1433 lines
148 KiB
XML
1433 lines
148 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.97.59559" ID-GBIF-Dataset="a466a75c-706c-4cbb-b591-99df3300f175" ID-GBIF-Taxon="177318571" ID-Pensoft-Pub="1860-0743-1-121" ID-Pensoft-UUID="D6B72C47EAF45F7A94A24A00BAFAF95B" ID-ZooBank="94AF3450CEAF4F0C8149C3F9BA9E405A" ModsDocID="1860-0743-1-121" checkinTime="1612984351702" checkinUser="pensoft" docAuthor="Molina-Acevedo, Isabel Cristina & Idris, Izwandy" docDate="2021" docId="3927268468CD501C8B929B4076438B58" docLanguage="en" docName="ZoosystEvol 97(1): 121-139" docOrigin="Zoosystematics and Evolution 97 (1)" docPubDate="2021-02-10" docSource="http://dx.doi.org/10.3897/zse.97.59559" docTitle="Marphysa teretiuscula" docType="treatment" docVersion="3" id="D6B72C47EAF45F7A94A24A00BAFAF95B" lastPageNumber="121" masterDocId="D6B72C47EAF45F7A94A24A00BAFAF95B" masterDocTitle="Unravelling the convoluted nomenclature of Marphysa simplex (Annelida, Eunicidae) with the proposal of a new name and the re-description of species" masterLastPageNumber="139" masterPageNumber="121" pageId="0" pageNumber="121" updateTime="1643426773792" updateUser="ExternalLinkService">
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<mods:title>Unravelling the convoluted nomenclature of Marphysa simplex (Annelida, Eunicidae) with the proposal of a new name and the re-description of species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Molina-Acevedo, Isabel Cristina</mods:namePart>
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<mods:affiliation>Estructura y Funcion del Bentos, Depto. Sistematica y Ecologia Acuatica, El Colegio de la Frontera Sur, Chetumal, Quintana Roo, Mexico & South China Sea Repository and Reference Centre, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Idris, Izwandy</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1516-8175</mods:nameIdentifier>
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<mods:affiliation>South China Sea Repository and Reference Centre, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-02-10</mods:number>
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<mods:number>97</mods:number>
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<mods:start>121</mods:start>
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<subSection pageId="0" pageNumber="121" type="results">
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<treatment ID-GBIF-Taxon="177318571" LSID="urn:lsid:plazi:treatment:3927268468CD501C8B929B4076438B58" httpUri="http://treatment.plazi.org/id/3927268468CD501C8B929B4076438B58" lastPageNumber="121" pageId="0" pageNumber="121">
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<subSubSection pageId="0" pageNumber="121" type="nomenclature">
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<taxonomicName LSID="39272684-68CD-501C-8B92-9B4076438B58" authority="(Schmarda, 1861 a)" baseAuthorityName="Schmarda" baseAuthorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Marphysa teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">Marphysa teretiuscula (Schmarda, 1861a)</taxonomicName>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">Figures 5</figureCitation>
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Marphysa teretiuscula (Schmarda, 1861 a). A, B. Chaetiger 3; C. Chaetiger 7; D. Chaetiger 12; E. Chaetiger 14; F. Chaetiger 47; G. Chaetiger 97; H. Chaetiger 143; I. Chaetiger 162; J. Chaetiger 44 before pygidium; K. Chaetiger 256. All chaetigers in anterior view. A, D, F, H. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); J. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); B, C, E, G, I, K. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). Arrows in F, H. indicate the button-shaped branchial stem. Scale bars: 0.2 mm (A, D, F, H, J); 0.1 mm (B, C, E, G, I, K)." figureDoi="10.3897/zse.97.59559.figure6" httpUri="https://binary.pensoft.net/fig/507571" pageId="0" pageNumber="121">, 6</figureCitation>
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Marphysa teretiuscula (Schmarda, 1861 a). A. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 47; B. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 189; C. Thick, isodont wide, with long and wide teeth, chaetiger 44 before pygidium; D. Thick, anodont wide, with long and wide teeth, chaetiger 256; E. Compound spinigers, chaetiger 44 before pygidium; F. Compound spiniger, chaetiger 211; G. Compound spinigers, chaetiger 47; H. Unidentate subacicular hook, chaetiger 47; I. Bidentate subacicular hook, chaetiger 73 before pygidium. A, B. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); C, E, I. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); D, F. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25); G, H. from M. simplex Crossland, 1903 syntype 2 (BNHM 1955.4.1.21 - 25). Scale bars: 20 μm (A-D, G); 60 μm (E); 50 μm (F, H, I)." figureDoi="10.3897/zse.97.59559.figure7" httpUri="https://binary.pensoft.net/fig/507572" pageId="0" pageNumber="121">, 7</figureCitation>
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Large chaetiger 10 (L 10) / Wide chaetiger 10 (W 10) - dependent variation of some morphological features in Marphysa teretiuscula (Schmarda, 1861 a). A. First chaetiger where the branchiae start (R ² = 0.7328, p = 1.65708 E- 05, n = 7); B. Last chaetiger where the postchaetal lobe is developed (R ² = 0.7976, p = 0.00028646, n = 7); C. Chaetiger where the subacicular hook starts (R ² = 0.6291, p = 2.02774 E- 07, n = 7)." figureDoi="10.3897/zse.97.59559.figure8" httpUri="https://binary.pensoft.net/fig/507573" pageId="0" pageNumber="121">, 8</figureCitation>
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution of branchial filaments throughout the body. A. Syntype 1 of Marphysa simplex Crossland, 1903 (BNHM 1924.3.1.1 - 2) with L 10: 8.6 mm, TL: 137 mm and 273 chaetigers; B. Syntype 1 of Marphysa furcellata Crossland, 1903 (BNHM 1924.3.1.139) with L 10: 6.3 mm, TL: 40 mm and 114 chaetigers; C. Syntype 1 of Marphysa macintoshi Crossland, 1903 (BNHM 1924.3.1.22 - 3) with L 10: 8.1 mm, TL: 18.8 mm and 262 chaetigers. Dotted blue line in B indicates the organism is incomplete." figureDoi="10.3897/zse.97.59559.figure9" httpUri="https://binary.pensoft.net/fig/507574" pageId="0" pageNumber="121">, 9</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological features of Marphysa species with reddish subacicular hook and three types of pectinate chaetae. Abbreviations: MF: Maxillary formula, roman numerals refer to number of maxilla; MxC: maxillary carriers; CIS: closing system; COp: cavity opening; PR-I: first peristomial ring; PR-II: second peristomial ring; Chaet: chaetiger; p / a: present / absent; AR: anterior region; MR: median region; PR: posterior region; SH: subacicular hook. INSS: Isodont narrow with short and slender teeth; INLS: Isodont narrow with long and slender teeth; INLT: Isodont narrow with long and thick teeth; IWSS: Isodont wide with short and slender teeth; IWLS: Isodont wide with long and slender teeth; IWLT: Isodont wide with long and thick teeth; AWLT: Anodont wide with long and thick teeth; AWLS: Anodont wide with long and slender teeth." pageId="0" pageNumber="121">Table 2</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName authorityName="Schmarda" authorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Eunice" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eunice teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">Eunice teretiuscula</taxonomicName>
|
||
Schmarda, 1861a: 129, pl. 32, fig. 59, text-figs a-d, f, OK, UK;
|
||
<bibRefCitation author="Grube, AE" journalOrPublisher="Naturwissenschaftliche Schriften Gesellschaft, Berlin" pageId="0" pageNumber="121" pagination="37 - 62" refId="B21" refString="Grube, AE, 1878. Untersuchungen ueber die Familie Eunicea. Naturwissenschaftliche Schriften Gesellschaft, Berlin 1878: 37 - 62" title="Untersuchungen ueber die Familie Eunicea." volume="1878" year="1878">Grube 1878</bibRefCitation>
|
||
: 59.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName baseAuthorityName="Schmarda" baseAuthorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Marphysa teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">Marphysa teretiuscula</taxonomicName>
|
||
-
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.122818" author="de Quatrefages, A" journalOrPublisher="Librairie Encyclopedique de Roret, Paris" pageId="0" pageNumber="121" refId="B10" refString="de Quatrefages, A, 1866. [1865] Histoire Naturelle des Anneles Marins et d'Eau Douce: Annelides et Gephyriens. Tome Premier. Collection des Suites a Buffon formant avec les Oeuvres de cet auteur un Cours Complet d'Histoire Naturelle. Librairie Encyclopedique de Roret, Paris, DOI: https://doi.org/10.5962/bhl.title.122818" title="[1865] Histoire Naturelle des Anneles Marins et d'Eau Douce: Annelides et Gephyriens. Tome Premier. Collection des Suites a Buffon formant avec les Oeuvres de cet auteur un Cours Complet d'Histoire Naturelle." url="https://doi.org/10.5962/bhl.title.122818" year="1866">de Quatrefages 1866</bibRefCitation>
|
||
: 337;
|
||
<bibRefCitation author="Ehlers, E" journalOrPublisher="Librairie Encyclopedique de Roret, Paris" pageId="0" pageNumber="121" refId="B11" refString="Ehlers, E, 1868. Die Borstenwürmer (AnnelidaChaetopoda) nach systematischen und anatomischen Untersuchungen dargestellt. Verlag von Wilhelm Engelmann, Leipzig, 269-748 pp." title="Die Borstenwu ̈ rmer (AnnelidaChaetopoda) nach systematischen und anatomischen Untersuchungen dargestellt. Verlag von Wilhelm Engelmann, Leipzig, 269 - 748 pp." year="1868">Ehlers 1868</bibRefCitation>
|
||
: 359;
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1469-7998.1903.tb08273.x" author="Crossland, C" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="121" pagination="169 - 176" refId="B5" refString="Crossland, C, 1903. On the marine fauna of Zanzibar and British East Africa, from collections made by Cyril Crossland in the years 1901 and 1902. Polychaeta. Part I. Proceedings of the Zoological Society of London 73 (1): 169 - 176, DOI: https://doi.org/10.1111/j.1469-7998.1903.tb08273.x" title="On the marine fauna of Zanzibar and British East Africa, from collections made by Cyril Crossland in the years 1901 and 1902. Polychaeta. Part I." url="https://doi.org/10.1111/j.1469-7998.1903.tb08273.x" volume="73" year="1903">Crossland 1903</bibRefCitation>
|
||
: 136; -
|
||
<bibRefCitation author="Hartman, O" journalOrPublisher="Allan Hancock Foundation Occasional Paper" pageId="0" pageNumber="121" pagination="1 - 628" refId="B24" refString="Hartman, O, 1959. Catalogue of the Polychaetous Annelids of the World. Parts 1 and 2. Allan Hancock Foundation Occasional Paper 23: 1 - 628" title="Catalogue of the Polychaetous Annelids of the World. Parts 1 and 2." volume="23" year="1959">Hartman 1959</bibRefCitation>
|
||
: 332; -
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2352.1.2" author="Glasby, CJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="121" pagination="29 - 45" refId="B19" refString="Glasby, CJ, Hutchings, PA, 2010. A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352: 29 - 45, DOI: https://doi.org/10.11646/zootaxa.2352.1.2" title="A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865." url="https://doi.org/10.11646/zootaxa.2352.1.2" volume="2352" year="2010">Glasby and Hutchings 2010</bibRefCitation>
|
||
: 32, 40-41, table 2;
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4377.2.3" author="Liu, Y" journalOrPublisher="Zootaxa" pageId="0" pageNumber="121" pagination="228 - 250" refId="B32" refString="Liu, Y, Hutchings, PA, Sun, S, 2017. Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934. Zootaxa 4263: 228 - 250, DOI: https://doi.org/10.11646/zootaxa.4377.2.3" title="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934." url="https://doi.org/10.11646/zootaxa.4377.2.3" volume="4263" year="2017">Liu et al. 2017</bibRefCitation>
|
||
: 244-247, table 3; -
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4377.2.3" author="Liu, Y" journalOrPublisher="Zootaxa" pageId="0" pageNumber="121" pagination="191 - 215" refId="B33" refString="Liu, Y, Hutchings, PA, Kupriyanova, E, 2018. Two new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern coast of China and redescription for Marphysa orientalis Treadwell, 1936. Zootaxa 4377: 191 - 215, DOI: https://doi.org/10.11646/zootaxa.4377.2.3" title="Two new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern coast of China and redescription for Marphysa orientalis Treadwell, 1936." url="https://doi.org/10.11646/zootaxa.4377.2.3" volume="4377" year="2018">Liu et al. 2018</bibRefCitation>
|
||
: 210-211, table 1.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Marphysa simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">Marphysa simplex</taxonomicName>
|
||
Crossland, 1903: 140-141, pl. 15, figs. 11-12, text-fig. 13.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="121">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName authorityName="Schmarda" authorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Eunice" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eunice teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Eunice teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
Schmarda, 1861a
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Sri Lanka · two specimens, one of them missing anterior end; Trincomalee, east of Sri Lanka; May 1853 to Jan 1854; L.K. Schmarda leg.;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">syntypes</emphasis>
|
||
NHMW type 1092.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa simplex</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Zanzibar · two adult specimens; 11 Jan 1934; Murray Exped. St. 104, Petersen Grab, V.310, 207 m;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">syntypes</emphasis>
|
||
BNHM type 1937.9.2.325.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="121">Other material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName baseAuthorityName="Schmarda" baseAuthorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a)
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">Mozambique · two specimens; Morrumbene Estuary; 16 Jan 1954; BNHM 1955.4.1.21-25.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
India · one specimen; Ratnagiri Creek, Shirgaon, Maharashtra;
|
||
<geoCoordinate degrees="17" direction="north" minutes="17" orientation="latitude" precision="1" seconds="13.78" value="17.28716">17°17'13.78"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73" direction="east" minutes="17" orientation="longitude" precision="1" seconds="13.87" value="73.287186">73°17'13.87"E</geoCoordinate>
|
||
; 18 Apr 1994; USNM 1128572 · one specimen; same data as for preceding; USNM 1128570.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="121">Comparative material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName authorityName="Crossland" authorityYear="1903" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa furcellata</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Zanzibar · two specimens; 1901; between tidemarks, 27.4 m; C. Crossland leg.;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">syntypes</emphasis>
|
||
BNHM 1924.3.1.139.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName authorityName="Crossland" authorityYear="1903" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa macintoshi</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Zanzibar · three specimens; 1901-1902; collected by digging in sand between tidemarks on both east and west coast of Zanzibar;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">syntypes</emphasis>
|
||
BNHM 1924.3.1.22-3, slide BNHM.1924.3.1.22A.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="description">
|
||
<paragraph pageId="0" pageNumber="121">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Syntype NHM type 1092 incomplete, gravid female, with 210 chaetigers, L10 = 9.3 mm, W10 = 5 mm TL = 860 mm (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5A-C</figureCitation>
|
||
). Anterior region with dorsum convex, flat ventre (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5C, E</figureCitation>
|
||
); body depressed from chaetiger 13 (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5C, E</figureCitation>
|
||
), widest at chaetiger 51, tapering after chaetiger 173.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 5.</emphasis>
|
||
<taxonomicName baseAuthorityName="Schmarda" baseAuthorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a).
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A.</emphasis>
|
||
Anterior end, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">B.</emphasis>
|
||
Anterior end, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C.</emphasis>
|
||
Anterior end, lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">D.</emphasis>
|
||
Anterior end, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">E.</emphasis>
|
||
Anterior end, lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">F.</emphasis>
|
||
Median region, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">G.</emphasis>
|
||
Pygidium, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">H.</emphasis>
|
||
Maxillary apparatus, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">I.</emphasis>
|
||
Left MI-II-III-IV-V, lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">J.</emphasis>
|
||
Attachment lamella in left side, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">K.</emphasis>
|
||
Attachment lamella in right side, lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">L.</emphasis>
|
||
Mandible, ventral view.
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A-C, H-L.</emphasis>
|
||
from
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861) syntype 1 (NHMW type 1092);
|
||
<emphasis bold="true" pageId="0" pageNumber="121">D-G.</emphasis>
|
||
from
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa simplex</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903 syntype 1 (BNHM 1955.4.1.21-25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A-C</emphasis>
|
||
); 3.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">D-F</emphasis>
|
||
); 1.16 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">G</emphasis>
|
||
); 3.0 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">H, I, L</emphasis>
|
||
); 1.13 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">J</emphasis>
|
||
); 0.9 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">K</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Prostomium bilobed, 4 mm long, 2.5 mm wide; lobes frontally rounded; median sulcus (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5A, B, D</figureCitation>
|
||
) shallow and deep ventrally. Prostomial appendages in semicircle, median antenna isolated by a gap. Palps reaching second chaetiger; lateral antennae reaching middle of third chaetiger; median antennae reaching fourth chaetiger. Palpophores and ceratophores ring-shaped, short, thick; palpostyles and ceratostyles tapering, slender. On right side with two palpostyle in the same palpophore (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5C</figureCitation>
|
||
). Eyes oval, brown, between palps and lateral antennae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Peristomium (2 mm long, 5.2 mm wide) wider than prostomium, first ring twice as long as second ring, separation between rings distinct on all sides (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5A-E</figureCitation>
|
||
). Ventral lip with slight central depression, with several shallow wrinkles (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Maxillary apparatus with MF = 1+1, 4+4, 5+0, 5+7, 1+1 (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5H</figureCitation>
|
||
). MI three times longer than length of maxillary carriers. MI forceps-like, MI four times longer than closing system (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5H, I</figureCitation>
|
||
); sclerotised ligament between MI and MII. MII wider than rest of maxillae, with triangular teeth; MII 3.2 times longer than cavity opening (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5H</figureCitation>
|
||
); ligament between left MII-MIII and right MII-MIV, slightly sclerotised. MIII with triangular teeth; with irregular attachment lamella, situated in centre of ventral edge of maxilla, slightly sclerotised (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5I</figureCitation>
|
||
). Left MIV with two teeth larger than rest of teeth; attachment lamella semicircle, wide, better developed in right portion, situated 2/3 along anterior edge of maxilla (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5J</figureCitation>
|
||
). Right MIV with four teeth larger than rest of teeth; attachment lamella semicircle, wide, better developed in central portion, situated 2/3 along anterior edge of maxilla (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5K</figureCitation>
|
||
). MV square, with a short triangular tooth. Mandibles dark; with calcareous cutting plates; sclerotised cutting plates brown, with nine growth rings (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Branchiae from chaetiger 32, with up to five long filaments; with two forms: palmate with short button-shaped branchial stem in anterior chaetigers (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Marphysa teretiuscula (Schmarda, 1861 a). A, B. Chaetiger 3; C. Chaetiger 7; D. Chaetiger 12; E. Chaetiger 14; F. Chaetiger 47; G. Chaetiger 97; H. Chaetiger 143; I. Chaetiger 162; J. Chaetiger 44 before pygidium; K. Chaetiger 256. All chaetigers in anterior view. A, D, F, H. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); J. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); B, C, E, G, I, K. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). Arrows in F, H. indicate the button-shaped branchial stem. Scale bars: 0.2 mm (A, D, F, H, J); 0.1 mm (B, C, E, G, I, K)." figureDoi="10.3897/zse.97.59559.figure6" httpUri="https://binary.pensoft.net/fig/507571" pageId="0" pageNumber="121">6F, H</figureCitation>
|
||
), pectinate in median chaetigers (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Marphysa teretiuscula (Schmarda, 1861 a). A, B. Chaetiger 3; C. Chaetiger 7; D. Chaetiger 12; E. Chaetiger 14; F. Chaetiger 47; G. Chaetiger 97; H. Chaetiger 143; I. Chaetiger 162; J. Chaetiger 44 before pygidium; K. Chaetiger 256. All chaetigers in anterior view. A, D, F, H. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); J. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); B, C, E, G, I, K. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). Arrows in F, H. indicate the button-shaped branchial stem. Scale bars: 0.2 mm (A, D, F, H, J); 0.1 mm (B, C, E, G, I, K)." figureDoi="10.3897/zse.97.59559.figure6" httpUri="https://binary.pensoft.net/fig/507571" pageId="0" pageNumber="121">6G</figureCitation>
|
||
). In second syntype, branchiae ending 25 chaetigers before pygidium. One filament in chaetigers 32L-34L; 2 in chaetigers 35L-39L; 3 in chaetigers 40L-48L; 3, 4 or 5 from chaetiger 49L to last chaetiger of the fragment. In second syntype, last 18 branchiae with one filament. Branchial filaments longer than dorsal cirri.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.59559.figure6" httpUri="https://binary.pensoft.net/fig/507571" pageId="0" pageNumber="121" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 6.</emphasis>
|
||
<taxonomicName baseAuthorityName="Schmarda" baseAuthorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a).
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A, B.</emphasis>
|
||
Chaetiger 3;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C.</emphasis>
|
||
Chaetiger 7;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">D.</emphasis>
|
||
Chaetiger 12;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">E.</emphasis>
|
||
Chaetiger 14;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">F.</emphasis>
|
||
Chaetiger 47;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">G.</emphasis>
|
||
Chaetiger 97;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">H.</emphasis>
|
||
Chaetiger 143;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">I.</emphasis>
|
||
Chaetiger 162;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">J.</emphasis>
|
||
Chaetiger 44 before pygidium;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">K.</emphasis>
|
||
Chaetiger 256. All chaetigers in anterior view.
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A, D, F, H.</emphasis>
|
||
from
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a) syntype 1 (NHMW type 1092);
|
||
<emphasis bold="true" pageId="0" pageNumber="121">J.</emphasis>
|
||
from
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a) syntype 2 (NHMW type 1092);
|
||
<emphasis bold="true" pageId="0" pageNumber="121">B, C, E, G, I, K.</emphasis>
|
||
from
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa simplex</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903 syntype 1 (BNHM 1955.4.1.21-25). Arrows in
|
||
<emphasis bold="true" pageId="0" pageNumber="121">F, H.</emphasis>
|
||
indicate the button-shaped branchial stem. Scale bars: 0.2 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A, D, F, H, J</emphasis>
|
||
); 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">B, C, E, G, I, K</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
First pair of parapodia small; best developed in chaetigers 11-56, following parapodia gradually decreasing in size. Dorsal cirri conical in all chaetigers; longer than ventral cirri in anterior and posterior chaetigers, shorter in median chaetigers; best developed in chaetigers 3-37, following gradually decreasing in size (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Marphysa teretiuscula (Schmarda, 1861 a). A, B. Chaetiger 3; C. Chaetiger 7; D. Chaetiger 12; E. Chaetiger 14; F. Chaetiger 47; G. Chaetiger 97; H. Chaetiger 143; I. Chaetiger 162; J. Chaetiger 44 before pygidium; K. Chaetiger 256. All chaetigers in anterior view. A, D, F, H. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); J. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); B, C, E, G, I, K. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). Arrows in F, H. indicate the button-shaped branchial stem. Scale bars: 0.2 mm (A, D, F, H, J); 0.1 mm (B, C, E, G, I, K)." figureDoi="10.3897/zse.97.59559.figure6" httpUri="https://binary.pensoft.net/fig/507571" pageId="0" pageNumber="121">6A-K</figureCitation>
|
||
). Prechaetal lobes short, in anterior chaetigers dorsal edge longer than ventral, in median-posterior chaetigers, as transverse fold (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Marphysa teretiuscula (Schmarda, 1861 a). A, B. Chaetiger 3; C. Chaetiger 7; D. Chaetiger 12; E. Chaetiger 14; F. Chaetiger 47; G. Chaetiger 97; H. Chaetiger 143; I. Chaetiger 162; J. Chaetiger 44 before pygidium; K. Chaetiger 256. All chaetigers in anterior view. A, D, F, H. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); J. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); B, C, E, G, I, K. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). Arrows in F, H. indicate the button-shaped branchial stem. Scale bars: 0.2 mm (A, D, F, H, J); 0.1 mm (B, C, E, G, I, K)." figureDoi="10.3897/zse.97.59559.figure6" httpUri="https://binary.pensoft.net/fig/507571" pageId="0" pageNumber="121">6A-K</figureCitation>
|
||
). Chaetal lobes rounded in first 35 chaetigers, shorter than other lobes, with aciculae emerging dorsal to mid-line; triangular from chaetiger 36, longer than other lobes, with aciculae emerging in mid-line (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Marphysa teretiuscula (Schmarda, 1861 a). A, B. Chaetiger 3; C. Chaetiger 7; D. Chaetiger 12; E. Chaetiger 14; F. Chaetiger 47; G. Chaetiger 97; H. Chaetiger 143; I. Chaetiger 162; J. Chaetiger 44 before pygidium; K. Chaetiger 256. All chaetigers in anterior view. A, D, F, H. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); J. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); B, C, E, G, I, K. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). Arrows in F, H. indicate the button-shaped branchial stem. Scale bars: 0.2 mm (A, D, F, H, J); 0.1 mm (B, C, E, G, I, K)." figureDoi="10.3897/zse.97.59559.figure6" httpUri="https://binary.pensoft.net/fig/507571" pageId="0" pageNumber="121">6A-K</figureCitation>
|
||
). Postchaetal lobes well developed in first 56 chaetigers; ovoid with dorsal edge longer than ventral edge in following chaetigers; progressively smaller from chaetiger 19; from chaetiger 57, inconspicuous (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Marphysa teretiuscula (Schmarda, 1861 a). A, B. Chaetiger 3; C. Chaetiger 7; D. Chaetiger 12; E. Chaetiger 14; F. Chaetiger 47; G. Chaetiger 97; H. Chaetiger 143; I. Chaetiger 162; J. Chaetiger 44 before pygidium; K. Chaetiger 256. All chaetigers in anterior view. A, D, F, H. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); J. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); B, C, E, G, I, K. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). Arrows in F, H. indicate the button-shaped branchial stem. Scale bars: 0.2 mm (A, D, F, H, J); 0.1 mm (B, C, E, G, I, K)." figureDoi="10.3897/zse.97.59559.figure6" httpUri="https://binary.pensoft.net/fig/507571" pageId="0" pageNumber="121">6A-K</figureCitation>
|
||
). Ventral cirri conical in first five chaetigers; from chaetiger 6 to last chaetiger of fragment with short oval swollen base and digitiform tip (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Marphysa teretiuscula (Schmarda, 1861 a). A, B. Chaetiger 3; C. Chaetiger 7; D. Chaetiger 12; E. Chaetiger 14; F. Chaetiger 47; G. Chaetiger 97; H. Chaetiger 143; I. Chaetiger 162; J. Chaetiger 44 before pygidium; K. Chaetiger 256. All chaetigers in anterior view. A, D, F, H. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); J. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); B, C, E, G, I, K. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). Arrows in F, H. indicate the button-shaped branchial stem. Scale bars: 0.2 mm (A, D, F, H, J); 0.1 mm (B, C, E, G, I, K)." figureDoi="10.3897/zse.97.59559.figure6" httpUri="https://binary.pensoft.net/fig/507571" pageId="0" pageNumber="121">6A-K</figureCitation>
|
||
). Second syntype with ventral cirri with short oval swollen base and digitiform tip up to 27 chaetigers before pygidium; digitiform in following ones, gradually decreasing in size posteriorly.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Aciculae blunt, basally reddish and translucent distally (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Marphysa teretiuscula (Schmarda, 1861 a). A, B. Chaetiger 3; C. Chaetiger 7; D. Chaetiger 12; E. Chaetiger 14; F. Chaetiger 47; G. Chaetiger 97; H. Chaetiger 143; I. Chaetiger 162; J. Chaetiger 44 before pygidium; K. Chaetiger 256. All chaetigers in anterior view. A, D, F, H. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); J. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); B, C, E, G, I, K. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). Arrows in F, H. indicate the button-shaped branchial stem. Scale bars: 0.2 mm (A, D, F, H, J); 0.1 mm (B, C, E, G, I, K)." figureDoi="10.3897/zse.97.59559.figure6" httpUri="https://binary.pensoft.net/fig/507571" pageId="0" pageNumber="121">6A-K</figureCitation>
|
||
). First two chaetigers with two aciculae; in chaetigers 3-5 with three or four; in chaetigers 6-47 with four or five; in chaetigers 48-139 with three; from chaetiger 140, with two. In second syntype, last 20 chaetigers with one acicula.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Limbate chaetae of two lengths in same chaetiger, dorsalmost chaetae longer; reduced in number around chaetiger 13. Three types of pectinate chaetae; from chaetiger 11 thin, isodont narrow chaetae, with short and slender teeth; in anterior chaetigers with 1-2 pectinate and with up to 21-22 teeth; in median-posterior chaetigers, with 20-25 pectinate and 30-32 teeth (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Marphysa teretiuscula (Schmarda, 1861 a). A. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 47; B. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 189; C. Thick, isodont wide, with long and wide teeth, chaetiger 44 before pygidium; D. Thick, anodont wide, with long and wide teeth, chaetiger 256; E. Compound spinigers, chaetiger 44 before pygidium; F. Compound spiniger, chaetiger 211; G. Compound spinigers, chaetiger 47; H. Unidentate subacicular hook, chaetiger 47; I. Bidentate subacicular hook, chaetiger 73 before pygidium. A, B. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); C, E, I. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); D, F. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25); G, H. from M. simplex Crossland, 1903 syntype 2 (BNHM 1955.4.1.21 - 25). Scale bars: 20 μm (A-D, G); 60 μm (E); 50 μm (F, H, I)." figureDoi="10.3897/zse.97.59559.figure7" httpUri="https://binary.pensoft.net/fig/507572" pageId="0" pageNumber="121">7A, B</figureCitation>
|
||
). In median-posterior chaetigers, 3-4 thick, isodont wide chaetae, with up to 16-18 long and wide teeth (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Marphysa teretiuscula (Schmarda, 1861 a). A. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 47; B. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 189; C. Thick, isodont wide, with long and wide teeth, chaetiger 44 before pygidium; D. Thick, anodont wide, with long and wide teeth, chaetiger 256; E. Compound spinigers, chaetiger 44 before pygidium; F. Compound spiniger, chaetiger 211; G. Compound spinigers, chaetiger 47; H. Unidentate subacicular hook, chaetiger 47; I. Bidentate subacicular hook, chaetiger 73 before pygidium. A, B. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); C, E, I. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); D, F. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25); G, H. from M. simplex Crossland, 1903 syntype 2 (BNHM 1955.4.1.21 - 25). Scale bars: 20 μm (A-D, G); 60 μm (E); 50 μm (F, H, I)." figureDoi="10.3897/zse.97.59559.figure7" httpUri="https://binary.pensoft.net/fig/507572" pageId="0" pageNumber="121">7C</figureCitation>
|
||
). In posterior chaetigers, 2-3 thick, anodont wide chaetae, with up to 6-7 long and thick teeth (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Marphysa teretiuscula (Schmarda, 1861 a). A. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 47; B. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 189; C. Thick, isodont wide, with long and wide teeth, chaetiger 44 before pygidium; D. Thick, anodont wide, with long and wide teeth, chaetiger 256; E. Compound spinigers, chaetiger 44 before pygidium; F. Compound spiniger, chaetiger 211; G. Compound spinigers, chaetiger 47; H. Unidentate subacicular hook, chaetiger 47; I. Bidentate subacicular hook, chaetiger 73 before pygidium. A, B. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); C, E, I. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); D, F. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25); G, H. from M. simplex Crossland, 1903 syntype 2 (BNHM 1955.4.1.21 - 25). Scale bars: 20 μm (A-D, G); 60 μm (E); 50 μm (F, H, I)." figureDoi="10.3897/zse.97.59559.figure7" httpUri="https://binary.pensoft.net/fig/507572" pageId="0" pageNumber="121">7D</figureCitation>
|
||
). Compound spinigers present in all chaetigers, with blades of two lengths in the same chaetiger, shorter ones more abundant (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Marphysa teretiuscula (Schmarda, 1861 a). A. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 47; B. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 189; C. Thick, isodont wide, with long and wide teeth, chaetiger 44 before pygidium; D. Thick, anodont wide, with long and wide teeth, chaetiger 256; E. Compound spinigers, chaetiger 44 before pygidium; F. Compound spiniger, chaetiger 211; G. Compound spinigers, chaetiger 47; H. Unidentate subacicular hook, chaetiger 47; I. Bidentate subacicular hook, chaetiger 73 before pygidium. A, B. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); C, E, I. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); D, F. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25); G, H. from M. simplex Crossland, 1903 syntype 2 (BNHM 1955.4.1.21 - 25). Scale bars: 20 μm (A-D, G); 60 μm (E); 50 μm (F, H, I)." figureDoi="10.3897/zse.97.59559.figure7" httpUri="https://binary.pensoft.net/fig/507572" pageId="0" pageNumber="121">7E, F, G</figureCitation>
|
||
). Subacicular hooks present from chaetiger 33 to 140, with continuous distribution, one or two per chaetiger (second one replacement); unidentate in anterior chaetigers (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Marphysa teretiuscula (Schmarda, 1861 a). A. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 47; B. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 189; C. Thick, isodont wide, with long and wide teeth, chaetiger 44 before pygidium; D. Thick, anodont wide, with long and wide teeth, chaetiger 256; E. Compound spinigers, chaetiger 44 before pygidium; F. Compound spiniger, chaetiger 211; G. Compound spinigers, chaetiger 47; H. Unidentate subacicular hook, chaetiger 47; I. Bidentate subacicular hook, chaetiger 73 before pygidium. A, B. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); C, E, I. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); D, F. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25); G, H. from M. simplex Crossland, 1903 syntype 2 (BNHM 1955.4.1.21 - 25). Scale bars: 20 μm (A-D, G); 60 μm (E); 50 μm (F, H, I)." figureDoi="10.3897/zse.97.59559.figure7" httpUri="https://binary.pensoft.net/fig/507572" pageId="0" pageNumber="121">7H</figureCitation>
|
||
), bidentate in median chaetigers, basally reddish translucent distally; with blunt teeth, distal and proximal teeth of similar sizes, booth teeth directed upwards (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Marphysa teretiuscula (Schmarda, 1861 a). A. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 47; B. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 189; C. Thick, isodont wide, with long and wide teeth, chaetiger 44 before pygidium; D. Thick, anodont wide, with long and wide teeth, chaetiger 256; E. Compound spinigers, chaetiger 44 before pygidium; F. Compound spiniger, chaetiger 211; G. Compound spinigers, chaetiger 47; H. Unidentate subacicular hook, chaetiger 47; I. Bidentate subacicular hook, chaetiger 73 before pygidium. A, B. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); C, E, I. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); D, F. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25); G, H. from M. simplex Crossland, 1903 syntype 2 (BNHM 1955.4.1.21 - 25). Scale bars: 20 μm (A-D, G); 60 μm (E); 50 μm (F, H, I)." figureDoi="10.3897/zse.97.59559.figure7" httpUri="https://binary.pensoft.net/fig/507572" pageId="0" pageNumber="121">7I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.59559.figure7" httpUri="https://binary.pensoft.net/fig/507572" pageId="0" pageNumber="121" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 7.</emphasis>
|
||
<taxonomicName baseAuthorityName="Schmarda" baseAuthorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a).
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A.</emphasis>
|
||
Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 47;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">B.</emphasis>
|
||
Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 189;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C.</emphasis>
|
||
Thick, isodont wide, with long and wide teeth, chaetiger 44 before pygidium;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">D.</emphasis>
|
||
Thick, anodont wide, with long and wide teeth, chaetiger 256;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">E.</emphasis>
|
||
Compound spinigers, chaetiger 44 before pygidium;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">F.</emphasis>
|
||
Compound spiniger, chaetiger 211;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">G.</emphasis>
|
||
Compound spinigers, chaetiger 47;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">H.</emphasis>
|
||
Unidentate subacicular hook, chaetiger 47;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">I.</emphasis>
|
||
Bidentate subacicular hook, chaetiger 73 before pygidium.
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A, B.</emphasis>
|
||
from
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a) syntype 1 (NHMW type 1092);
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C, E, I.</emphasis>
|
||
from
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a) syntype 2 (NHMW type 1092);
|
||
<emphasis bold="true" pageId="0" pageNumber="121">D, F.</emphasis>
|
||
from
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa simplex</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903 syntype 1 (BNHM 1955.4.1.21-25);
|
||
<emphasis bold="true" pageId="0" pageNumber="121">G, H.</emphasis>
|
||
from
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. simplex</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903 syntype 2 (BNHM 1955.4.1.21-25). Scale bars: 20
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A-D, G</emphasis>
|
||
); 60
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="121">E</emphasis>
|
||
); 50
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="121">F, H, I</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
In second syntype, pygidium with dorsal pairs of anal cirri, as long as last 12 chaetigers; ventral pair of anal cirri short, as long as last three chaetigers (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="variation">
|
||
<paragraph pageId="0" pageNumber="121">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Material examined varied in the following features: L10 = 3.1-12.4 mm, W10 = 0.8-5 mm, TChae = 88-265. Palps reaching middle of first peristomial ring or first chaetiger; lateral antennae reaching first or middle of first chaetiger; median antenna reaching middle of first or second chaetiger. Maxillary formula: MII 4-6+4-7, MIII 5-8, MIV 4-5+7-9. MI is 3-3.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than maxillary carriers; MI is 4.4-5.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than closing system; MII is 2.7-3.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than cavity opening. Branchiae starting from chaetigers 15-32 and disappearing 7-12 chaetigers before pygidium. The maximum number of branchial filaments varies from two to six. Postchaetal lobes well developed in first 20-56 chaetigers. Ventral cirri with swollen base starting from chaetigers 4-8 and disappearing 34-68 chaetigers before pygidium. Start of subacicular hooks from chaetigers 23-38.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Regression analyses showed a correlation between L10/W10 and the first branchiate chaetiger (R² = 0.7328,
|
||
<emphasis italics="true" pageId="0" pageNumber="121">p</emphasis>
|
||
= 1.65708E-05, n = 7, Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Large chaetiger 10 (L 10) / Wide chaetiger 10 (W 10) - dependent variation of some morphological features in Marphysa teretiuscula (Schmarda, 1861 a). A. First chaetiger where the branchiae start (R ² = 0.7328, p = 1.65708 E- 05, n = 7); B. Last chaetiger where the postchaetal lobe is developed (R ² = 0.7976, p = 0.00028646, n = 7); C. Chaetiger where the subacicular hook starts (R ² = 0.6291, p = 2.02774 E- 07, n = 7)." figureDoi="10.3897/zse.97.59559.figure8" httpUri="https://binary.pensoft.net/fig/507573" pageId="0" pageNumber="121">8A</figureCitation>
|
||
), the last chaetiger with developed postchaetal lobe (R² = 0.7976,
|
||
<emphasis italics="true" pageId="0" pageNumber="121">p</emphasis>
|
||
= 0.00028646, n = 7, Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Large chaetiger 10 (L 10) / Wide chaetiger 10 (W 10) - dependent variation of some morphological features in Marphysa teretiuscula (Schmarda, 1861 a). A. First chaetiger where the branchiae start (R ² = 0.7328, p = 1.65708 E- 05, n = 7); B. Last chaetiger where the postchaetal lobe is developed (R ² = 0.7976, p = 0.00028646, n = 7); C. Chaetiger where the subacicular hook starts (R ² = 0.6291, p = 2.02774 E- 07, n = 7)." figureDoi="10.3897/zse.97.59559.figure8" httpUri="https://binary.pensoft.net/fig/507573" pageId="0" pageNumber="121">8B</figureCitation>
|
||
) and the first chaetiger with subacicular hook (R² = 0.6291,
|
||
<emphasis italics="true" pageId="0" pageNumber="121">p</emphasis>
|
||
= 2.02774E-07, n = 7, Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Large chaetiger 10 (L 10) / Wide chaetiger 10 (W 10) - dependent variation of some morphological features in Marphysa teretiuscula (Schmarda, 1861 a). A. First chaetiger where the branchiae start (R ² = 0.7328, p = 1.65708 E- 05, n = 7); B. Last chaetiger where the postchaetal lobe is developed (R ² = 0.7976, p = 0.00028646, n = 7); C. Chaetiger where the subacicular hook starts (R ² = 0.6291, p = 2.02774 E- 07, n = 7)." figureDoi="10.3897/zse.97.59559.figure8" httpUri="https://binary.pensoft.net/fig/507573" pageId="0" pageNumber="121">8C</figureCitation>
|
||
). Most of the specimens were incomplete and regression analysis regarding the maximum number of branchial filaments in the body could not be performed.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.59559.figure8" httpUri="https://binary.pensoft.net/fig/507573" pageId="0" pageNumber="121" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 8.</emphasis>
|
||
Large chaetiger 10 (L10)/Wide chaetiger 10 (W10)-dependent variation of some morphological features in
|
||
<taxonomicName baseAuthorityName="Schmarda" baseAuthorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a).
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A.</emphasis>
|
||
First chaetiger where the branchiae start (R² = 0.7328,
|
||
<emphasis italics="true" pageId="0" pageNumber="121">p</emphasis>
|
||
= 1.65708E-05, n = 7);
|
||
<emphasis bold="true" pageId="0" pageNumber="121">B.</emphasis>
|
||
Last chaetiger where the postchaetal lobe is developed (R² = 0.7976,
|
||
<emphasis italics="true" pageId="0" pageNumber="121">p</emphasis>
|
||
= 0.00028646, n = 7);
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C.</emphasis>
|
||
Chaetiger where the subacicular hook starts (R² = 0.6291,
|
||
<emphasis italics="true" pageId="0" pageNumber="121">p</emphasis>
|
||
= 2.02774E-07, n = 7).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="distribution">
|
||
<paragraph pageId="0" pageNumber="121">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">Sri Lanka, Maharashtra (India), Zanzibar.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="habitat">
|
||
<paragraph pageId="0" pageNumber="121">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Unknown.
|
||
<bibRefCitation author="Schmarda, LK" journalOrPublisher="Erster Band (zweite halfte) Turbellarian, Rotatorien un Anneliden" pageId="0" pageNumber="121" pagination="1 - 164" refId="B46" refString="Schmarda, LK, 1861a. Neue wirbellose Thiere beobachtet und gesammelt auf einer Reise un die Erdr 1853 bis 1857. Erster Band (zweite halfte) Turbellarian, Rotatorien un Anneliden 2: 1 - 164" title="Neue wirbellose Thiere beobachtet und gesammelt auf einer Reise un die Erdr 1853 bis 1857." volume="2" year="1861 a">Schmarda (1861a)</bibRefCitation>
|
||
did not indicate the habitat of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="remarks">
|
||
<paragraph pageId="0" pageNumber="121">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<bibRefCitation author="Schmarda, LK" journalOrPublisher="Erster Band (zweite halfte) Turbellarian, Rotatorien un Anneliden" pageId="0" pageNumber="121" pagination="1 - 164" refId="B46" refString="Schmarda, LK, 1861a. Neue wirbellose Thiere beobachtet und gesammelt auf einer Reise un die Erdr 1853 bis 1857. Erster Band (zweite halfte) Turbellarian, Rotatorien un Anneliden 2: 1 - 164" title="Neue wirbellose Thiere beobachtet und gesammelt auf einer Reise un die Erdr 1853 bis 1857." volume="2" year="1861 a">Schmarda (1861a)</bibRefCitation>
|
||
collected
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(firstly in the genus
|
||
<taxonomicName class="Polychaeta" family="Eunicidae" genus="Eunice" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eunice" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Eunice</emphasis>
|
||
</taxonomicName>
|
||
) in the east of Ceylon (now Sri Lanka) during a series of expeditions around the world to collect fauna and flora (
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.85313" author="Schmarda, LK" journalOrPublisher="Wilhelm Engelmann, Leipzig" pageId="0" pageNumber="121" refId="B45" refString="Schmarda, LK, 1859. Neue wirbellose Tiere beobachtet und gesammelt auf einer Reise um die Erde 1853 bis 1857. Erster Band: Neue Turbellarien, Rotatorien und Anneliden. Erste Haelfte. Wilhelm Engelmann, Leipzig, DOI: https://doi.org/10.5962/bhl.title.85313" title="Neue wirbellose Tiere beobachtet und gesammelt auf einer Reise um die Erde 1853 bis 1857. Erster Band: Neue Turbellarien, Rotatorien und Anneliden. Erste Haelfte." url="https://doi.org/10.5962/bhl.title.85313" year="1859">Schmarda 1859</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.5252/zoosystema2019v41a24" author="Villalobos-Guerrero, TF" journalOrPublisher="Zoosystema" pageId="0" pageNumber="121" pagination="465 - 496" refId="B52" refString="Villalobos-Guerrero, TF, 2019. Redescription of two overlooked species of the Perinereis nuntia complex and morphological delimitation of P. nuntia (Savigny in Lamarck, 1818) from the Red Sea (Annelida, Nereididae). Zoosystema 41 (24): 465 - 496, DOI: https://doi.org/10.5252/zoosystema2019v41a24" title="Redescription of two overlooked species of the Perinereis nuntia complex and morphological delimitation of P. nuntia (Savigny in Lamarck, 1818) from the Red Sea (Annelida, Nereididae)." url="https://doi.org/10.5252/zoosystema2019v41a24" volume="41" year="2019">Villalobos-Guerrero 2019</bibRefCitation>
|
||
). The syntypes label only states
|
||
<normalizedToken originalValue="‘Trincomalie’">'Trincomalie'</normalizedToken>
|
||
(Trincomalee) as the collecting site, but no collecting date is given. However, the expedition notes (
|
||
<bibRefCitation author="Schmarda, LK" journalOrPublisher="George Westermann, Braunschweig (Brunswick), Vol. 1" pageId="0" pageNumber="121" refId="B47" refString="Schmarda, LK, 1861b. Reise um die Erde in den Jahre 1853-1857. George Westermann, Braunschweig (Brunswick), Vol. 1" title="Reise um die Erde in den Jahre 1853 - 1857." year="1861 b">Schmarda 1861b</bibRefCitation>
|
||
) state that he visited Ceylon from May 1853 to January 1854, whereby, based on this information, the syntypes of
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
were most likely collected during this time.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1469-7998.1903.tb08273.x" author="Crossland, C" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="121" pagination="169 - 176" refId="B5" refString="Crossland, C, 1903. On the marine fauna of Zanzibar and British East Africa, from collections made by Cyril Crossland in the years 1901 and 1902. Polychaeta. Part I. Proceedings of the Zoological Society of London 73 (1): 169 - 176, DOI: https://doi.org/10.1111/j.1469-7998.1903.tb08273.x" title="On the marine fauna of Zanzibar and British East Africa, from collections made by Cyril Crossland in the years 1901 and 1902. Polychaeta. Part I." url="https://doi.org/10.1111/j.1469-7998.1903.tb08273.x" volume="73" year="1903">Crossland (1903)</bibRefCitation>
|
||
described
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. macintoshi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. simplex</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
from Zanzibar. These species were differentiated, based on the shape of the prostomium and the pectinate chaetae. However, some authors considered these features irrelevant over time and proposed several synonyms between them or other species from distant regions. For instance,
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.8154" author="Fauvel, P" journalOrPublisher="Archives de zoologie experimentale et generale" pageId="0" pageNumber="121" pagination="315 - 473" refId="B14" refString="Fauvel, P, 1919. Annelides Polychetes de Madagascar, de Djibouti, et du Golfe Persique. Archives de zoologie experimentale et generale 58: 315 - 473, DOI: https://doi.org/10.5962/bhl.part.8154" title="Annelides Polychetes de Madagascar, de Djibouti, et du Golfe Persique." url="https://doi.org/10.5962/bhl.part.8154" volume="58" year="1919">Fauvel (1919)</bibRefCitation>
|
||
considered
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
to be a junior synonym of
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. sanguinea" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="sanguinea">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. sanguinea</emphasis>
|
||
</taxonomicName>
|
||
Montagu, 1813. On the contrary,
|
||
<bibRefCitation author="Day, JH" journalOrPublisher="Annals of the Natal Museum" pageId="0" pageNumber="121" pagination="59 - 129" refId="B7" refString="Day, JH, 1957. The Polychaeta Fauna of South Africa. Part 4. New species and records from Natal and Mocambique. Annals of the Natal Museum 14 (1): 59 - 129" title="The Polychaeta Fauna of South Africa. Part 4. New species and records from Natal and Mocambique." volume="14" year="1957">Day (1957)</bibRefCitation>
|
||
indicated that
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. sanguinea" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="sanguinea">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. sanguinea</emphasis>
|
||
</taxonomicName>
|
||
differed from
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
by having bidentate subacicular hooks, whereas, in the latter species, they are unidentate. However, Day regarded
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. simplex</emphasis>
|
||
</taxonomicName>
|
||
(Crossland). Later,
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1469-7998.1962.tb01597.x" author="Day, JH" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="121" pagination="627 - 656" refId="B8" refString="Day, JH, 1962. Polychaeta from several localities in the western Indian Ocean. Proceedings of the Zoological Society of London 139 (4): 627 - 656, DOI: https://doi.org/10.1111/j.1469-7998.1962.tb01597.x" title="Polychaeta from several localities in the western Indian Ocean." url="https://doi.org/10.1111/j.1469-7998.1962.tb01597.x" volume="139" year="1962">Day (1962)</bibRefCitation>
|
||
pointed out that
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. simplex</emphasis>
|
||
</taxonomicName>
|
||
(Crossland) were synonyms of
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. macintoshi</emphasis>
|
||
</taxonomicName>
|
||
, considering that the
|
||
<normalizedToken originalValue="prostomium’s">prostomium's</normalizedToken>
|
||
shape was insufficient to differentiate them. More recently,
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2352.1.2" author="Glasby, CJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="121" pagination="29 - 45" refId="B19" refString="Glasby, CJ, Hutchings, PA, 2010. A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352: 29 - 45, DOI: https://doi.org/10.11646/zootaxa.2352.1.2" title="A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865." url="https://doi.org/10.11646/zootaxa.2352.1.2" volume="2352" year="2010">Glasby and Hutchings (2010)</bibRefCitation>
|
||
recognised that an entire prostomium is useful to distinguish
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. macintoshi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. simplex</emphasis>
|
||
</taxonomicName>
|
||
(Crossland). Simultaneously,
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2352.1.2" author="Glasby, CJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="121" pagination="29 - 45" refId="B19" refString="Glasby, CJ, Hutchings, PA, 2010. A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352: 29 - 45, DOI: https://doi.org/10.11646/zootaxa.2352.1.2" title="A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865." url="https://doi.org/10.11646/zootaxa.2352.1.2" volume="2352" year="2010">Glasby and Hutchings (2010)</bibRefCitation>
|
||
compared
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. simplex</emphasis>
|
||
</taxonomicName>
|
||
(Crossland) and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
, but they also did not detect morphological differences between them. After examining the type materials, we confirm the validity of
|
||
<normalizedToken originalValue="Crossland’s">Crossland's</normalizedToken>
|
||
species
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. macintoshi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
and the synonymy of
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. simplex</emphasis>
|
||
</taxonomicName>
|
||
(Crossland) with
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(see Figs
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Marphysa teretiuscula (Schmarda, 1861 a). A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Anterior end, dorsal view; E. Anterior end, lateral view; F. Median region, ventral view; G. Pygidium, dorsal view; H. Maxillary apparatus, dorsal view; I. Left MI-II-III-IV-V, lateral view; J. Attachment lamella in left side, dorsal view; K. Attachment lamella in right side, lateral view; L. Mandible, ventral view. A-C, H-L. from M. teretiuscula (Schmarda, 1861) syntype 1 (NHMW type 1092); D-G. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25). al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV; al-MIV-L: attachment lamella MIV on left side; al-MIV-R: attachment lamella MIV on right side; MI-R: Maxilla I on right side; MII-R: Maxilla II on right side. Scale bars: 2.3 mm (A-C); 3.1 mm (D-F); 1.16 mm (G); 3.0 mm (H, I, L); 1.13 mm (J); 0.9 mm (K)." figureDoi="10.3897/zse.97.59559.figure5" httpUri="https://binary.pensoft.net/fig/507570" pageId="0" pageNumber="121">5</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Marphysa teretiuscula (Schmarda, 1861 a). A. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 47; B. Thin, isodont narrow pectinate, with short and slender teeth, chaetiger 189; C. Thick, isodont wide, with long and wide teeth, chaetiger 44 before pygidium; D. Thick, anodont wide, with long and wide teeth, chaetiger 256; E. Compound spinigers, chaetiger 44 before pygidium; F. Compound spiniger, chaetiger 211; G. Compound spinigers, chaetiger 47; H. Unidentate subacicular hook, chaetiger 47; I. Bidentate subacicular hook, chaetiger 73 before pygidium. A, B. from M. teretiuscula (Schmarda, 1861 a) syntype 1 (NHMW type 1092); C, E, I. from M. teretiuscula (Schmarda, 1861 a) syntype 2 (NHMW type 1092); D, F. from Marphysa simplex Crossland, 1903 syntype 1 (BNHM 1955.4.1.21 - 25); G, H. from M. simplex Crossland, 1903 syntype 2 (BNHM 1955.4.1.21 - 25). Scale bars: 20 μm (A-D, G); 60 μm (E); 50 μm (F, H, I)." figureDoi="10.3897/zse.97.59559.figure7" httpUri="https://binary.pensoft.net/fig/507572" pageId="0" pageNumber="121">7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName baseAuthorityName="Schmarda" baseAuthorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. borradailei" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="borradailei">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. borradailei</emphasis>
|
||
</taxonomicName>
|
||
Pillai, 1958 from Sri Lanka and the Indian Ocean,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
from Zanzibar,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. gravelyi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="gravelyi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. gravelyi</emphasis>
|
||
</taxonomicName>
|
||
Southern, 1921 from Chilka Lake, India,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. macintoshi</emphasis>
|
||
</taxonomicName>
|
||
from Zanzibar and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. madrasi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="madrasi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. madrasi</emphasis>
|
||
</taxonomicName>
|
||
Hutchings, Lavesque, Priscilla, Daffe, Malathi & Glasby, 2020 from Ennore Creek, India by having compound spinigers and inhabiting the same geographical area. However,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
bears only subacicular chaetae compound spinigers, while
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. borradailei" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="borradailei">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. borradailei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. gravelyi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="gravelyi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. gravelyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. madrasi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="madrasi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. madrasi</emphasis>
|
||
</taxonomicName>
|
||
have both subacicular spinigers and limbate chaetae. Furthermore,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
has distinct bilobed prostomium, in contrast to an entire prostomium in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. macintoshi</emphasis>
|
||
</taxonomicName>
|
||
. Moreover,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
has palmate branchiae with a short button-shaped branchial stem in the anterior region, the postchaetal lobe is rounded in the first three chaetigers and the subacicular hooks are reddish basally and translucent distally. In contrast,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
has pectinate branchiae in the anterior region, digitiform postchaetal lobes in the first chaetigers and translucent subacicular hooks. In addition,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. macintoshi</emphasis>
|
||
</taxonomicName>
|
||
differ by distributing the branchial filaments throughout the body. In
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
, the maximum number of five branchial filaments is present only in a small/low number of chaetigers (between chaetiger 86 and 106), while in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. macintoshi</emphasis>
|
||
</taxonomicName>
|
||
, the maximum number of five branchial filaments (in each species) is found in a larger number of chaetigers (in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. furcellata</emphasis>
|
||
</taxonomicName>
|
||
from chaetiger 80 to 120+ and in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. macintoshi</emphasis>
|
||
</taxonomicName>
|
||
from chaetiger 105 to 236; Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution of branchial filaments throughout the body. A. Syntype 1 of Marphysa simplex Crossland, 1903 (BNHM 1924.3.1.1 - 2) with L 10: 8.6 mm, TL: 137 mm and 273 chaetigers; B. Syntype 1 of Marphysa furcellata Crossland, 1903 (BNHM 1924.3.1.139) with L 10: 6.3 mm, TL: 40 mm and 114 chaetigers; C. Syntype 1 of Marphysa macintoshi Crossland, 1903 (BNHM 1924.3.1.22 - 3) with L 10: 8.1 mm, TL: 18.8 mm and 262 chaetigers. Dotted blue line in B indicates the organism is incomplete." figureDoi="10.3897/zse.97.59559.figure9" httpUri="https://binary.pensoft.net/fig/507574" pageId="0" pageNumber="121">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.59559.figure9" httpUri="https://binary.pensoft.net/fig/507574" pageId="0" pageNumber="121" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 9.</emphasis>
|
||
Distribution of branchial filaments throughout the body.
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A.</emphasis>
|
||
Syntype 1 of
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa simplex</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903 (BNHM 1924.3.1.1-2) with L10: 8.6 mm, TL: 137 mm and 273 chaetigers;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">B.</emphasis>
|
||
Syntype 1 of
|
||
<taxonomicName authorityName="Crossland" authorityYear="1903" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa furcellata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="furcellata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa furcellata</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903 (BNHM 1924.3.1.139) with L10: 6.3 mm, TL: 40 mm and 114 chaetigers;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C.</emphasis>
|
||
Syntype 1 of
|
||
<taxonomicName authorityName="Crossland" authorityYear="1903" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa macintoshi" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="macintoshi">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa macintoshi</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903 (BNHM 1924.3.1.22-3) with L10: 8.1 mm, TL: 18.8 mm and 262 chaetigers. Dotted blue line in B indicates the organism is incomplete.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName baseAuthorityName="Schmarda" baseAuthorityYear="1861" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. americana" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="americana">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. americana</emphasis>
|
||
</taxonomicName>
|
||
Monro, 1933,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. angelensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="angelensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. angelensis</emphasis>
|
||
</taxonomicName>
|
||
Fauchald, 1970,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. depressa" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="depressa">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. depressa</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a),
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. emiliae" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="emiliae">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. emiliae</emphasis>
|
||
</taxonomicName>
|
||
Molina-Acevedo and Carrera-Parra, 2017,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. nobilis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="nobilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. nobilis</emphasis>
|
||
</taxonomicName>
|
||
Treadwell, 1917,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. sanguinea" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="sanguinea">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. sanguinea</emphasis>
|
||
</taxonomicName>
|
||
(Montagu, 1913) and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. tripectinata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="tripectinata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. tripectinata</emphasis>
|
||
</taxonomicName>
|
||
Liu, Hutchings & Sun, 2017 in having reddish subacicular hooks, the presence of compound spinigers and the absence of subacicular limbate chaetae. However,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
has palmate branchiae with a short bottom-stem in the anterior region, contrary to
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. americana" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="americana">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. americana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. angelensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="angelensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. angelensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. depressa" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="depressa">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. depressa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. emiliae" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="emiliae">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. emiliae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. nobilis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="nobilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. nobilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. sanguinea" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="sanguinea">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. sanguinea</emphasis>
|
||
</taxonomicName>
|
||
which have pectinate branchiae throughout the body. Furthermore,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
has compound spinigers in all chaetigers, while in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. depressa" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="depressa">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. depressa</emphasis>
|
||
</taxonomicName>
|
||
, the spinigers are restricted to the anterior region. In addition,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
has the postchaetal lobe rounded in the first three chaetigers, while it is conical in the first three parapodia of
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. americana" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="americana">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. americana</emphasis>
|
||
</taxonomicName>
|
||
and digitiform in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. angelensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="angelensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. angelensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. depressa" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="depressa">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. depressa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. emiliae" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="emiliae">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. emiliae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. sanguinea" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="sanguinea">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. sanguinea</emphasis>
|
||
</taxonomicName>
|
||
. Moreover,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
has distinctly longer branchial filaments than in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. angelensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="angelensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. angelensis</emphasis>
|
||
</taxonomicName>
|
||
. Additionally,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
has the subacicular hook as wide as the acicula, in contrast to that half as wide as acicula in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. nobilis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="nobilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. nobilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. tripectinata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="tripectinata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. tripectinata</emphasis>
|
||
</taxonomicName>
|
||
. The comparison of
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
with similar species is provided in Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological features of Marphysa species with reddish subacicular hook and three types of pectinate chaetae. Abbreviations: MF: Maxillary formula, roman numerals refer to number of maxilla; MxC: maxillary carriers; CIS: closing system; COp: cavity opening; PR-I: first peristomial ring; PR-II: second peristomial ring; Chaet: chaetiger; p / a: present / absent; AR: anterior region; MR: median region; PR: posterior region; SH: subacicular hook. INSS: Isodont narrow with short and slender teeth; INLS: Isodont narrow with long and slender teeth; INLT: Isodont narrow with long and thick teeth; IWSS: Isodont wide with short and slender teeth; IWLS: Isodont wide with long and slender teeth; IWLT: Isodont wide with long and thick teeth; AWLT: Anodont wide with long and thick teeth; AWLS: Anodont wide with long and slender teeth." httpUri="http://table.plazi.org/id/459CA3C9C009D7FC17D2C4808E2C9319" pageId="0" pageNumber="121" tableUuid="459CA3C9C009D7FC17D2C4808E2C9319">2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="459CA3C9C009D7FC17D2C4808E2C9319" httpUri="http://table.plazi.org/id/459CA3C9C009D7FC17D2C4808E2C9319" pageId="0" pageNumber="121" start="Table 2" startId="T2">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Table 2.</emphasis>
|
||
Morphological features of
|
||
<taxonomicName class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa</emphasis>
|
||
</taxonomicName>
|
||
species with reddish subacicular hook and three types of pectinate chaetae. Abbreviations: MF: Maxillary formula, roman numerals refer to number of maxilla; MxC: maxillary carriers; CIS: closing system; COp: cavity opening; PR-I: first peristomial ring; PR-II: second peristomial ring; Chaet: chaetiger; p/a: present/absent; AR: anterior region; MR: median region; PR: posterior region; SH: subacicular hook. INSS: Isodont narrow with short and slender teeth; INLS: Isodont narrow with long and slender teeth; INLT: Isodont narrow with long and thick teeth; IWSS: Isodont wide with short and slender teeth; IWLS: Isodont wide with long and slender teeth; IWLT: Isodont wide with long and thick teeth; AWLT: Anodont wide with long and thick teeth; AWLS: Anodont wide with long and slender teeth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<table pageId="0" pageNumber="121">
|
||
<tr pageId="0" pageNumber="121">
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">Morphological feature</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. americana" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="americana">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. americana</emphasis>
|
||
</taxonomicName>
|
||
Monro, 1933
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. angelensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="angelensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. angelensis</emphasis>
|
||
</taxonomicName>
|
||
Fauchald, 1970
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. depressa" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="depressa">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. depressa</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a)
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. emiliae" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="emiliae">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. emiliae</emphasis>
|
||
</taxonomicName>
|
||
Molina-Acevedo & Carrera-Parra, 2017
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. nobilis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="nobilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. nobilis</emphasis>
|
||
</taxonomicName>
|
||
Treadwell, 1917
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. sanguinea" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="sanguinea">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. sanguinea</emphasis>
|
||
</taxonomicName>
|
||
(Montagu, 1913)
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. teretiuscula" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="teretiuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. teretiuscula</emphasis>
|
||
</taxonomicName>
|
||
(Schmarda, 1861a)
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. tripectinata" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="tripectinata">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. tripectinata</emphasis>
|
||
</taxonomicName>
|
||
Liu, Hutchings & Sun, 2017
|
||
</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Source of information</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">holotype BNHM 1932.12.24.554-556</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">holotype LACM-AHF POLY 285; and additional material</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">syntypes NHM 1044</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">holotype ECOSUR0180; paratype ECOSUR0181</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">holotype AMNH VI-1915-1350, additional material</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">neotype BNHM 1867.1.7.24; additional material</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
syntypes NHM 1092,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. simplex</emphasis>
|
||
</taxonomicName>
|
||
Crossland, 1903 syntypes BNHM 1937.9.2.325
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">paratypes AM W.49071, AM W.49072</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Size (mm): L10, W10</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">12, 5.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2.1-5.4, 17</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4.2-11.5, 1.9-4.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">3.5, 1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">6 -13.6, 6.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">11.5-20.4, 7.2-11</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5.5-12.4; 0.8-5</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">11-12.3, 4-5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Prostomium: shape</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Palps: reaching</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-I</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II, Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-I, Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-I, PR-II</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-I, PR-II</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-I, Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Lateral antennae: reaching</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II, Chaet 2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1, 2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II, Chaet 2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II, Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Median antennae: reaching</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1, 3</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1, 3</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1, 3</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II, Chaet 2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1, 2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1, 2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Peduncle in prostomial appendages</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Eyes</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">MF: MII, MIII, MIV</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5+4, 7+0, 3+9</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">3-5+4-5, 4-6, 3-4+6-8</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">3-4+4, 4-5, 3-5+6-8</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4+4, 5+0, 3+8</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">3-4+3-4, 5-6, 3-4+7-8</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4+4-5, 5-6, 3-4+6-8</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4-6+4-7, 5-8, 4-5+7-9</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5+5-6, 7, 4-5+8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">MI vs. MxC: proportion</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
2.1-3.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
2-2.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
2.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
2.9-3.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3-3.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
2.6-2.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">MI vs. CIS: proportion</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3-4.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3.5-4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
4.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3.8-5.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
4.4-5.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
4-4.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">MII vs COp: proportion</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
4.3-6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3.1-5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3.7-4.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
2.7-3.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3.6-3.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Branchiae: shaped</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">pectinate</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">pectinate</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">pectinate</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">pectinate</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">pectinate</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">pectinate</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">palmate/pectinate</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">palmate/pectinate</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Branchiae: start chaetiger; last chaetiger before pygidium</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">45; 28</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">9-14; 3-31</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">26-44; 20-39</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">8-12; 10-13</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">17-27; 34-37</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">21-25; 9-18</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">18-32; 7-12</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">13-20; 16</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Branchial filaments: numbers, length of the filaments</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">12, long</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2-6, short</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2-4, long</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2-5, long</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4-6; long</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5-6; long</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4-6; long</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5-6; long</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Dorsal cirri: shape</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR, MR: conical, PR: digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: digitiform, MR: digitiform with swollen base, PR: conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: digitiform, MR, PR: conical</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Prechaetal lobe: shape</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">transverse fold</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">transverse fold</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">transverse fold</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">transverse fold</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR, MR, PR: transverse fold</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR, MR, PR: transverse fold</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: dorsal edge longer, MR, PR: transverse fold</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: dorsal edge longer, MR, PR: transverse fold</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaetal lobe: shape</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: rounded, MR, PR: triangular</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: rounded, MR, PR: triangular</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: rounded, MR, PR: triangular</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: rounded, MR, PR: triangular</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: rounded, MR, PR: triangular</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: rounded, MR, PR: triangular</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: rounded, MR, PR: triangular</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: rounded, MR, PR: triangular</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Developed postchaetal lobe: end chaetiger</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">92</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">18-61</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">27-60</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">42</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">39-71</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">50-70</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">20-56</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">50-131</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Poschaetal lobe: shape in body regions</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1-4: conical; from Chaet 5: rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1-4: digitiform, from Chaet 5: rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1-4: digitiform, from Chaet 5: auricular</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1-4: digitiform, from Chaet 5: rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1-4: digitiform, from Chaet 5: ovoid</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">ovoid dorsal edge longer</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">rounded with dorsal edge longer</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Ventral cirri in first chaetigers: shape</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Ventral cirri with swollen base: start chaetiger; last chaetiger before pygidium</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5; 45</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4-7; 28-58</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5-6; 22-28</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4-7; 39-44</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">10-11; 44</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5-8; 8-18</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4-8; 34-68</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">6-8; 151</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Ventral cirri in most posterior chaetigers: shape</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Aciculae: shape; colour</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Subacicular limbate chaetae</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Pectinate chaetae: type in AR; MR, PR</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INLS; IWSS, IWLT, AWLT</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INST; IWSS, AWLS</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INSS; IWLT, AWLT</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">IWSS; AWLT</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INLS; IWSS, AWLS</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INLS; IWSS, AWLS</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INLS; INLS, IWLT, AWLT</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INLS; IWLS, AWLS</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Pectinate chaetae: number per type</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">3-4; 3-4, 2-3, 1-2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">1-2; 3-4, 2-3</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2-3; 3-4, 3-4</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">6-8; 3-4</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2-3; 10-12, 6-7</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">1-2; 18-20, 4-5</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">1-2; 20-25, 3-4, 2-3</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">1-2; 16-17, 4-5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Pectinate chaetae teeth: number per type</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">12; 16, 16, 11</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">15; 18, 8-9</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">8-9; 14, 13-14</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">20-22; 13</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">16-17; 17, 16</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">10; 18, 10-12</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">21-22; 30-32, 16-18, 6-7</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">18; 25, 15-17</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Spiniger blade: length in AR</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">similar length</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Spiniger blade: length in MR-PR</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">similar length</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2 lengths</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Spiniger: distribution</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">all chaet</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">all chaet</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR only</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">all chaet</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">all chaet</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">all chaet</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">all chaet</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">all chaet</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Falciger: (p/a); distribution</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">a; NA</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">p; all chaet or AR</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">p; all chaet</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">p; AR</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Subacicular hook: start chaetiger</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">117</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">14-29</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">33-68</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">21-28</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">31-94</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">74-286</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">30-38</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">62-115</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Subacicular hook: shape; colour</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bidentate, reddish basally and translucent distally</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bidentate, reddish basally and translucent distally</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bidentate, reddish basally and translucent distally</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bidentate, reddish basally and translucent distally</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">unidentate, reddish basally and translucent distally</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bidentate, reddish basally and translucent distally</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bidentate, reddish basally and translucent distally</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bidentate, reddish basally and translucent distally</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Width acicula vs. SH in MR-PR: proportion</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">similar width</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">similar width</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">similar width</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">similar width</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
Acicula 2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
wider than SH
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
Acicula 2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
wider than SH
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">similar width</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
Acicula 2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
wider than SH
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Subacicular hook: distribution</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">discontinuous</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">continue</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">continue</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">continue</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">discontinuous</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">discontinuous</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">discontinuous</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">continuous</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |