294 lines
31 KiB
XML
294 lines
31 KiB
XML
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<mods:title id="0E0713D7C37915584FDF7C2B222FA571">Phylogeny of the genus Yumtaax Boucher (Coleoptera, Passalidae, Proculini): Taxonomic and evolutionary implications with descriptions of three new species</mods:title>
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<mods:namePart id="BF991F5571D70028D802C28C89EC719C">Beza-Beza, Cristian Fernando</mods:namePart>
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<mods:namePart id="D3C3351945BCFEDCC627A027DE0E8F1E">Beck, James</mods:namePart>
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<mods:namePart id="39995D42B458EDC00A58227F02FDA14D">Reyes-Castillo, Pedro</mods:namePart>
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<mods:namePart id="3DBDBE8D8A1BD47936592477EF6CCFD8">Jameson, Mary Liz</mods:namePart>
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<mods:date id="49EE8841A80F7585049A65A10BCDCF50">2017</mods:date>
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<mods:identifier id="6F6444F3AFBFFC11859C36979399FD43" type="DOI">http://dx.doi.org/10.3897/zookeys.667.10716</mods:identifier>
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<treatment id="421B3BDF5260042DD19716C396F96837" ID-GBIF-Taxon="128345062" LSID="urn:lsid:plazi:treatment:421B3BDF5260042DD19716C396F96837" httpUri="http://treatment.plazi.org/id/421B3BDF5260042DD19716C396F96837" lastPageId="21" lastPageNumber="116" pageId="18" pageNumber="113">
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<subSubSection id="A5EF4FC9E44A19E577E428A13B53CEFF" pageId="18" pageNumber="113" type="nomenclature">
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<paragraph id="9E1015BAD47E817567B6CC1CB03ABCE6" pageId="18" pageNumber="113">
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<taxonomicName id="72008EB1C3BD4693BC8BF0961632EE6F" ID-CoL="5CTFF" authority="Truqui, 1857" authorityName="Truqui" authorityYear="1857" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax laticornis" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="laticornis">Yumtaax laticornis (Truqui, 1857)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="3FC0EF0CA888EC3ACA14DF662A7F8433" pageId="18" pageNumber="113" type="reference_group">
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<paragraph id="65108193F029EFF812CE7BDDFF05AF00" pageId="18" pageNumber="113">
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<taxonomicName id="54F4C03FFB94CD6B1F390DD6E8653E7F" class="Insecta" family="Passalidae" genus="Passalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passalus laticornis" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="laticornis">Passalus laticornis</taxonomicName>
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Truqui, 1857: 262, 316.
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</paragraph>
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<paragraph id="2D4A993B7A3BC33783A4E4AABB4EC17D" pageId="18" pageNumber="113">
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<taxonomicName id="84B015C119A03A557A72F524936BB5AB" class="Insecta" family="Passalidae" genus="Pseudacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudacanthus laticornis" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="laticornis">Pseudacanthus laticornis</taxonomicName>
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(Truqui) [comb. n. by
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<bibRefCitation id="99CF775A19A84948F1A49D94BC6787BF" author="Kaup, JJ" journalOrPublisher="Berliner Entomologische zeitschrift," pageId="29" pageNumber="124" pagination="1 - 125" title="Monographie der Passaliden." url="https://doi.org/10.1002/mmnd.18710150306" volume="15" year="1871">Kaup 1871</bibRefCitation>
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||
: 72, 74].
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</paragraph>
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<paragraph id="13DD77FBAE6CB4D73178317A7E68A53D" pageId="18" pageNumber="113">
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<taxonomicName id="ED373C8BD074A4DAB333A0ACB5992B12" class="Insecta" family="Passalidae" genus="Petrejoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Petrejoides laticornis" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="laticornis">Petrejoides laticornis</taxonomicName>
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(Truqui) [comb. n. by
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<bibRefCitation id="1EABCA33B808DCE82354F1699421EA8D" pageId="18" pageNumber="113">Reyes-Castillo 1970</bibRefCitation>
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||
: 125].
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</paragraph>
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<paragraph id="9B05F308D4666EB0BA5E3CA93EC7A2D8" pageId="18" pageNumber="113">
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<taxonomicName id="812D238F1AFD6279DBC17858EB3E63E7" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax laticornis" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="laticornis">Yumtaax laticornis</taxonomicName>
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(Truqui) [comb. n. by
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<bibRefCitation id="EF9FE6A3DFC2D3AAC767A15585558675" author="Boucher, S" journalOrPublisher="Annale de la Societe Entomologique de France" pageId="28" pageNumber="123" pagination="237 - 604" title="Evolution et phylogenie des coleopteres Passalidae (Scarabaeoidea)." url="https://doi.org/10.1080/00379271.2005.10697444" volume="41" year="2006">Boucher 2006</bibRefCitation>
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: 348].
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</paragraph>
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</subSubSection>
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<subSubSection id="A40D1DB96B51C276CE25A0E65096662F" pageId="18" pageNumber="113" type="materials_examined">
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<paragraph id="D3C59E46FF8230504E9EB7D2EB50C037" pageId="18" pageNumber="113">Material examined.</paragraph>
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<paragraph id="E2AADAE35E71BCB9AB4FC0BABACD78A1" pageId="18" pageNumber="113">Holotype and 31 non-type specimens.</paragraph>
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<paragraph id="74F01C25B92452431151C7D748FA79F7" pageId="18" pageNumber="113">
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Holotype ♂. MEXICO: Jacale, 1708 (
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<normalizedToken id="071F42D5F2A3656B065112F806332826" originalValue="Sallé">Salle</normalizedToken>
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) (BMNH).
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</paragraph>
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<paragraph id="CD5C5B6DAEC1C1838F11A720C23C949F" pageId="18" pageNumber="113">
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Non-type specimens (31 total): 2 ♂, 20 ♀, 9 unknown. MEXICO: Veracruz, Calcahualco, Tecuanapa, bosque
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<normalizedToken id="D87F27994300F99D2CC582C3F91B09AA" originalValue="mesófilo">mesofilo</normalizedToken>
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, alt. 2200 m, VI 1992 (
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<normalizedToken id="D215FD4ADE9C8E43E9D6D0A434B53F6A" originalValue="Capistrán">Capistran</normalizedToken>
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, Delgado) (IEXA; CFBB).
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</paragraph>
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</subSubSection>
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<subSubSection id="6ABEB8A6BB3E412E8ECC35D1CF4743E4" pageId="18" pageNumber="113" type="diagnosis">
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<paragraph id="3DB10BD7FE9F64512DD9AC5CD3F909A8" pageId="18" pageNumber="113">Diagnosis.</paragraph>
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<paragraph id="0FA053AB85B157B129A51DB7DC3A51B2" pageId="18" pageNumber="113">
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<taxonomicName id="F40F01CC7D4D77515B8A2710BFE20DA7" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax laticornis" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="laticornis">Yumtaax laticornis</taxonomicName>
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is a large (24.5-33.0 mm) brachypterous species and is part of the
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<taxonomicName id="59598CF1874FC82A5EB6511E6AA56280" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax laticornis" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="species" species="laticornis">Yumtaax laticornis</taxonomicName>
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clade (Fig. 4). This species is diagnosed by the following character combination: the clypeus is vertical (shared with
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<taxonomicName id="B2506EDD6211CE0FD657389DE04ECC6B" lsidName="Y. veracrucensis" pageId="18" pageNumber="113" rank="species" species="veracrucensis">Y. veracrucensis</taxonomicName>
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,
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<taxonomicName id="30B14D405BE36CA83DC129BBC380CF0E" lsidName="Y. cameliae" pageId="18" pageNumber="113" rank="species" species="cameliae">Y. cameliae</taxonomicName>
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; inclined in other members of
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<taxonomicName id="9EF265AB3FEA1D824C2FB07FD85DA4FC" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="genus">Yumtaax</taxonomicName>
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) and the anterior border is straight (shared with other members of
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<taxonomicName id="DA3D36F27D3EBF17F67A7D4FFF222097" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="genus">Yumtaax</taxonomicName>
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except for
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<taxonomicName id="09BAA7083F1F441E8DC1695C6CDC803F" lsidName="Y. recticornis" pageId="18" pageNumber="113" rank="species" species="recticornis">Y. recticornis</taxonomicName>
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and
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<taxonomicName id="D4E31E2A534E9F83BE25E57BE5E8A1DF" lsidName="Y. olmecae" pageId="18" pageNumber="113" rank="species" species="olmecae">Y. olmecae</taxonomicName>
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with concave anterior border of clypeus); mesofrontal structure (MFS) of the
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<normalizedToken id="F7460315856A3C430860DBF01C60125C" originalValue="“falsus”">"falsus"</normalizedToken>
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type (see
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<bibRefCitation id="49DBEB23F15B2B7A243871CEF0427931" pageId="18" pageNumber="113">Reyes-Castillo 1970</bibRefCitation>
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) (shared with all members of
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<taxonomicName id="7745B7AEAF1D8CA54697D0AC4BC4CCB2" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="genus">Yumtaax</taxonomicName>
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except
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<taxonomicName id="CC898948F8253350B5AEBB75E8E8F425" lsidName="Y. cameliae" pageId="18" pageNumber="113" rank="species" species="cameliae">Y. cameliae</taxonomicName>
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which has the MFS of the
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<normalizedToken id="4FBB2ED105E543F124DD10C6327EC7D1" originalValue="“striatopunctatus”">"striatopunctatus"</normalizedToken>
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type), with the central tooth largely free (shared with
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<taxonomicName id="490BFE70916A73EAA9B8F52C747DF86A" lsidName="Y. recticornis" pageId="18" pageNumber="113" rank="species" species="recticornis">Y. recticornis</taxonomicName>
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,
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<taxonomicName id="47CC0FF7F0B8F294D58DBA21E7042F7A" lsidName="Y. veracrucensis" pageId="18" pageNumber="113" rank="species" species="veracrucensis">Y. veracrucensis</taxonomicName>
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,
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<taxonomicName id="994411FF00474E5771010F92F1FF99A1" lsidName="Y. cameliae" pageId="18" pageNumber="113" rank="species" species="cameliae">Y. cameliae</taxonomicName>
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,
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<taxonomicName id="EC535BFB7EB246DAF656F0267BED0314" lsidName="Y. mazatecus" pageId="18" pageNumber="113" rank="species" species="mazatecus">Y. mazatecus</taxonomicName>
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; fused with frontal ridges in
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<taxonomicName id="E24AAD7B509973BE10210C13C433B603" lsidName="Y. jimenezi" pageId="18" pageNumber="113" rank="species" species="jimenezi">Y. jimenezi</taxonomicName>
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,
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<taxonomicName id="73950E390923677EC859A903366404B4" lsidName="Y. imbellis" pageId="18" pageNumber="113" rank="species" species="imbellis">Y. imbellis</taxonomicName>
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||
,
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<taxonomicName id="B804F474EC66B8DC6B8CCCBF8FF0869F" lsidName="Y. nebulosus" pageId="18" pageNumber="113" rank="species" species="nebulosus">Y. nebulosus</taxonomicName>
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,
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<taxonomicName id="65BABC7FEEB4ADC9D3341D9A3B33238F" lsidName="Y. olmecae" pageId="18" pageNumber="113" rank="species" species="olmecae">Y. olmecae</taxonomicName>
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), directed anteriorly (shared with
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<taxonomicName id="99CBBCA573CA82D40A3223A702EF47C6" lsidName="Y. veracrucensis" pageId="18" pageNumber="113" rank="species" species="veracrucensis">Y. veracrucensis</taxonomicName>
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; directed dorsally in
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<taxonomicName id="BD688045D0E38A1D7E6BD5AF8121DC2D" lsidName="Y. recticornis" pageId="18" pageNumber="113" rank="species" species="recticornis">Y. recticornis</taxonomicName>
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||
,
|
||
<taxonomicName id="9AC7F538D4AF9CCF32909CE7EC42DD76" lsidName="Y. imbellis" pageId="18" pageNumber="113" rank="species" species="imbellis">Y. imbellis</taxonomicName>
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||
,
|
||
<taxonomicName id="F49D9678D8339A18A2FE3A7425E9AD57" lsidName="Y. nebulosus" pageId="18" pageNumber="113" rank="species" species="nebulosus">Y. nebulosus</taxonomicName>
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||
,
|
||
<taxonomicName id="3343DF0D1B81EEA950CE39A72DCD78BB" lsidName="Y. olmecae" pageId="18" pageNumber="113" rank="species" species="olmecae">Y. olmecae</taxonomicName>
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||
; directed dorsally and anteriorly in
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||
<taxonomicName id="FA83903D71655FBB25FFDA0A5C314DFA" lsidName="Y. jimenezi" pageId="18" pageNumber="113" rank="species" species="jimenezi">Y. jimenezi</taxonomicName>
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||
,
|
||
<taxonomicName id="BB0096007CE4949425CD9C2406B5BE1A" lsidName="Y. mazatecus" pageId="18" pageNumber="113" rank="species" species="mazatecus">Y. mazatecus</taxonomicName>
|
||
; elevated in the posterior half bending abruptly forward in the anterior half in
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||
<taxonomicName id="924994B803AF19931C26DC0B7280240B" lsidName="Y. cameliae" pageId="18" pageNumber="113" rank="species" species="cameliae">Y. cameliae</taxonomicName>
|
||
), and reaching the frontoclypeal suture (shared with
|
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<taxonomicName id="0C8E6B2E186CD7BE3F4424B536DDECF3" lsidName="Y. veracrucensis" pageId="18" pageNumber="113" rank="species" species="veracrucensis">Y. veracrucensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="9AF733D11FEE3BD9F66C270E73307C6F" lsidName="Y. cameliae" pageId="18" pageNumber="113" rank="species" species="cameliae">Y. cameliae</taxonomicName>
|
||
; not reaching the clypeus in other members of
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<taxonomicName id="D1A2903DF44E87F738D17FD27B5787E9" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="genus">Yumtaax</taxonomicName>
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); eyes are strongly reduced (shared with
|
||
<taxonomicName id="CB8A62D7B2F6A8899637177F2CCD1372" lsidName="Y. mazatecus" pageId="18" pageNumber="113" rank="species" species="mazatecus">Y. mazatecus</taxonomicName>
|
||
; eyes large in
|
||
<taxonomicName id="3AB77C284C09335BA0A273AE666F8CBE" lsidName="Y. recticornis" pageId="18" pageNumber="113" rank="species" species="recticornis">Y. recticornis</taxonomicName>
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||
,
|
||
<taxonomicName id="78C751E82819957E3D8A072C22DDD661" lsidName="Y. imbellis" pageId="18" pageNumber="113" rank="species" species="imbellis">Y. imbellis</taxonomicName>
|
||
; eyes moderately reduced in
|
||
<taxonomicName id="E4B108848FB236112AF3F029EBC74C4F" lsidName="Y. veracrucensis" pageId="18" pageNumber="113" rank="species" species="veracrucensis">Y. veracrucensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4005B7B70C81AF1CD4CC708533514551" lsidName="Y. cameliae" pageId="18" pageNumber="113" rank="species" species="cameliae">Y. cameliae</taxonomicName>
|
||
,
|
||
<taxonomicName id="062F0E3D6E7B4E0E2A9D655B97FE0EC3" lsidName="Y. jimenezi" pageId="18" pageNumber="113" rank="species" species="jimenezi">Y. jimenezi</taxonomicName>
|
||
,
|
||
<taxonomicName id="DF6BB3C582E472C6E15E8144B2FD934E" lsidName="Y. nebulosus" pageId="18" pageNumber="113" rank="species" species="nebulosus">Y. nebulosus</taxonomicName>
|
||
,
|
||
<taxonomicName id="8254742B003B881ED467B46C53D7C7A9" lsidName="Y. olmecae" pageId="18" pageNumber="113" rank="species" species="olmecae">Y. olmecae</taxonomicName>
|
||
); and the scutellum is punctate (smooth in other members of
|
||
<taxonomicName id="97137EF36A0990FA78936D5E4C8C3862" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax" order="Coleoptera" pageId="18" pageNumber="113" phylum="Arthropoda" rank="genus">Yumtaax</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ABF6B5974F0DAB9CAC56CF2CDD5CF59B" pageId="18" pageNumber="113" type="description">
|
||
<paragraph id="6F3F460BEF9584DC94779D9F310653A0" pageId="18" pageNumber="113">Dimensions</paragraph>
|
||
<paragraph id="7BD1E4F9953B485D38FB9BD25B9FD934" pageId="18" pageNumber="113">
|
||
(mm) (n = 19). Total length 24.5-33.0 (
|
||
<normalizedToken id="6933ECDAF3DCE14AF3C074293B7C39C4" originalValue="χ">x</normalizedToken>
|
||
= 29.5); elytral length 14.0-17.5 (
|
||
<normalizedToken id="54054B6BFF9B8BEC84886669F3B67528" originalValue="χ">x</normalizedToken>
|
||
= 16.5); pronotal length 6.0-9.0 (
|
||
<normalizedToken id="F6A9654B356FF7625FB30BA10EE45588" originalValue="χ">x</normalizedToken>
|
||
= 8.0); pronotal width 8.0-11.0 (
|
||
<normalizedToken id="F612CE43FD5E286D3635568559B955C3" originalValue="χ">x</normalizedToken>
|
||
= 10.0); humeral width 7.0-10.0 (
|
||
<normalizedToken id="58025AC55099D4D2BC0E62416B99660F" originalValue="χ">x</normalizedToken>
|
||
= 9.0).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6BE8167893DB81DF3717174F5F524AE3" lastPageId="20" lastPageNumber="115" pageId="18" pageNumber="113" type="redescription">
|
||
<paragraph id="A556C6CD887AF6F9B0CAAD06ACFC49F8" pageId="18" pageNumber="113">Redescription of holotype</paragraph>
|
||
<paragraph id="C556D23823D442CA08AEB5EEFCD91019" lastPageId="20" lastPageNumber="115" pageId="18" pageNumber="113">
|
||
(Fig. 7). Head (Fig. 7C). Labrum: anterior border concave, dorsal surface smooth and glabrous medially, punctate and setose apicolaterally, apically, and basally; anterior edge excavated. Clypeus: vertical, rectangular, flat, shiny, and smooth. Frontoclypeal suture: straight, and shiny; external tubercles
|
||
<pageBreakToken id="EAA55EA5F36CFDCE80945E0CF2369791" pageId="19" pageNumber="114" start="start">rounded</pageBreakToken>
|
||
weak, directed anteriorly. Frontal area: vertical, flat, smooth and shiny, frontal ridge absent. Frontal fossae: punctate and glabrous. Mesofrontal structure (MFS): of the
|
||
<normalizedToken id="CC896DC7AF2D0E4063F81110461C2C7B" originalValue="“falsus”">"falsus"</normalizedToken>
|
||
type (see
|
||
<bibRefCitation id="F6DE4848D227C2D7F3E7C6DB532FA13A" pageId="19" pageNumber="114">Reyes-Castillo 1970</bibRefCitation>
|
||
); base subparallel and narrower than
|
||
<normalizedToken id="B65A608C8FEDCC1133DA65581947EAF9" originalValue="MFS’">MFS'</normalizedToken>
|
||
lateral ridge; center horn long with apex acute, largely free and directed anteriorly (Fig. 7D), surpassing posterior margin of clypeus, dorsally without micro-punctures; base of center horn wide narrowing gradually until apex; dorsal fossa absent at the base of MFS. Occipital fossa: shallow posteriorly and deeper laterally, not connected with the frontal fossae. Posterior occipital sulcus concave. Supraorbital ridge: bituberculate, tubercles of similar size; posterior half of supraorbital ridge bifurcated. Canthus: with apex rounded, covering more than 1/3 of the eye, expanded distally. Eyes: strongly
|
||
<pageBreakToken id="201C2212E95FA22426C8D619DD624908" pageId="20" pageNumber="115" start="start">reduced</pageBreakToken>
|
||
(distal edge of the eye shorter than the distal edge of the canthus), width = 0.4 mm (each eye). Head width (between posterior tubercles of the supraorbital ridge) = 5.0 mm. Ratio of sums of both eyes widths/total head width = 0.16; postocular area punctate and setose. Ligula: tridentate, central tooth surpassing apex of lateral teeth, lateral teeth rounded; setose punctures present in discal area; posterior border straight. Mentum: lateral lobes rounded and wide, with setose punctures. Basomedial portion protruding ventrally; anterior border at middle straight; basal fossae present, with setose punctures. Hypostomal process: with lateral depression; separated from mentum by a distance larger than the wide of the anterior width of hypostomal process. Infraocular ridge (e.g., Fig. 3): short and wide anteriorly, narrow posteriorly. Mandible: with 3 apical teeth; internal tooth of left mandible bidentate; dorsal tooth occupies more than half length of the mandible. Pronotum: anterior angles rounded. Anterior fossae of marginal sulcus punctate. Lateral fossae without punctures. Marginal groove with punctures. Prosternum: opaque. Prosternellum with anterior half and lateral edges opaque and posterior half and middle area shiny. Scutellum: punctate and glabrous. Mesosternum: with anterior-lateral areas opaque. Metasternum: with setae anterolaterally, lacking punctures in lateral margins of the metasternal disc. Lateral fossae wide glabrous posteriorly with setose punctures anteriorly. Elytra: anterior border straight. Meeting point of striae 1-10 (see
|
||
<bibRefCitation id="6BDD8B176B83D3A962F691FA00B31A3F" pageId="20" pageNumber="115">Reyes-Castillo 1970</bibRefCitation>
|
||
) with one line of punctures. Wings: reduced. Legs (Fig. 7E): femur I with longitudinal anteroventral groove strongly developed, reaching the distal end of the femur, posterioventral half pubescent; setae short, dense, reddish. Abdomen: last sternite with marginal groove complete (Fig. 7B). Aedeagus (Fig. 7F, G, and H) (Description based on non-type material): dorsal view phallus globose (wider than long). Ventral view lateral edges of the phallus surpassing the laterodistal edges of the parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EC2AD9CB713BB96B5B7E2BB80912D446" pageId="20" pageNumber="115" type="variation">
|
||
<paragraph id="D8085E2AA20A830D90221992803E5123" pageId="20" pageNumber="115">Variation.</paragraph>
|
||
<paragraph id="D823BBC8917B037BA991E125D0923E99" pageId="20" pageNumber="115">The non-type material differs from the holotype in the following characters: internal tubercles obsolete to strongly developed; frontal ridges obsolete to strongly developed; frontal area glabrous to sparsely setose; ratio of eyes versus head width varies from 0.13-0.23; pronotum laterally with or without strong punctures, even at the individual level (right vs left side of the pronotum); prosternelum completely opaque or opaque and shiny.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="20C41CB9F9C989BB875B733171D59AE7" pageId="20" pageNumber="115" type="distribution">
|
||
<paragraph id="320C3E693D18CDDDF8AC39B7290F2633" pageId="20" pageNumber="115">Distribution.</paragraph>
|
||
<paragraph id="57709953D625431958CC398B245D3990" pageId="20" pageNumber="115">
|
||
This species is known from cloud forest (bosque
|
||
<normalizedToken id="1BABAB137E5D61854FE281C5C3644578" originalValue="mesófilo">mesofilo</normalizedToken>
|
||
, 2200 m elevation) at Orizaba Peak, Veracruz, Mexico. In the original description,
|
||
<bibRefCitation id="3F762498A1826007885FE83BFBD0B4AE" pageId="20" pageNumber="115">Truqui (1857)</bibRefCitation>
|
||
cited one specimen collected by
|
||
<normalizedToken id="836DF0417B35E92E2B65BD8E2899656C" originalValue="Sallé">Salle</normalizedToken>
|
||
from Jacal near the Orizaba Volcano. This locality corresponds to El Jacal, Coscomatepec, Orizaba Peak (
|
||
<bibRefCitation id="44BD76832851ADFFFC6E9762E04BF32D" pageId="20" pageNumber="115">Reyes-Castillo 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="414742E9558AB09FDC2E965B8165FA8F" pageId="20" pageNumber="115">MEXICO: Veracruz: Calcahualco (Tecuanapa, road from Calcahualco to the Pico de Orizaba), Jacale, Pico de Orizaba.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04FF786F3EBCF67366956B60735651D7" lastPageId="21" lastPageNumber="116" pageId="20" pageNumber="115" type="remarks">
|
||
<paragraph id="C8917DBAE95BE3C018A5C4FE1DDB33C4" pageId="20" pageNumber="115">Remarks.</paragraph>
|
||
<paragraph id="357B4F7A358908734D8904FF7D2B0CB4" lastPageId="21" lastPageNumber="116" pageId="20" pageNumber="115">
|
||
<bibRefCitation id="DC65805C57691238516A85AA3DEA752A" author="Castillo, C" journalOrPublisher="Acta Zoologica Mexicana (nueva serie)" pageId="29" pageNumber="124" pagination="1 - 84" title="Biosistematica del genero Petrejoides Kuwert (Coleoptera, Lamellicornia, Passalidae)." volume="4" year="1984">Castillo and Reyes-Castillo (1984)</bibRefCitation>
|
||
redescribed
|
||
<taxonomicName id="32BE5BE687F9ABB02012FA6BFD1E19F7" lsidName="Y. laticornis" pageId="20" pageNumber="115" rank="species" species="laticornis">Y. laticornis</taxonomicName>
|
||
without examining type specimens. We compared two specimens of
|
||
<taxonomicName id="ADC3C6C08C25367E1D5E1D981D4F90A6" lsidName="Y. laticornis" pageId="20" pageNumber="115" rank="species" species="laticornis">Y. laticornis</taxonomicName>
|
||
determined by Castillo and Reyes-Castillo with the holotype specimen designated by M. E. Bachus at The Natural History Museum, London. Close examination of the holotype and results of the phylogenetic analysis (Fig. 4 and Suppl. materials 2-4) provide evidence that
|
||
<taxonomicName id="C28A1A75931AEC5CD6AA4AF3914B762D" lsidName="Y. laticornis" pageId="20" pageNumber="115" rank="species" species="laticornis">Y. laticornis</taxonomicName>
|
||
is not conspecific with
|
||
<taxonomicName id="C53AF879BE36F6E66546088B81C2E9FF" lsidName="Y. laticornis" pageId="20" pageNumber="115" rank="species" species="laticornis">Y. laticornis</taxonomicName>
|
||
sensu
|
||
<bibRefCitation id="58A9979D7EC8CC8F94808F0264C9C84B" author="Castillo, C" journalOrPublisher="Acta Zoologica Mexicana (nueva serie)" pageId="29" pageNumber="124" pagination="1 - 84" title="Biosistematica del genero Petrejoides Kuwert (Coleoptera, Lamellicornia, Passalidae)." volume="4" year="1984">Castillo and Reyes-Castillo (1984)</bibRefCitation>
|
||
(
|
||
<taxonomicName id="7DCF0E85873DCA67CD04F0931AE80BDD" genus="Y." lsidName="Y." pageId="21" pageNumber="116" rank="genus">
|
||
<pageBreakToken id="9B68106E81C1B88DFAA8D33FEDCEAE9C" pageId="21" pageNumber="116" start="start">Y</pageBreakToken>
|
||
.
|
||
</taxonomicName>
|
||
LM in Suppl. materials 2-4). Specimens described as
|
||
<taxonomicName id="9CD367D594C83A47E3945863A7A1556E" lsidName="Y. laticornis" pageId="21" pageNumber="116" rank="species" species="laticornis">Y. laticornis</taxonomicName>
|
||
by
|
||
<bibRefCitation id="50636985CE9120BEB213AD02CF934986" author="Castillo, C" journalOrPublisher="Acta Zoologica Mexicana (nueva serie)" pageId="29" pageNumber="124" pagination="1 - 84" title="Biosistematica del genero Petrejoides Kuwert (Coleoptera, Lamellicornia, Passalidae)." volume="4" year="1984">Castillo and Reyes-Castillo (1984)</bibRefCitation>
|
||
correspond with
|
||
<taxonomicName id="92034D16AD5A0A51BEA929E3D66FECDB" lsidName="Y. cameliae" pageId="21" pageNumber="116" rank="species" species="cameliae">Y. cameliae</taxonomicName>
|
||
sp. n. (
|
||
<taxonomicName id="EB5E17F84CD3D11E97DBC7293354E9D3" genus="Y." lsidName="Y." pageId="21" pageNumber="116" rank="genus">Y.</taxonomicName>
|
||
LM in Suppl. materials 2-4), and the holotype of
|
||
<taxonomicName id="2655790D63C4A74006BD7862D08D2224" lsidName="Y. laticornis" pageId="21" pageNumber="116" rank="species" species="laticornis">Y. laticornis</taxonomicName>
|
||
corresponds with
|
||
<taxonomicName id="4183464210E380659B99B66990D57981" genus="Y." lsidName="Y." pageId="21" pageNumber="116" rank="genus">Y.</taxonomicName>
|
||
CM (Suppl. materials 2-4). The overall length of the holotype specimen is 30.0 mm, and this falls within the size range for
|
||
<taxonomicName id="CA943534AF0373B239FD51C7DE3DCD26" lsidName="Y. laticornis" pageId="21" pageNumber="116" rank="species" species="laticornis">Y. laticornis</taxonomicName>
|
||
(25.0-33.0 mm), but not within the range for
|
||
<taxonomicName id="352A6ED9CA370FD2B6386F34362B800A" lsidName="Y. cameliae" pageId="21" pageNumber="116" rank="species" species="cameliae">Y. cameliae</taxonomicName>
|
||
(22.5-25.5 mm). Furthermore, based on distribution and biogeography,
|
||
<taxonomicName id="EB853682D0A9CF0926F47D73FA8AD59F" lsidName="Y. cameliae" pageId="21" pageNumber="116" rank="species" species="cameliae">Y. cameliae</taxonomicName>
|
||
has been collected only in the type locality where suitable habitat for the species occurs. This area is geographically isolated from the distribution area of
|
||
<taxonomicName id="44085D69C1E5AC377C897A7F14FFA581" lsidName="Y. laticornis" pageId="21" pageNumber="116" rank="species" species="laticornis">Y. laticornis</taxonomicName>
|
||
.
|
||
<taxonomicName id="F765B4B72C59289A6A6D514DD4C42F2E" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax laticornis" order="Coleoptera" pageId="21" pageNumber="116" phylum="Arthropoda" rank="species" species="laticornis">Yumtaax laticornis</taxonomicName>
|
||
' inclusion in the
|
||
<taxonomicName id="831A51853CEB9DEBFA9D2829977F05D1" lsidName="Y. laticornis" pageId="21" pageNumber="116" rank="species" species="laticornis">Y. laticornis</taxonomicName>
|
||
clade (is strongly supported (PP 1.0/BS 100) (Fig. 4).
|
||
</paragraph>
|
||
<caption id="06908626CF79D5F5CC1F48D62B85EE18" pageId="21" pageNumber="116">
|
||
<paragraph id="1BD3099376BA18B8BA850FC45A9688B4" pageId="21" pageNumber="116">
|
||
Figure 7.
|
||
<taxonomicName id="F060951E17469A9332ECA0BA3C31AF6F" class="Insecta" family="Passalidae" genus="Yumtaax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yumtaax laticornis" order="Coleoptera" pageId="21" pageNumber="116" phylum="Arthropoda" rank="species" species="laticornis">Yumtaax laticornis</taxonomicName>
|
||
(Truqui), holotype: A dorsal habitus B ventral habitus C dorsal view of pronotum and head D lateral view of pronotum and head E lateral view of right mesotibia F lateral view of aedeagus G ventral view of aedeagus H dorsal view of aedeagus.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |