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<mods:title id="D7A5BDB6A9651CCE5011AE95DABBC2C4">Tortonian teleost otoliths from northern Italy: taxonomic synthesis and stratigraphic significance</mods:title>
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<mods:namePart id="DF9D3A6850DBA57FD91D497341C4E9BF">Lin, Chien-Hsiang</mods:namePart>
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<mods:namePart id="026B57A8278F5572066C496E70DF4786">Brzobohatý, Rostislav</mods:namePart>
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<treatment id="61599612947BCE2B7A29A538FE919738" ID-DOI="http://doi.org/10.5281/zenodo.3848341" ID-GBIF-Taxon="164159945" ID-Zenodo-Dep="3848341" LSID="urn:lsid:plazi:treatment:61599612947BCE2B7A29A538FE919738" httpUri="http://treatment.plazi.org/id/61599612947BCE2B7A29A538FE919738" lastPageNumber="30" pageId="29" pageNumber="30">
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<taxonomicName id="2EF05C87947BCE2B7A29A538FC7192DB" ID-CoL="45XVM" authority="Johnson, 1865" authorityName="Johnson" authorityYear="1865" box="[614,973,367,394]" class="Actinopterygii" family="Gempylidae" genus="Nealotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="tripes">
<emphasis id="DB84FB16947BCE2B7A29A538FCA092DB" bold="true" box="[614,796,367,394]" italics="true" pageId="29" pageNumber="30">Nealotus tripes</emphasis>
Johnson, 1865
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An otolith, named
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<emphasis id="DB84FB16947BCE2B79D8A5F8FD759298" box="[407,713,431,457]" italics="true" pageId="29" pageNumber="30">Prometichthys prometheus</emphasis>
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by
<bibRefCitation id="8D615AF5947BCE2B7ABAA5F8FC13929B" author="Lin C. - H. &amp; Girone A. &amp; Nolf D." box="[757,943,431,458]" pageId="29" pageNumber="30" pagination="249 - 261" refId="ref22326" refString="Lin C. - H., Girone A. &amp; Nolf D. 2015. Tortonian fish otoliths from turbiditic deposits in northern Italy: taxonomic and stratigraphic significance. Geobios 48: 249 - 261. https: // doi. org / 10.1016 / j. geobios. 2015.03.003" type="journal article" year="2015">
Lin
<emphasis id="DB84FB16947BCE2B7B6AA5E7FCE49298" box="[805,856,431,457]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. (2015
</bibRefCitation>
: fig. 7(1)), is also assigned to
<taxonomicName id="2EF05C87947BCE2B7D44A5E7FAD29298" authorityName="Johnson" authorityYear="1865" box="[1291,1390,431,457]" class="Actinopterygii" family="Gempylidae" genus="Nealotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="tripes">
<emphasis id="DB84FB16947BCE2B7D44A5E7FAA29298" box="[1291,1310,432,457]" italics="true" pageId="29" pageNumber="30">N</emphasis>
.
<emphasis id="DB84FB16947BCE2B7D62A5E7FAD29298" box="[1325,1390,432,457]" italics="true" pageId="29" pageNumber="30">tripes</emphasis>
</taxonomicName>
: three additional specimens from Mondovi (one figured on
<figureCitation id="71CB3B81947BCE2B7BC2A583FC4B92BF" box="[909,1015,468,494]" captionStart="Fig" captionStartId="28.[189,232,1788,1814]" captionTargetBox="[189,1398,612,1746]" captionTargetId="figure@28.[189,1398,612,1746]" captionTargetPageId="28" captionText="Fig. 12. Tortonian fish otoliths from northern Italy. AC. “Gobius” bicornutus (Lin, Girone &amp; Nolf, 2015), SantAgata Fossili (IRSNB P 9836P 9838). D. Aphanopus carbo Lowe, 1839, Costa Vescovato (IRSNB P 9839). E. Nealotus tripes Johnson, 1865, Mondovi, Madonna della Neve (IRSNB P 9840). FG. Arnoglossus kokeni (Bassoli &amp; Schubert, 1906), Torrente Stirone (IRSNB P 9841P 9842). HI. Gobiidae indet., SantAgata Fossili (IRSNB P 9843P 9844). JK. Microchirus aff. variegatus (Donovan, 1808), Torrente Stirone (IRSNB P 98459846). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829329" httpUri="https://zenodo.org/record/3829329/files/figure.png" pageId="29" pageNumber="30">Fig. 12E</figureCitation>
) present a very similar spindle shape, and seem to be nearest to those of the Recent
<taxonomicName id="2EF05C87947BCE2B7B0CA5ADFC169142" authorityName="Johnson" authorityYear="1865" box="[835,938,505,531]" class="Actinopterygii" family="Gempylidae" genus="Nealotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="tripes">
<emphasis id="DB84FB16947BCE2B7B0CA5ADFCEA9142" box="[835,854,506,531]" italics="true" pageId="29" pageNumber="30">N</emphasis>
.
<emphasis id="DB84FB16947BCE2B7B26A5ADFC169142" box="[873,938,506,531]" italics="true" pageId="29" pageNumber="30">tripes</emphasis>
</taxonomicName>
(see
<bibRefCitation id="8D615AF5947BCE2B7BA1A5AEFBD09142" author="Nolf D." box="[1006,1132,505,531]" pageId="29" pageNumber="30" refId="ref22613" refString="Nolf D. 2013. The Diversity of Fish Otoliths, Past and Present. Royal Belgian Institute of Natural Sciences, Brussels." type="book" year="2013">Nolf 2013</bibRefCitation>
: pl. 327). The otoliths of
<taxonomicName id="2EF05C87947BCE2B78ABA649FDF69169" authority="(Cuvier, 1832)" baseAuthorityName="Cuvier" baseAuthorityYear="1832" box="[228,586,541,568]" class="Actinopterygii" family="Gempylidae" genus="Prometichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="prometheus">
<emphasis id="DB84FB16947BCE2B78ABA649FE319169" box="[228,397,542,568]" italics="true" pageId="29" pageNumber="30">P. prometheus</emphasis>
(Cuvier, 1832)
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also show a reasonable resemblance, but the sulcus is straighter in
<taxonomicName id="2EF05C87947BCE2B7891A614FEF8910D" authorityName="Johnson" authorityYear="1865" box="[222,324,579,605]" class="Actinopterygii" family="Gempylidae" genus="Nealotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="tripes">
<emphasis id="DB84FB16947BCE2B7891A614FF4D910D" box="[222,241,579,604]" italics="true" pageId="29" pageNumber="30">N</emphasis>
.
<emphasis id="DB84FB16947BCE2B794CA614FEF8910D" box="[259,324,579,604]" italics="true" pageId="29" pageNumber="30">tripes</emphasis>
</taxonomicName>
, and the rostrum in
<taxonomicName id="2EF05C87947BCE2B7A71A614FD5B910D" baseAuthorityName="Cuvier" baseAuthorityYear="1832" box="[574,743,578,605]" class="Actinopterygii" family="Gempylidae" genus="Prometichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="prometheus">
<emphasis id="DB84FB16947BCE2B7A71A614FDEC910D" box="[574,592,579,604]" italics="true" pageId="29" pageNumber="30">P</emphasis>
.
<emphasis id="DB84FB16947BCE2B7A2DA615FD5B910D" box="[610,743,578,604]" italics="true" pageId="29" pageNumber="30">prometheus</emphasis>
</taxonomicName>
is more prominent with a conspicuous excisura notch.
</paragraph>
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<emphasis id="DB84FB16947BCE2B78F2A6D4FB8591F1" bold="true" box="[189,1081,643,672]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="2EF05C87947BCE2B78F2A6D4FEDC91CE" box="[189,352,643,671]" class="Actinopterygii" family="Bothidae" genus="Arnoglossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="overview">An overview</taxonomicName>
on the taxonomic composition of investigated localities
</emphasis>
</paragraph>
<paragraph id="E94F2704947BCE2B78F2A6EEFE919738" blockId="29.[189,1399,697,1129]" pageId="29" pageNumber="30">
A list of taxa recognised in this study is given in
<tableCitation id="A47212BF947BCE2B7A90A6EEFC8F9182" box="[735,819,697,723]" captionStart="Table 1" captionStartId="5.[186,255,550,578]" captionTargetId="graphics@5.[175,1927,629,1746]" captionTargetPageId="5" captionText="Table 1 (continued on next pages). List of otolith-based fish taxa from the Tortonian deposits in northern Italy." httpUri="http://table.plazi.org/id/BD8F778C9463CE3378F5A671FA759113" pageId="29" pageNumber="30" tableUuid="BD8F778C9463CE3378F5A671FA759113">Table 1</tableCitation>
(page 6 and following), with all of them illustrated by at least one specimen. The present study, based on more than 5600 otoliths, reveals the presence of at least 109 taxa belonging to 38 families. Of these, 88 taxa were identified at species level. Among all the investigated localities, SantAgata Fossili and Montegibbio are the most diversified at family level, consisting of 20 and 18 families, respectively. The most abundant material came from Mondovi, Madonna della Neve, where more than 1600 specimens were collected. Furthermore, this locality is also characterised by many true oceanic mesopelagic taxa, such as microstomatids, sternoptychids and phosichthyids. At the Torrente Stirone locality, the otolith assemblage is most diversified at species level (51 taxa), of which the most abundant and even dominant taxa are neritic gobiids and congrids. Finally, the assemblages recognised at the Alba, Tanaro, Costa Vescovato, Gallo DAlba, SantAlosio, and Stazzano localities show the typical characteristics of deep-water taxa, with a near-absence of neritic elements.
</paragraph>
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