157 lines
17 KiB
XML
157 lines
17 KiB
XML
<document id="A26DAD2E7B95F5BEDE8F64D133C70B06" ID-CLB-Dataset="36828" ID-DOI="10.11646/zootaxa.3824.1.1" ID-GBIF-Dataset="df227efc-13d9-421e-92ed-f9aa417b0cf7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286404" ID-ZooBank="023D78D0-4182-48D2-BAEB-CDA6473CF585" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460443577100" checkinUser="plazi" docAuthor="Webster, Mark & Bohach, Lisa L." docDate="2014" docId="B10C8793FFE2FFA861B5FAE3FB9082D5" docLanguage="en" docName="zt03824p066.pdf" docOrigin="Zootaxa 3824 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lochmanolenellus primus" docType="treatment" docVersion="6" lastPageNumber="32" masterDocId="4D35FFEBFFFDFF886122FFE2FFE18379" masterDocTitle="Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia" masterLastPageNumber="66" masterPageNumber="1" pageNumber="32" updateTime="1698333190807" updateUser="ExternalLinkService">
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<mods:mods id="902340DB06C861803F68A2843168598D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B20D6E8ECD0D51AE5BE39E0C37F3C133">
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<mods:title id="EE3692FED16F9118F815858E735F1AE0">Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia</mods:title>
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<mods:name id="75455C4A6F6D2509F80A823CB9D22EEF" type="personal">
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<mods:namePart id="88A6596C1BD67F478636A12B499A629C">Webster, Mark</mods:namePart>
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<mods:name id="1D10C5864D9B315B3AE276735E9F61B3" type="personal">
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<mods:roleTerm id="A2733D607217F92E509FD7C0FB403762">Author</mods:roleTerm>
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<mods:namePart id="EE1767F7A0BB5CE55EFF573234D81D41">Bohach, Lisa L.</mods:namePart>
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<mods:typeOfResource id="111F1B8D8454A51BA2D486C58D80FA23">text</mods:typeOfResource>
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<mods:title id="044E4D5A32F111F399066C77C0A413C1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="371C04F1BF1E0AF9FDF486BFD67A6668">
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<mods:date id="14E1E525313C545026DE6CECE2CDAADB">2014</mods:date>
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<mods:detail id="B147AE32F5463D3D895D471E312B8A17" type="volume">
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<mods:number id="3B028265486719ADA70CE81267F886EF">3824</mods:number>
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<mods:detail id="8924B13A0B7D46569A65EC58738B6887" type="issue">
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<mods:number id="BF936FCE35C915CECD45277DDC0079D3">1</mods:number>
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<mods:classification id="ED0C5831A97D6A13E27231E7A4D90157">journal article</mods:classification>
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<mods:identifier id="98BFA7817FAAE4F6E769B371788251FE" type="CLB-Dataset">36828</mods:identifier>
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<mods:identifier id="0AA2D53B37BF420FCCC2D0C01191829F" type="DOI">10.11646/zootaxa.3824.1.1</mods:identifier>
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<mods:identifier id="F7A152E49FE307AEA579EC877E8AA3B9" type="GBIF-Dataset">df227efc-13d9-421e-92ed-f9aa417b0cf7</mods:identifier>
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<mods:identifier id="F5EE770E8F840A58EE759B4944A7BD63" type="ISSN">1175-5326</mods:identifier>
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<treatment id="B10C8793FFE2FFA861B5FAE3FB9082D5" ID-DOI="http://doi.org/10.5281/zenodo.6129726" ID-GBIF-Taxon="127668556" ID-Zenodo-Dep="6129726" LSID="urn:lsid:plazi:treatment:B10C8793FFE2FFA861B5FAE3FB9082D5" httpUri="http://treatment.plazi.org/id/B10C8793FFE2FFA861B5FAE3FB9082D5" lastPageId="32" lastPageNumber="32" pageId="31" pageNumber="32">
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<subSubSection id="71BF650EFFE2FF9761B5FAE3FC0486FB" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="391A3685FFE2FF9761B5FAE3FDEE8662" blockId="31.[151,527,1281,1341]" box="[151,527,1281,1307]" pageId="31" pageNumber="32">
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<heading id="625281E9FFE2FF9761B5FAE3FDEE8662" bold="true" box="[151,527,1281,1307]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
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<emphasis id="0BD1EA97FFE2FF9761B5FAE3FDEE8662" bold="true" box="[151,527,1281,1307]" pageId="31" pageNumber="32">
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<taxonomicName id="FEA54D06FFE2FF9761B5FAE3FE188662" box="[151,505,1281,1307]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
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<emphasis id="0BD1EA97FFE2FF9761B5FAE3FE998662" bold="true" box="[151,376,1281,1307]" italics="true" pageId="31" pageNumber="32">Lochmanolenellus</emphasis>
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cf.
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<emphasis id="0BD1EA97FFE2FF976086FAE3FE188662" bold="true" box="[420,505,1281,1307]" italics="true" pageId="31" pageNumber="32">primus</emphasis>
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</taxonomicName>
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1
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="391A3685FFE2FF9761B5FAC6FEDB8645" blockId="31.[151,527,1281,1341]" box="[151,314,1316,1341]" pageId="31" pageNumber="32">Fig. 10.1–10.3</paragraph>
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<paragraph id="391A3685FFE2FF9761B5FA8EFC0486FB" blockId="31.[151,997,1388,1410]" box="[151,997,1388,1410]" pageId="31" pageNumber="32">
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1952
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<taxonomicName id="FEA54D06FFE2FF9761F2FA8FFDA386FB" authority="Lochman" authorityName="Lochman" box="[208,578,1388,1410]" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">
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<emphasis id="0BD1EA97FFE2FF9761F2FA8FFE3D86FB" box="[208,476,1389,1410]" italics="true" pageId="31" pageNumber="32">Wanneria mexicana prima</emphasis>
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Lochman
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</taxonomicName>
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in Cooper
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<emphasis id="0BD1EA97FFE2FF976390FA8FFD3E86FB" box="[690,735,1388,1410]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
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. (part), pl. 18, fig. 1 only.
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</paragraph>
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</subSubSection>
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<subSubSection id="71BF650EFFE2FF9761B5FA4CFAA4852E" pageId="31" pageNumber="32" type="materials_examined">
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<paragraph id="391A3685FFE2FF9761B5FA4CFDB6854A" blockId="31.[151,1436,1454,2019]" pageId="31" pageNumber="32">
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<emphasis id="0BD1EA97FFE2FF9761B5FA4CFEE786BE" bold="true" box="[151,262,1454,1479]" pageId="31" pageNumber="32">Material.</emphasis>
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<collectionCode id="5FB4AE40FFE2FF97602DFA4DFE8186BE" LSID="urn:lsid:biocol.org:col:34871" box="[271,352,1455,1479]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="32">USNM</collectionCode>
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115683 (Fig. 10.2; also figured by Lochman in
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<bibRefCitation id="5D344B74FFE2FF9762A9FA4DFB8D86BE" author="Cooper" box="[907,1132,1454,1479]" pageId="31" pageNumber="32" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
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Cooper
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<emphasis id="0BD1EA97FFE2FF9762C5FA4DFBFB86BF" box="[999,1050,1454,1478]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
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., 1952
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</bibRefCitation>
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, pl. 18, fig. 1) and
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<collectionCode id="5FB4AE40FFE2FF976469FA4DFA7D86BE" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,1455,1479]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="32">USNM</collectionCode>
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115684 (a lot collection of two limestone slabs, both bearing several cephala). Quantitative morphometric and/or qualitative descriptive data were recorded from first-hand examination of
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<collectionCode id="5FB4AE40FFE2FF97652AFA15FBB88576" LSID="urn:lsid:biocol.org:col:34871" box="[1032,1113,1527,1551]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="32">USNM</collectionCode>
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115683 and the three best preserved cephala from
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<collectionCode id="5FB4AE40FFE2FF976087F9F9FE17854A" LSID="urn:lsid:biocol.org:col:34871" box="[421,502,1563,1587]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="32">USNM</collectionCode>
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115684.
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</paragraph>
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<paragraph id="391A3685FFE2FF9761E5F9DCFAA4852E" blockId="31.[151,1436,1454,2019]" box="[199,1349,1598,1623]" pageId="31" pageNumber="32">
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<emphasis id="0BD1EA97FFE2FF9761E5F9DCFEB8852E" bold="true" box="[199,345,1598,1623]" pageId="31" pageNumber="32">Occurrence.</emphasis>
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<collectingCountry id="41B27615FFE2FF976042F9DDFE32852E" box="[352,467,1599,1623]" name="Mexico" pageId="31" pageNumber="32">MEXICO</collectingCountry>
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:
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<emphasis id="0BD1EA97FFE2FF9760FFF9DDFD97852F" box="[477,630,1599,1622]" italics="true" pageId="31" pageNumber="32">Sonora State:</emphasis>
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<collectionCode id="5FB4AE40FFE2FF97635EF9DDFD2C852E" LSID="urn:lsid:biocol.org:col:34871" box="[636,717,1599,1623]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="32">USNM</collectionCode>
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locality 801c. See
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<taxonomicName id="FEA54D06FFE2FF976281F9DCFB27852F" box="[931,1222,1598,1622]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
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<emphasis id="0BD1EA97FFE2FF976281F9DCFB27852F" box="[931,1222,1598,1622]" italics="true" pageId="31" pageNumber="32">Lochmanolenellus primus</emphasis>
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</taxonomicName>
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for details.
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</paragraph>
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</subSubSection>
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<subSubSection id="71BF650EFFE2FFA861E5F980FB9082D5" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="discussion">
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<paragraph id="391A3685FFE2FFA861E5F980FB9082D5" blockId="31.[151,1436,1454,2019]" lastBlockId="32.[151,1436,151,429]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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<emphasis id="0BD1EA97FFE2FF9761E5F980FEAD8502" bold="true" box="[199,332,1634,1659]" pageId="31" pageNumber="32">Discussion.</emphasis>
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USNM 115683 and USNM 115684 from the Caborca area were listed as
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<typeStatus id="E61E8827FFE2FF9765B3F981FB1D8502" box="[1169,1276,1635,1659]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
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of “
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<taxonomicName id="FEA54D06FFE2FF976412F981FEB185E7" class="Trilobita" family="Holmiidae" genus="Wanneria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="mexicana" subSpecies="prima">
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<emphasis id="0BD1EA97FFE2FF976412F981FEB185E7" italics="true" pageId="31" pageNumber="32">Wanneria mexicana prima</emphasis>
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</taxonomicName>
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” by Lochman (in
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<bibRefCitation id="5D344B74FFE2FF976309F965FCF185E6" author="Cooper" box="[555,784,1670,1695]" pageId="31" pageNumber="32" refString="Cooper, G. A., Arellano, A. R. V., Johnson, J. H., Okulitch, V. J., Stoyanow, A. & Lochman, C. (1952) Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico. Smithsonian Miscellaneous Collections, 119 (1), 1 - 184." type="journal article" year="1952">
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Cooper
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<emphasis id="0BD1EA97FFE2FF9763AAF965FD5C85E7" box="[648,701,1670,1694]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
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., 1952
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</bibRefCitation>
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, p. 97). All these specimens are very worn and mostly/ entirely exfoliated, and none would be expected to preserve the prosopon or the lateral swellings on the glabella characteristic of the genus (although USNM 115683 does show traces of the reticulate prosopon in the anterior cephalic border furrow [Fig. 10.3]). However, all cephala in lot collection USNM 115684 bear genal and intergenal spines that flare quite strongly laterally, as do at least the genal spines of USNM 115683 (the intergenal spines of this specimen are not preserved; Fig. 10.2). USNM 115683 and at least one specimen within lot collection USNM 115684 (Fig. 10.1) seem to possess an extraocular platform (the other specimens within the lot collection are too poorly preserved to determine the state of this character). Such traits are indicative of
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<taxonomicName id="FEA54D06FFE2FF9765B3F861FABF84E2" box="[1169,1374,1923,1947]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="0BD1EA97FFE2FF9765B3F861FABF84E2" box="[1169,1374,1923,1947]" italics="true" pageId="31" pageNumber="32">Lochmanolenellus</emphasis>
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</taxonomicName>
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. The placement of the genal spine bases and the orientation of the distal portion of the posterior cephalic margin of these specimens are similar to
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<taxonomicName id="FEA54D06FFE2FF9760E5F829FD0E849B" box="[455,751,1995,2019]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="primus">
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<emphasis id="0BD1EA97FFE2FF9760E5F829FD0E849B" box="[455,751,1995,2019]" italics="true" pageId="31" pageNumber="32">Lochmanolenellus primus</emphasis>
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</taxonomicName>
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, with which the specimens co-occur. The best preserved specimens (Fig. 10.1, 10.2) differ from the
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<typeStatus id="E61E8827FFDDFFA863BDFF75FCE083C9" box="[671,769,151,176]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="FEA54D06FFDDFFA86213FF7BFC5183D6" box="[817,944,152,176]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="primus">
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<emphasis id="0BD1EA97FFDDFFA86213FF7BFC5183D6" box="[817,944,152,176]" italics="true" pageId="32" pageNumber="33">Lo. primus</emphasis>
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</taxonomicName>
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in possessing a proportionally wider (tr.) extraocular area with a more subdued extraocular platform, a less strongly dorsally rounded anterior cephalic border, and proportionally shorter ocular lobes (with posterior tips located transversely opposite the lateral margins of L2 rather than S1). However, these two specimens are more than twice as large as the
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<typeStatus id="E61E8827FFDDFFA865B4FEE6FB198264" box="[1174,1272,260,285]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
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of
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<taxonomicName id="FEA54D06FFDDFFA86403FEE7FA7D8265" box="[1313,1436,261,284]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="primus">
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<emphasis id="0BD1EA97FFDDFFA86403FEE7FA7D8265" box="[1313,1436,261,284]" italics="true" pageId="32" pageNumber="33">Lo. primus</emphasis>
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</taxonomicName>
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(sagittal cephalic lengths approximately
|
||
<quantity id="FE5D9B60FFDDFFA8637CFECAFD238246" box="[606,706,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="32" pageNumber="33" unit="mm" value="19.8">19.8 mm</quantity>
|
||
and
|
||
<quantity id="FE5D9B60FFDDFFA863DAFECAFCBF8246" box="[760,862,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="32" pageNumber="33" unit="mm" value="22.4">22.4 mm</quantity>
|
||
, respectively, versus approximately
|
||
<quantity id="FE5D9B60FFDDFFA865D7FECAFAAA8246" box="[1269,1355,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="32" pageNumber="33" unit="mm" value="9.7">9.7 mm</quantity>
|
||
for the
|
||
<typeStatus id="E61E8827FFDDFFA861B5FEAEFF1C821C" box="[151,253,332,357]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
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), and the differences might be attributable to ontogenetic allometry given the known patterns of shape change within other species of the genus (see following).
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<collectingCountry id="41B27615FFDDFFA86265FE92FC8782F1" box="[839,870,368,392]" name="American Samoa" pageId="32" pageNumber="33">As</collectingCountry>
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a result of these morphological differences and ambiguities, all these specimens are herein conservatively identified as
|
||
<taxonomicName id="FEA54D06FFDDFFA8629AFE77FBB582D5" box="[952,1108,404,429]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="primus">
|
||
<emphasis id="0BD1EA97FFDDFFA8629AFE77FC3C82D5" box="[952,989,405,428]" italics="true" pageId="32" pageNumber="33">Lo.</emphasis>
|
||
cf.
|
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<emphasis id="0BD1EA97FFDDFFA86525FE77FBB582D5" box="[1031,1108,405,428]" italics="true" pageId="32" pageNumber="33">primus</emphasis>
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</taxonomicName>
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1.
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</paragraph>
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</subSubSection>
|
||
</treatment>
|
||
</document> |