed from the TARI and NMNS collections vaguely resemble Kuoh’s illustrations, but it will be necessary to examine his
<typeStatus id="98688850FFF7897379D7A04DBF2CFEB5" box="[1245,1295,616,642]" pageId="33" pageNumber="34">type</typeStatus>
material in order to make a positive identification. These specimens ultimately were not included in the phylogenetic analysis.
</paragraph>
</subSubSection>
</treatment>
</document>
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