treatments-xml/data/11/57/87/11578788FE5AFFC47DB5FF1216E3DFB1.xml
2024-06-21 12:29:42 +02:00

263 lines
32 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="61B86FACFDB78DE68A8DC2D7D61D8D1E" ID-DOI="10.5281/zenodo.193406" ID-GBIF-Dataset="5e8043a5-e685-4a21-b696-0f7a706ab223" ID-ISSN="1175-5326" ID-Zenodo-Dep="193406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460138703176" checkinUser="plazi" docAuthor="Toledo, Mario, Spangler, Paul J. &amp; Balke, Michael" docDate="2010" docId="11578788FE5AFFC47DB5FF1216E3DFB1" docLanguage="en" docName="zt02347p058.pdf" docOrigin="Zootaxa 2347" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laccodytes apalodes Guignot 1955" docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="ED6EFFF0FE5CFFC27D22FFF3160ED852" masterDocTitle="Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)" masterLastPageNumber="58" masterPageNumber="37" pageNumber="43" updateTime="1698235333712" updateUser="plazi">
<mods:mods id="309C0F5E97829842B73A23232CDB67A8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="11B6CE93074B45567908D3CD6A6E8791">
<mods:title id="52115C1AD22E26D661DEBE6072F04CB4">Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="C2A34DA15916750CEC2D670C6A1AEF66" type="personal">
<mods:role id="E6D08C359AED5405152343FDA41A571F">
<mods:roleTerm id="29260B5697D26DB2E9225B65377CF994">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="89AAC3F3292F5796116A24A9189EC11B">Toledo, Mario</mods:namePart>
</mods:name>
<mods:name id="8A4904DBF5E6ACF4972B056C6E8C4F40" type="personal">
<mods:role id="57510A7724F7408B9D89675D403DCEEA">
<mods:roleTerm id="4B0577DE944A4B52C9375006B088EF70">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F4AABB99A49207C969BB9808BE529086">Spangler, Paul J.</mods:namePart>
</mods:name>
<mods:name id="BCA3A2A41E5ABBD13D7D5B76E5CA1CDF" type="personal">
<mods:role id="187B5C814FC8CD1AD72B01440EA88FE4">
<mods:roleTerm id="02F3FB6B45A01E96402C43F6EAD9A8F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="156C0C04ED829F94710DFA0D4800DCE0">Balke, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource id="315138060290A75B9E36F8D1DDAD2CDA">text</mods:typeOfResource>
<mods:relatedItem id="215207CAE0DA726B244F507ED14E27CA" type="host">
<mods:titleInfo id="FB87BEC4BF3ED8D215224CFD802001D0">
<mods:title id="56E0FF315AAF1F1E02DD1A911EACDB96">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9777599AD0FE95E8AD02AA41649E0977">
<mods:date id="AE992CF9449E960B9F7775DDCAFA68B8">2010</mods:date>
<mods:detail id="ACA7448CE5FF49EBA96E0FD8DA009769" type="volume">
<mods:number id="6C2570B22A779E6F4F21FF67A33B3196">2347</mods:number>
</mods:detail>
<mods:extent id="6E95F05F4BD9A896FE9F459BCB9E0CE0" unit="page">
<mods:start id="BCF0E0F1A728C6BBDD1E0C65789BC9AE">37</mods:start>
<mods:end id="3EE4128F3410C6FD77946CAF0A98F788">58</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="85228F441A1166597E87ED11CE377853">journal article</mods:classification>
<mods:identifier id="7460DB865B848E67FA5656B5B54FC472" type="DOI">10.5281/zenodo.193406</mods:identifier>
<mods:identifier id="97EA76EBA9DBD1D21272C27EB78C3013" type="GBIF-Dataset">5e8043a5-e685-4a21-b696-0f7a706ab223</mods:identifier>
<mods:identifier id="23D16C19B9DFC9FC97424C83B2115829" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D81ECF5BD9B62255FBACB06D7C9E6BC6" type="Zenodo-Dep">193406</mods:identifier>
</mods:mods>
<treatment id="11578788FE5AFFC47DB5FF1216E3DFB1" ID-DOI="http://doi.org/10.5281/zenodo.6198364" ID-GBIF-Taxon="124955823" ID-Zenodo-Dep="6198364" LSID="urn:lsid:plazi:treatment:11578788FE5AFFC47DB5FF1216E3DFB1" httpUri="http://treatment.plazi.org/id/11578788FE5AFFC47DB5FF1216E3DFB1" lastPageNumber="43" pageId="6" pageNumber="43">
<subSubSection id="D1E46515FE5AFFC47DB5FF121482D94C" pageId="6" pageNumber="43" type="nomenclature">
<paragraph id="9941369EFE5AFFC47DB5FF12144AD8A8" blockId="6.[151,652,224,286]" box="[151,580,224,251]" pageId="6" pageNumber="43">
<heading id="C20981F2FE5AFFC47DB5FF12144AD8A8" bold="true" box="[151,580,224,251]" fontSize="11" level="1" pageId="6" pageNumber="43" reason="1">
<taxonomicName id="5EFE4D1DFE5AFFC47DB5FF12144AD8A8" ID-CoL="3RLZF" authority="Guignot, 1955" authorityName="Guignot" authorityYear="1955" box="[151,580,224,251]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="AB8AEA8CFE5AFFC47DB5FF12144AD8A8" bold="true" box="[151,580,224,251]" pageId="6" pageNumber="43">
<emphasis id="AB8AEA8CFE5AFFC47DB5FF121780D8A9" bold="true" box="[151,398,225,251]" italics="true" pageId="6" pageNumber="43">Laccodytes apalodes</emphasis>
<bibRefCitation id="FD6F4B6FFE5AFFC47CB6FF13144AD8A8" author="Guignot" box="[404,580,224,250]" pageId="6" pageNumber="58" refString="Guignot, F. (1955) Description de nouveaux Dytiscidae principalement de l'Amerique du Sud. Bulletin de l'Institut royal des Sciences naturelles de Belgique, 31 (27), 1 - 12." type="journal article" year="1955">Guignot, 1955</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="9941369EFE5AFFC47DB5FEF71482D94C" blockId="6.[151,652,224,286]" box="[151,652,260,286]" pageId="6" pageNumber="43">Figs (4, 7, 9, 12, 15, 2224, 28, 44, 54, 62)</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE5AFFC47DB5FEA21280D93A" box="[151,1166,337,360]" pageId="6" pageNumber="43" type="reference_group">
<paragraph id="9941369EFE5AFFC47DB5FEA21280D93A" blockId="6.[151,1166,337,360]" box="[151,1166,337,360]" pageId="6" pageNumber="43">
<treatmentCitationGroup id="B9EE11B0FE5AFFC47DB5FEA21280D93A" box="[151,1166,337,360]" pageId="6" pageNumber="43">
<taxonomicName id="5EFE4D1DFE5AFFC47DB5FEA21432D93A" ID-CoL="3RLZF" authority="Guignot (1955: 8)" authorityName="Guignot" authorityPageNumber="8" authorityYear="1955" box="[151,572,337,360]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="AB8AEA8CFE5AFFC47DB5FEA2177BD93A" box="[151,373,337,360]" italics="true" pageId="6" pageNumber="43">Laccodytes apalodes</emphasis>
<treatmentCitation id="185F108FFE5AFFC47C5EFEA21432D93A" author="Guignot" box="[380,572,337,360]" page="8" pageId="6" pageNumber="43" year="1955">
<bibRefCitation id="FD6F4B6FFE5AFFC47C5EFEA21432D93A" author="Guignot" box="[380,572,337,360]" pageId="6" pageNumber="58" refString="Guignot, F. (1955) Description de nouveaux Dytiscidae principalement de l'Amerique du Sud. Bulletin de l'Institut royal des Sciences naturelles de Belgique, 31 (27), 1 - 12." type="journal article" year="1955">Guignot (1955: 8)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(original description);
<treatmentCitation id="185F108FFE5AFFC47E11FEA21206D93A" author="Nilsson" box="[819,1032,337,360]" page="239" pageId="6" pageNumber="43" year="2001">
<bibRefCitation id="FD6F4B6FFE5AFFC47E11FEA21206D93A" author="Nilsson" box="[819,1032,337,360]" pageId="6" pageNumber="58" refString="Nilsson, A. N. (2001) World catalogue of insects. Vol. 3. Dytiscidae Coleoptera. Stenstrup, Apollo Books, 395 pp." type="book" year="2001">Nilsson (2001: 239)</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE5AFFC47DB5FE641460DC4E" pageId="6" pageNumber="43" type="materials_examined">
<paragraph id="9941369EFE5AFFC47DB5FE6414A3D9E0" blockId="6.[151,1437,407,2019]" box="[151,685,407,434]" pageId="6" pageNumber="43">
<emphasis id="AB8AEA8CFE5AFFC47DB5FE641733D9E3" bold="true" box="[151,317,407,433]" pageId="6" pageNumber="43">
<typeStatus id="4645883CFE5AFFC47DB5FE6416DAD9E3" box="[151,212,407,433]" pageId="6" pageNumber="43">Type</typeStatus>
locality.
</emphasis>
French
<collectingCountry id="E1E9760EFE5AFFC47CBEFE6B17F2D9E0" box="[412,508,408,434]" name="Guyana" pageId="6" pageNumber="43">Guyana</collectingCountry>
: Cayenne area.
</paragraph>
<paragraph id="9941369EFE5AFFC47DE4FE4D126ADA1E" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
<emphasis id="AB8AEA8CFE5AFFC47DE4FE4D1770D98A" bold="true" box="[198,382,446,472]" pageId="6" pageNumber="43">
<typeStatus id="4645883CFE5AFFC47DE4FE4D170DD98A" box="[198,259,446,472]" pageId="6" pageNumber="43">Type</typeStatus>
material.
</emphasis>
<typeStatus id="4645883CFE5AFFC47CA5FE4D17FAD98A" box="[391,500,446,472]" pageId="6" pageNumber="43" type="holotype">Holotype</typeStatus>
Ƥ (
<collectionCode id="FFEFAE5BFE5AFFC47F06FE4D148CD98A" LSID="urn:lsid:biocol.org:col:34988" box="[548,642,446,472]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="43">MNHN</collectionCode>
, Guignot collection):
<collectingCountry id="E1E9760EFE5AFFC47EAAFE4D15EDD98A" box="[904,995,446,472]" name="French Guiana" pageId="6" pageNumber="43">Guyane</collectingCountry>
Fr. [française], Env. [environs],
<date id="ED40105EFE5AFFC47848FE4D16DCD9AD" pageId="6" pageNumber="43" value="1937-08">VIII 1937</date>
// Museum Paris, Coll. Guignot //
<taxonomicName id="5EFE4D1DFE5AFFC47F47FE161555D9AD" box="[613,859,485,511]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="apalodes">Laccodytes apalodes</taxonomicName>
. The
<typeStatus id="4645883CFE5AFFC47EB8FE16120FD9AD" box="[922,1025,485,511]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
was studied and illustrated by Paul Spangler in 1980, and the redescription of
<taxonomicName id="5EFE4D1DFE5AFFC47FAAFDFF151FDA77" box="[648,785,524,549]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="AB8AEA8CFE5AFFC47FAAFDFF151FDA77" box="[648,785,524,549]" italics="true" pageId="6" pageNumber="43">L. apalodes</emphasis>
</taxonomicName>
is based on the notes taken by Paul Spangler back then, as well as on
<collectionCode id="FFEFAE5BFE5AFFC47C17FDC11784DA1E" LSID="urn:lsid:biocol.org:col:34871" box="[309,394,562,588]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="43">USNM</collectionCode>
specimens which agree well with these notes on the
<typeStatus id="4645883CFE5AFFC47ED7FDC11251DA1E" box="[1013,1119,562,588]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="9941369EFE5AFFC47DE4FDAB17E3DAB5" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
<emphasis id="AB8AEA8CFE5AFFC47DE4FDAB17FBDA20" bold="true" box="[198,501,600,626]" pageId="6" pageNumber="43">Other material studied.</emphasis>
<collectingCountry id="E1E9760EFE5AFFC47F23FDAA146FDA21" box="[513,609,601,627]" name="Guyana" pageId="6" pageNumber="43">Guyana</collectingCountry>
: Mazaruni-Potaro Distr., Takutu Mts., Takutu lumber camp,
<geoCoordinate id="FCCA5059FE5AFFC47868FDAA1395DA21" box="[1354,1435,601,627]" direction="north" orientation="latitude" pageId="6" pageNumber="43" precision="925" value="6.25">6°15'N</geoCoordinate>
<geoCoordinate id="FCCA5059FE5AFFC47DB5FD73170ADAC8" box="[151,260,640,666]" direction="west" orientation="longitude" pageId="6" pageNumber="43" precision="925" value="-59.0">59°00'W</geoCoordinate>
,
<date id="ED40105EFE5AFFC47C33FD7317A2DAC8" box="[273,428,640,666]" pageId="6" pageNumber="43" value="1982-12-30">30.XII.1982</date>
, W.E. Steiner, J.E. Lowry &amp; G.W. Williams (6 exs
<collectionCode id="FFEFAE5BFE5AFFC47909FD731287DAC8" LSID="urn:lsid:biocol.org:col:34871" box="[1067,1161,640,666]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="43">USNM</collectionCode>
); Takutu Mts.,
<geoCoordinate id="FCCA5059FE5AFFC47864FD731393DAC8" box="[1350,1437,640,666]" direction="north" orientation="latitude" pageId="6" pageNumber="43" precision="925" value="6.25">6°15'N</geoCoordinate>
<geoCoordinate id="FCCA5059FE5AFFC47DB5FD5516E1DA92" box="[151,239,678,704]" direction="west" orientation="longitude" pageId="6" pageNumber="43" precision="925" value="-59.083332">59°5'W</geoCoordinate>
, Earthwatch research expedition,
<date id="ED40105EFE5AFFC47F5FFD55153ADA92" box="[637,820,678,704]" pageId="6" pageNumber="43" value="1983-12-03" valueMax="1983-12-19" valueMin="1983-12-03">3.19.XII.1983</date>
, P.J. Spangler, W.E. Steiner &amp; R.A. Faitoute (59 exs
<collectionCode id="FFEFAE5BFE5AFFC47DB5FD3E16D6DAB5" box="[151,216,717,743]" pageId="6" pageNumber="43">CMT</collectionCode>
,
<collectionCode id="FFEFAE5BFE5AFFC47DC1FD3E1735DAB5" LSID="urn:lsid:biocol.org:col:34871" box="[227,315,717,743]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="43">USNM</collectionCode>
,
<collectionCode id="FFEFAE5BFE5AFFC47C64FD3E178BDAB5" box="[326,389,717,743]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="43">ZSM</collectionCode>
,
<collectionCode id="FFEFAE5BFE5AFFC47CB3FD3E17EFDAB5" box="[401,481,717,743]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="6" pageNumber="43">NMW</collectionCode>
).
</paragraph>
<paragraph id="9941369EFE5AFFC47DE4FD0716F3DB66" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
<collectingCountry id="E1E9760EFE5AFFC47DE4FD071735DB5C" box="[198,315,756,782]" name="Suriname" pageId="6" pageNumber="43">Suriname</collectingCountry>
:
<taxonomicName id="5EFE4D1DFE5AFFC47C6BFD07140FDB5C" authority="Creek" authorityName="Creek" box="[329,513,756,782]" class="Insecta" family="Scarabaeidae" genus="Carolina" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">Carolina Creek</taxonomicName>
,
<quantity id="5E069B7BFE5AFFC47F32FD071453DB5C" box="[528,605,756,782]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="43" unit="km" value="10.0">10 km</quantity>
from Zanderij, waterhole in forest st.,
<date id="ED40105EFE5AFFC47915FD0712CEDB5C" box="[1079,1216,756,782]" pageId="6" pageNumber="43" value="1962-11-18">18.XI.1962</date>
, B. Malkin (2 exs
<collectionCode id="FFEFAE5BFE5AFFC47DB5FCE916FFDB66" LSID="urn:lsid:biocol.org:col:34871" box="[151,241,794,820]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="43">USNM</collectionCode>
).
</paragraph>
<paragraph id="9941369EFE5AFFC47DE7FCB21460DC4E" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
<collectingCountry id="E1E9760EFE5AFFC47DE7FCB2174ADB09" box="[197,324,833,859]" name="Venezuela" pageId="6" pageNumber="43">Venezuela</collectingCountry>
: TF Amazonas, Cerro de la Neblina, basecamp,
<quantity id="5E069B7BFE5AFFC47EABFCB215D6DB09" box="[905,984,833,859]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="43" unit="m" value="140.0">140 m</quantity>
,
<geoCoordinate id="FCCA5059FE5AFFC47EC7FCB21237DB09" box="[997,1081,833,859]" direction="north" orientation="latitude" pageId="6" pageNumber="43" precision="925" value="0.8333333">0°50'N</geoCoordinate>
<geoCoordinate id="FCCA5059FE5AFFC47963FCB212A4DB09" box="[1089,1194,833,859]" direction="west" orientation="longitude" pageId="6" pageNumber="43" precision="925" value="-66.15">66°09'W</geoCoordinate>
,
<date id="ED40105EFE5AFFC47994FCB21399DB09" box="[1206,1431,833,859]" pageId="6" pageNumber="43" value="1984-11-24" valueMax="1984-12-01" valueMin="1984-11-24">24.XI.1.XII.1984</date>
, R.L. Brown (1 ex.
<collectionCode id="FFEFAE5BFE5AFFC47C50FC9B17C2DBD0" LSID="urn:lsid:biocol.org:col:34871" box="[370,460,872,898]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="43">USNM</collectionCode>
); Cerro de la Neblina, basecamp,
<quantity id="5E069B7BFE5AFFC47E76FC9B15AFDBD0" box="[852,929,872,898]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="43" unit="m" value="140.0">140 m</quantity>
,
<geoCoordinate id="FCCA5059FE5AFFC47E8EFC9B15F0DBD0" box="[940,1022,872,898]" direction="north" orientation="latitude" pageId="6" pageNumber="43" precision="925" value="0.8333333">0°50'N</geoCoordinate>
<geoCoordinate id="FCCA5059FE5AFFC47927FC9B1262DBD0" box="[1029,1132,872,898]" direction="west" orientation="longitude" pageId="6" pageNumber="43" precision="925" value="-66.166664">66°10'W</geoCoordinate>
,
<date id="ED40105EFE5AFFC47955FC9B12FADBD0" box="[1143,1268,872,898]" pageId="6" pageNumber="43" value="1985-02-19">19.II.1985</date>
, P.J. Spangler, &amp; A.R. Faitoute (2 exs
<collectionCode id="FFEFAE5BFE5AFFC47C8CFC7D1406DBFA" LSID="urn:lsid:biocol.org:col:34871" box="[430,520,910,936]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="43">USNM</collectionCode>
); Amazonas state,
<geoCoordinate id="FCCA5059FE5AFFC47FC4FC7D1565DBFA" box="[742,875,910,936]" direction="north" orientation="latitude" pageId="6" pageNumber="43" precision="1" value="5.0617833">5°03.707'N</geoCoordinate>
<geoCoordinate id="FCCA5059FE5AFFC47E56FC7D1201DBFA" box="[884,1039,910,936]" direction="west" orientation="longitude" pageId="6" pageNumber="43" precision="1" value="-67.779465">67°46.768'W</geoCoordinate>
,
<quantity id="5E069B7BFE5AFFC4793EFC7D1255DBFA" box="[1052,1115,910,936]" metricMagnitude="1" metricUnit="m" metricValue="9.2" pageId="6" pageNumber="43" unit="m" value="92.0">92 m</quantity>
, river near Orinoco/Sipapo confluent,
<date id="ED40105EFE5AFFC47C34FC461784DB9D" box="[278,394,949,975]" pageId="6" pageNumber="43" value="2009-01-15">15.I.2009</date>
, shallow backwaters, A. Short (locality number VZ09-0115-01C) (1 ex.
<collectionCode id="FFEFAE5BFE5AFFC479CEFC461320DB9D" box="[1260,1326,949,975]" pageId="6" pageNumber="43">KUL</collectionCode>
); Bolivar state,
<geoCoordinate id="FCCA5059FE5AFFC47DFDFC2F176DDBA4" box="[223,355,988,1014]" direction="north" orientation="latitude" pageId="6" pageNumber="43" precision="1" value="6.58694">6.58694°N</geoCoordinate>
<geoCoordinate id="FCCA5059FE5AFFC47C49FC2F17F8DBA4" box="[363,502,988,1014]" direction="west" orientation="longitude" pageId="6" pageNumber="43" precision="5" value="-67.0912">67.0912°W</geoCoordinate>
, Rio Caripito, near Rio Orinoco,
<date id="ED40105EFE5AFFC47EACFC2F120BDBA4" box="[910,1029,988,1014]" pageId="6" pageNumber="43" value="2009-01-12">12.I.2009</date>
, A. Short &amp; K.B. Miller (locality number VZ09-0112-02°) (12 exs
<collectionCode id="FFEFAE5BFE5AFFC47F02FBF1146FDC4E" box="[544,609,1026,1052]" pageId="6" pageNumber="43">KUL</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE5AFFC47DE4FBDB170BDF3D" pageId="6" pageNumber="43" type="diagnosis">
<paragraph id="9941369EFE5AFFC47DE4FBDB15D1DC38" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
<emphasis id="AB8AEA8CFE5AFFC47DE4FBDB1748DC10" bold="true" box="[198,326,1064,1090]" pageId="6" pageNumber="43">Diagnosis.</emphasis>
Habitus (
<figureCitation id="01C52A1BFE5AFFC47C95FBDA1401DC11" box="[439,527,1065,1091]" captionStart="FIGURES 28 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="6" pageNumber="43">Fig. 28</figureCitation>
). Body length 1.71.8 mm. Body slightly drop-shaped, tip of elytron truncate.
<collectingCountry id="E1E9760EFE5AFFC47DB5FBA316DCDC38" box="[151,210,1104,1130]" name="India" pageId="6" pageNumber="43">Hind</collectingCountry>
angle of pronotum produced backwards, acute and needle-shaped.
</paragraph>
<paragraph id="9941369EFE5AFFC47DE4FB8512A2DD79" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
Color. Head and pronotum yellow; base of pronotum usually with a thin dark band. Elytron blackish with contrasting yellow marks: a subbasal transverse band; a small patch on the sutural angle; and an apical mark which is expanded forwards along the margin to the middle of the elytron. The subbasal band reaches the margin and the anterior angle of elytron. Sometimes, the subbasal band and the apical patch fuse marginally in the middle of elytron (
<figureCitation id="01C52A1BFE5AFFC47CBFFAE217FBDD79" box="[413,501,1297,1323]" captionStart="FIGURES 28 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="6" pageNumber="43">Fig. 28</figureCitation>
). Appendages of body and venter yellowish to ferruginous.
</paragraph>
<paragraph id="9941369EFE5AFFC47DE4FACB14EEDD94" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
Sculpture. The entire surface of the beetle apparently with a faint MR or shagreenation (
<date id="ED40105EFE5AFFC4783CFACB135EDD00" box="[1310,1360,1336,1362]" pageId="6" pageNumber="43" value="1940-10">40x</date>
). Few punctures visible on head and metaventrite, more densely punctate on elytron (
<figureCitation id="01C52A1BFE5AFFC47914FAAD1281DD2A" box="[1078,1167,1374,1400]" captionStart="FIGURES 17 25" captionStartId="4.[151,270,1905,1929]" captionTargetBox="[163,1426,187,1879]" captionTargetId="figure@4.[163,1428,187,1880]" captionTargetPageId="4" captionText="FIGURES 17 25. SEMs of MR in some Laccodytes species: 17) detail of elytron of L. americanus (magn. 2100 x); 18) idem, (magn. 2600 x); 19) metacoxa (MTX) and metaventrite (MTS) of L. americanus; 20) idem, detail at higher magnification; 21) idem, detail of metaventrite (MTX) at higher magnification; 22) detail of elytron of L. apalodes; 23) detail of metacoxa of L. apalodes; 24) idem, detail at higher magnification; 25) detail of metacoxa of L. neblinae." httpUri="https://zenodo.org/record/193409/files/figure.png" pageId="6" pageNumber="43">Fig. 22</figureCitation>
). No longitudinal lines or longish meshes visible. Metacoxal shagreenation consisting of network of impressed lines densely packed with what appear to be tiny sensillae (
<figureCitation id="01C52A1BFE5AFFC47F70FA5F14D6DD94" box="[594,728,1452,1478]" captionStart="FIGURES 17 25" captionStartId="4.[151,270,1905,1929]" captionTargetBox="[163,1426,187,1879]" captionTargetId="figure@4.[163,1428,187,1880]" captionTargetPageId="4" captionText="FIGURES 17 25. SEMs of MR in some Laccodytes species: 17) detail of elytron of L. americanus (magn. 2100 x); 18) idem, (magn. 2600 x); 19) metacoxa (MTX) and metaventrite (MTS) of L. americanus; 20) idem, detail at higher magnification; 21) idem, detail of metaventrite (MTX) at higher magnification; 22) detail of elytron of L. apalodes; 23) detail of metacoxa of L. apalodes; 24) idem, detail at higher magnification; 25) detail of metacoxa of L. neblinae." httpUri="https://zenodo.org/record/193409/files/figure.png" pageId="6" pageNumber="43">Figs 23, 24</figureCitation>
)
</paragraph>
<paragraph id="9941369EFE5AFFC47DE4FA211407DED5" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
Structures. Pronotum with narrow lateral bead; posterior angles produced backwards, needle-shaped, acute. Prosternum and base of prosternal process with small ridge; prosternal process broadly carinate and with a long, needle-like, acute tip. Epipleuron broad up to level of sternite 6 or 7. Metatarsomeres 14 with apicolateral angle distinctly lobed (
<figureCitation id="01C52A1BFE5AFFC47F19F9B5149ADE32" box="[571,660,1606,1632]" captionStart="FIGURES 3 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="6" pageNumber="43">Fig. 12</figureCitation>
).
<collectingCountry id="E1E9760EFE5AFFC47F88F9B514EBDE32" box="[682,741,1606,1632]" name="India" pageId="6" pageNumber="43">Hind</collectingCountry>
lobes of metacoxal process rather straight, with a deep Vshaped medial incision (
<figureCitation id="01C52A1BFE5AFFC47C91F99E17F5DED5" box="[435,507,1645,1671]" captionStart="FIGURES 3 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="6" pageNumber="43">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="9941369EFE5AFFC47DE4F967143EDF70" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
Male. Pro- and mesotarsi slightly dilated.
<collectingCountry id="E1E9760EFE5AFFC47F8CF96714E7DEFC" box="[686,745,1684,1710]" name="India" pageId="6" pageNumber="43">Hind</collectingCountry>
margin of sternite 7 rounded (
<figureCitation id="01C52A1BFE5AFFC4796AF96712AEDEFC" box="[1096,1184,1684,1710]" captionStart="FIGURES 54 61" captionStartId="12.[151,273,1107,1131]" captionTargetBox="[153,1431,196,1081]" captionTargetId="figure@12.[151,1436,194,1083]" captionTargetPageId="12" captionText="FIGURES 54 61. Last abdominal segment (left male; right female) in Laccodytes species: 54) L. apalodes; 55) L. rondonia; 56) L. americanus; 57) L. phalacroides; 58) L. takutuanus; 59) L. neblinae; 60) L. bassignanii; 61) Laccodytes sp. 1 female." httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="6" pageNumber="43">Fig. 54</figureCitation>
). Aedeagus (
<figureCitation id="01C52A1BFE5AFFC47814F9671381DEFC" box="[1334,1423,1684,1710]" captionStart="FIGURES 44 47" captionStartId="10.[151,270,1823,1847]" captionTargetBox="[180,1399,210,1765]" captionTargetId="figure@10.[171,1415,197,1798]" captionTargetPageId="10" captionText="FIGURES 44 47. Male genitals in Laccodytes species: 44) L. apalodes: a) median lobe of aedeagus in lateral view, b c) right and left parameres; 45) L. rondonia: idem; 46) L. americanus: idem; 47) L. phalacroides: a) apical half of median lobe of aedeagus in dorsal view, b) median lobe in lateral view, c d) right and left parameres;" httpUri="https://zenodo.org/record/193413/files/figure.png" pageId="6" pageNumber="43">Fig. 44</figureCitation>
): median lobe elongate, narrowed at tip, broadened in the middle from which a membranous expansion emerges. Parameres of different size, both with a long apical seta: the right larger and subtriangular, the left elongate and twisted near the base.
</paragraph>
<paragraph id="9941369EFE5AFFC47DE4F8DD170BDF3D" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
Female.
<collectingCountry id="E1E9760EFE5AFFC47C0EF8DD1769DF1A" box="[300,359,1838,1864]" name="India" pageId="6" pageNumber="43">Hind</collectingCountry>
margin of sternite 7 protruding medially, ending with a deep U- or V-shaped emargination (
<figureCitation id="01C52A1BFE5AFFC47DBDF8A616F9DF3D" box="[159,247,1877,1903]" captionStart="FIGURES 54 61" captionStartId="12.[151,273,1107,1131]" captionTargetBox="[153,1431,196,1081]" captionTargetId="figure@12.[151,1436,194,1083]" captionTargetPageId="12" captionText="FIGURES 54 61. Last abdominal segment (left male; right female) in Laccodytes species: 54) L. apalodes; 55) L. rondonia; 56) L. americanus; 57) L. phalacroides; 58) L. takutuanus; 59) L. neblinae; 60) L. bassignanii; 61) Laccodytes sp. 1 female." httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="6" pageNumber="43">Fig. 54</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE5AFFC47DE4F8881209DFC4" box="[198,1031,1915,1942]" pageId="6" pageNumber="43" type="distribution">
<paragraph id="9941369EFE5AFFC47DE4F8881209DFC4" blockId="6.[151,1437,407,2019]" box="[198,1031,1915,1942]" pageId="6" pageNumber="43">
<emphasis id="AB8AEA8CFE5AFFC47DE4F8881751DFC7" bold="true" box="[198,351,1915,1941]" pageId="6" pageNumber="43">Distribution</emphasis>
(
<figureCitation id="01C52A1BFE5AFFC47C4DF88F17C9DFC4" box="[367,455,1916,1942]" captionStart="FIGURE 62" captionStartId="18.[151,255,1941,1965]" captionText="FIGURE 62. Known distribution of Laccodytes species." pageId="6" pageNumber="43">Fig. 62</figureCitation>
). French
<collectingCountry id="E1E9760EFE5AFFC47F16F88F149CDFC4" box="[564,658,1916,1942]" name="Guyana" pageId="6" pageNumber="43">Guyana</collectingCountry>
,
<collectingCountry id="E1E9760EFE5AFFC47FBCF88F14F2DFC4" box="[670,764,1916,1942]" name="Guyana" pageId="6" pageNumber="43">Guyana</collectingCountry>
,
<collectingCountry id="E1E9760EFE5AFFC47E2AF88F1575DFC4" box="[776,891,1916,1942]" name="Suriname" pageId="6" pageNumber="43">Suriname</collectingCountry>
,
<collectingCountry id="E1E9760EFE5AFFC47EA5F88F120DDFC4" box="[903,1027,1916,1942]" name="Venezuela" pageId="6" pageNumber="43">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE5AFFC47DE4F85116E3DFB1" pageId="6" pageNumber="43" type="biology_ecology">
<paragraph id="9941369EFE5AFFC47DE4F85116E3DFB1" blockId="6.[151,1437,407,2019]" pageId="6" pageNumber="43">
<emphasis id="AB8AEA8CFE5AFFC47DE4F8511724DFEE" bold="true" box="[198,298,1954,1980]" pageId="6" pageNumber="43">Biology.</emphasis>
In
<collectingCountry id="E1E9760EFE5AFFC47C76F85117BCDFEE" box="[340,434,1954,1980]" name="Guyana" pageId="6" pageNumber="43">Guyana</collectingCountry>
, the species was collected at blacklight in forest clearing, close to a stream, and in a stream.
</paragraph>
</subSubSection>
</treatment>
</document>