180 lines
23 KiB
XML
180 lines
23 KiB
XML
<document id="EC805D0A8ECDE88EAF4F711C32C94304" ID-CLB-Dataset="27548" ID-DOI="10.11646/zootaxa.4555.2.3" ID-GBIF-Dataset="2373752d-6333-4f9d-9476-6f2e163a67da" ID-ISSN="1175-5326" ID-Zenodo-Dep="2624293" ID-ZooBank="A7CF86E1-C763-4082-B1C2-9B8B66428142" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554272273913" checkinUser="plazi" docAuthor="Magalhães, Wagner F., Rizzo, Alexandra E. & Bailey-Brock, Julie H." docDate="2019" docId="03E5E958FFDC916F62CFFC3FFD80F840" docLanguage="en" docName="zootaxa.4555.2.3.pdf" docOrigin="Zootaxa 4555 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polyophthalmus Quatrefages 1850" docType="treatment" docVersion="13" lastPageNumber="220" masterDocId="FFDC9120FFD791646258FF8FFFD1FFCE" masterDocTitle="Opheliidae (Annelida: Polychaeta) from the western Pacific islands, including five new species" masterLastPageNumber="235" masterPageNumber="209" pageNumber="220" updateTime="1698716406436" updateUser="ExternalLinkService">
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<mods:mods id="6DC4B33E1229ADF1B349ACE49C1B0156" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="19AA916C512EEDC554AFB65D39CB5A7E">
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<mods:title id="177AC8FE280783F8174264F0944CCF57">Opheliidae (Annelida: Polychaeta) from the western Pacific islands, including five new species</mods:title>
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<mods:namePart id="7204847EA7ABE9A092B444487B8BA0AE">Magalhães, Wagner F.</mods:namePart>
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<mods:namePart id="29C7E149D36045B2D3978304EF91BD14">Rizzo, Alexandra E.</mods:namePart>
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<mods:name id="4757E412B42310E874BF1919C9E93D2A" type="personal">
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<mods:namePart id="97DE325D8BE22E6EE08724A8E09F92A8">Bailey-Brock, Julie H.</mods:namePart>
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<mods:typeOfResource id="6A708CFE92491AD95CE0B471992484EC">text</mods:typeOfResource>
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<mods:title id="9A84CB5C13646C06AA12808265355AA6">Zootaxa</mods:title>
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<mods:part id="57C8AE71702722FCE82E033EB4099C52">
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<mods:date id="0C3D6D4E06FC454C497A1065C8D6CF83">2019</mods:date>
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<mods:number id="577D03AFEF01958949CE14F0F924537C">2019-02-13</mods:number>
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<mods:number id="E0B283831BFD3C3C7FC5E74D935801DE">4555</mods:number>
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<mods:identifier id="60B027FF3E09A66EFF8BE63DB84BDB25" type="DOI">10.11646/zootaxa.4555.2.3</mods:identifier>
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<treatment id="03E5E958FFDC916F62CFFC3FFD80F840" ID-DOI="http://doi.org/10.5281/zenodo.5282925" ID-GBIF-Taxon="154533565" ID-Zenodo-Dep="5282925" LSID="urn:lsid:plazi:treatment:03E5E958FFDC916F62CFFC3FFD80F840" httpUri="http://treatment.plazi.org/id/03E5E958FFDC916F62CFFC3FFD80F840" lastPageNumber="220" pageId="11" pageNumber="220">
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<subSubSection id="C3560BC5FFDC916F62CFFC3FFD4DFC04" box="[151,668,943,970]" pageId="11" pageNumber="220" type="nomenclature">
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<paragraph id="8BF3584EFFDC916F62CFFC3FFD4DFC04" blockId="11.[151,668,943,970]" box="[151,668,943,970]" pageId="11" pageNumber="220">
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<heading id="D0BBEF22FFDC916F62CFFC3FFD4DFC04" bold="true" box="[151,668,943,970]" fontSize="11" level="1" pageId="11" pageNumber="220" reason="1">
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<emphasis id="B938845CFFDC916F62CFFC3FFD4DFC04" bold="true" box="[151,668,943,970]" pageId="11" pageNumber="220">
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Genus
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<taxonomicName id="4C4C23CDFFDC916F62B6FC20FD4DFC04" ID-CoL="84SK5" authority="Quatrefages, 1850" authorityName="Quatrefages" authorityYear="1850" box="[238,668,943,970]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="genus">
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<emphasis id="B938845CFFDC916F62B6FC20FE62FC07" bold="true" box="[238,435,943,969]" italics="true" pageId="11" pageNumber="220">Polyophthalmus</emphasis>
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<bibRefCitation id="EFDD25BFFFDC916F63E2FC3FFD4DFC04" author="Quatrefages, A. de" box="[442,668,944,970]" pageId="11" pageNumber="220" pagination="5 - 46" refId="ref13906" refString="Quatrefages, A. de. (1850) Etudes sur les types inferieurs de l'embranchement des Anneles. Memoire sur la famille des Polyphthalmiens (Polyphthalmea Nob.). Annales des Cciences Naturelles, Paris, (series 3), 13, 5 - 46." type="journal article" year="1850">Quatrefages, 1850</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3560BC5FFDC916F62CFFC79FC96FBC2" box="[151,839,1014,1037]" pageId="11" pageNumber="220" type="materials_examined">
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<paragraph id="8BF3584EFFDC916F62CFFC79FC96FBC2" blockId="11.[151,839,1014,1037]" box="[151,839,1014,1037]" pageId="11" pageNumber="220">
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<emphasis id="B938845CFFDC916F62CFFC79FECBFBC2" bold="true" box="[151,282,1014,1036]" pageId="11" pageNumber="220">
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<typeStatus id="54F7E6ECFFDC916F62CFFC79FF1AFBC2" box="[151,203,1014,1036]" pageId="11" pageNumber="220">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="4C4C23CDFFDC916F637FFC77FDFAFBC2" authority="Dujardin, 1839" authorityName="Dujardin" authorityYear="1839" box="[295,555,1014,1037]" class="Polychaeta" family="Opheliidae" genus="Nais" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="picta">
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<emphasis id="B938845CFFDC916F637FFC77FE5CFBC3" box="[295,397,1016,1037]" italics="true" pageId="11" pageNumber="220">Nais picta</emphasis>
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<bibRefCitation id="EFDD25BFFFDC916F63CAFC79FDFAFBC2" author="Dujardin, F." box="[402,555,1014,1036]" pageId="11" pageNumber="220" pagination="287 - 294" refId="ref12745" refString="Dujardin, F. (1839) Observations sur quelques Annelides marines. Annales des Sciences Naturelles, Paris, Ser. 2, 11, 287 - 294." type="journal article" year="1839">Dujardin, 1839</bibRefCitation>
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</taxonomicName>
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, by subsequent designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3560BC5FFDC916F62CFFBB6FD44FAE4" pageId="11" pageNumber="220" type="diagnosis">
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<paragraph id="8BF3584EFFDC916F62CFFBB6FD44FAE4" blockId="11.[151,1437,1081,1934]" pageId="11" pageNumber="220">
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<emphasis id="B938845CFFDC916F62CFFBB6FEC0FB9C" bold="true" box="[151,273,1081,1106]" pageId="11" pageNumber="220">Diagnosis.</emphasis>
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Body elongated, not divided into distinct regions; ventral groove and two lateral grooves present. Segments indistinctly annulated. Prostomium broadly rounded, palpode absent; subdermal eyespots; a pair of large, eversible nuchal organs divided into a ventral and a dorsal region. Branchiae entirely absent. Segmental lateral eyes present between parapodia. Parapodial lobes reduced, notopodial and neuropodial capillaries emerging from body wall. Interparapodial ciliated sensory pits present and a ventral lobe may be present. Noto- and neuropodial lappets absent. Noto- and neuropodial small fascicles of simple capillary chaetae. Pygidium with short anal funnel bearing marginal papillae and a pair of basal papillae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3560BC5FFDC916F629FFABAFD80F840" pageId="11" pageNumber="220" type="discussion">
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<paragraph id="8BF3584EFFDC916F629FFABAFD80F840" blockId="11.[151,1437,1081,1934]" pageId="11" pageNumber="220">
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<emphasis id="B938845CFFDC916F629FFABAFEEAFA80" bold="true" box="[199,315,1333,1358]" pageId="11" pageNumber="220">Remarks.</emphasis>
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<bibRefCitation id="EFDD25BFFFDC916F6319FAB9FE25FA80" author="Hartman, O." box="[321,500,1333,1358]" pageId="11" pageNumber="220" pagination="1 - 628" refId="ref13049" refString="Hartman, O. (1959) Catalogue of the polychaetous annelids of the world. Parts 1 and 2. Allan Hancock Foundation Occasional Paper, 23, 1 - 628." type="journal article" year="1959">Hartman (1959)</bibRefCitation>
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synonymized thirteen
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<taxonomicName id="4C4C23CDFFDC916F60ACFABAFC7BFA80" authorityName="Quatrefages" authorityYear="1850" box="[756,938,1333,1358]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="genus">
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<emphasis id="B938845CFFDC916F60ACFABAFC7BFA80" box="[756,938,1333,1358]" italics="true" pageId="11" pageNumber="220">Polyophthalmus</emphasis>
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</taxonomicName>
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species with
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<taxonomicName id="4C4C23CDFFDC916F6619FAB9FAB2FA80" authority="(Dujardin, 1839)" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[1089,1379,1333,1358]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="pictus">
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<emphasis id="B938845CFFDC916F6619FAB9FB71FA83" box="[1089,1184,1334,1357]" italics="true" pageId="11" pageNumber="220">P. pictus</emphasis>
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(
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<bibRefCitation id="EFDD25BFFFDC916F66F6FABAFA8BFA80" author="Dujardin, F." box="[1198,1370,1333,1358]" pageId="11" pageNumber="220" pagination="287 - 294" refId="ref12745" refString="Dujardin, F. (1839) Observations sur quelques Annelides marines. Annales des Sciences Naturelles, Paris, Ser. 2, 11, 287 - 294." type="journal article" year="1839">Dujardin, 1839</bibRefCitation>
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)
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</taxonomicName>
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. The author only considered
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<taxonomicName id="4C4C23CDFFDC916F63C5FAD5FD1CFABC" authority="Kukenthal, 1887" authorityName="Kukenthal" authorityYear="1887" box="[413,717,1369,1394]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="striatus">
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<emphasis id="B938845CFFDC916F63C5FAD5FDDDFABF" box="[413,524,1370,1393]" italics="true" pageId="11" pageNumber="220">P. striatus</emphasis>
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<bibRefCitation id="EFDD25BFFFDC916F604AFAD6FD1CFABC" author="Kukenthal, W." box="[530,717,1369,1394]" pageId="11" pageNumber="220" pagination="361 - 373" refId="ref13264" refString="Kukenthal, W. (1887) Die Opheliaceen der expedition der Vettore Pisani. Jenaische Zeitschrift fur Naturwissenschaft, 21, 361 - 373." type="journal article" year="1887">Kükenthal, 1887</bibRefCitation>
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</taxonomicName>
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as a valid and distinct species from
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<taxonomicName id="4C4C23CDFFDC916F6606FAD5FB6DFABF" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[1118,1212,1370,1393]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="pictus">
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<emphasis id="B938845CFFDC916F6606FAD5FB6DFABF" box="[1118,1212,1370,1393]" italics="true" pageId="11" pageNumber="220">P. pictus</emphasis>
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</taxonomicName>
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.
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<bibRefCitation id="EFDD25BFFFDC916F6692FAD6FA4DFABC" author="Michaelsen, W." box="[1226,1436,1369,1394]" pageId="11" pageNumber="220" pagination="1 - 23" refId="ref13355" refString="Michaelsen, W. (1892) Polychaeten von Ceylon. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 9 (2), 1 - 23." type="journal article" year="1892">Michaelsen (1892)</bibRefCitation>
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described
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<taxonomicName id="4C4C23CDFFDC916F6353FAF1FE65FA58" authorityName="Michaelsen" authorityYear="1892" box="[267,436,1405,1430]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="longisetosus">
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<emphasis id="B938845CFFDC916F6353FAF1FE65FA58" box="[267,436,1405,1430]" italics="true" pageId="11" pageNumber="220">P. longisetosus</emphasis>
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</taxonomicName>
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based on a specimen collected in the pelagic environment with long capillaries on last five segments and it was most likely an epitoke. The species
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<taxonomicName id="4C4C23CDFFDC916F6113FA2DFAB9FA74" authority="Purschke, Ding & Muller, 1995" authorityName="Purschke, Ding & Muller" authorityYear="1995" box="[843,1384,1441,1466]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="qingdaoensis">
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<emphasis id="B938845CFFDC916F6113FA2DFC2DFA74" box="[843,1020,1441,1466]" italics="true" pageId="11" pageNumber="220">P. qingdaoensis</emphasis>
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<bibRefCitation id="EFDD25BFFFDC916F665CFA2EFAB9FA74" author="Purschke, G. & Ding, Z. & Muller, M. C." box="[1028,1384,1441,1466]" pageId="11" pageNumber="220" pagination="229 - 241" refId="ref13839" refString="Purschke, G., Ding, Z. & Muller, M. C. (1995) Ultrastructural differences as a taxonomic marker: the segmental ocelli of Polyophthalmus pictus and Polyophthalmus qingdaoensis sp. n. (Polychaeta, Opheliidae). Zoomorphology, 115 (4), 229 - 241. https: // doi. org / 10.1007 / BF 00393803" type="journal article" year="1995">Purschke, Ding & Müller, 1995</bibRefCitation>
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</taxonomicName>
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was recently described and distinguished from
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<taxonomicName id="4C4C23CDFFDC916F60DCFA49FD32FA13" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[644,739,1478,1501]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="pictus">
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<emphasis id="B938845CFFDC916F60DCFA49FD32FA13" box="[644,739,1478,1501]" italics="true" pageId="11" pageNumber="220">P. pictus</emphasis>
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</taxonomicName>
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mostly on the basis of the ultrastructure, size, position and number of the segmental eyes (ocelli), and number of anal papillae.
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<bibRefCitation id="EFDD25BFFFDC916F61F2FA65FBABF9CC" author="Sene Silva, G." box="[938,1146,1513,1538]" pageId="11" pageNumber="220" refId="ref14104" refString="Sene Silva, G. (2007) Filogenia de Opheliidae (Annelida: Polychaeta). Unpublished Thesis presented for the degree, Doctor of Sciences, in Zoology, Universidade Federal do Parana, Curitiba, Brazil, 95 pp." type="book" year="2007">Sene Silva (2007)</bibRefCitation>
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, in an unpublished Ph.D. dissertation, examined the
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<typeStatus id="54F7E6ECFFDC916F639DF981FE24F9E8" box="[453,501,1550,1574]" pageId="11" pageNumber="220">type</typeStatus>
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material of
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<taxonomicName id="4C4C23CDFFDC916F6027F981FC1CF9E8" authority="Hartman, 1960" authorityName="Hartman" authorityYear="1960" box="[639,973,1549,1574]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="translucens">
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<emphasis id="B938845CFFDC916F6027F981FCCCF9E8" box="[639,797,1549,1574]" italics="true" pageId="11" pageNumber="220">P. translucens</emphasis>
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Hartman, 1960
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</taxonomicName>
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and suggested it should be synonymized with
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<taxonomicName id="4C4C23CDFFDC916F6280F9BEFD16F984" authority="Stop-Bowitz, 1948" authorityName="Stop-Bowitz" authorityYear="1948" box="[216,711,1585,1610]" class="Polychaeta" family="Opheliidae" genus="Ophelina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="abranchiata">
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<emphasis id="B938845CFFDC916F6280F9BEFE08F984" box="[216,473,1585,1610]" italics="true" pageId="11" pageNumber="220">Ophelina abranchiata</emphasis>
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<bibRefCitation id="EFDD25BFFFDC916F63B0F9BEFD16F984" author="Stop-Bowitz, C." box="[488,711,1585,1610]" pageId="11" pageNumber="220" pagination="3 - 58" refId="ref14147" refString="Stop-Bowitz, C. (1948) Sur les polychetes arctiques des familles des Glyceriens, des Opheliens, des Scalibregmiens et des Flabelligeriens. TromsO Museums Arshefter, 66, 3 - 58." type="journal article" year="1948">Stop-Bowitz, 1948</bibRefCitation>
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</taxonomicName>
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.
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<bibRefCitation id="EFDD25BFFFDC916F6082F9BEFBB3F984" author="Purschke, G. & Ding, Z. & Muller, M. C." box="[730,1122,1585,1610]" pageId="11" pageNumber="220" pagination="229 - 241" refId="ref13839" refString="Purschke, G., Ding, Z. & Muller, M. C. (1995) Ultrastructural differences as a taxonomic marker: the segmental ocelli of Polyophthalmus pictus and Polyophthalmus qingdaoensis sp. n. (Polychaeta, Opheliidae). Zoomorphology, 115 (4), 229 - 241. https: // doi. org / 10.1007 / BF 00393803" type="journal article" year="1995">Purschke, Ding & Müller (1995)</bibRefCitation>
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described
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<taxonomicName id="4C4C23CDFFDC916F66B1F9BEFEF9F9A0" authorityName="Purschke, Ding & Muller" authorityYear="1995" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="qingdaoensis">
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<emphasis id="B938845CFFDC916F66B1F9BEFEF9F9A0" italics="true" pageId="11" pageNumber="220">Polyophthalmus qingdaoensis</emphasis>
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</taxonomicName>
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based on ultrastructural differences of the segmental eyes and discussed that several
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<taxonomicName id="4C4C23CDFFDC916F66B1F9DAFA4DF9A0" authorityName="Quatrefages" authorityYear="1850" box="[1257,1436,1621,1646]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="genus">
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<emphasis id="B938845CFFDC916F66B1F9DAFA4DF9A0" box="[1257,1436,1621,1646]" italics="true" pageId="11" pageNumber="220">Polyophthalmus</emphasis>
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</taxonomicName>
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species synonymized with
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<taxonomicName id="4C4C23CDFFDC916F63E5F9F5FDC8F95F" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[445,537,1658,1681]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="pictus">
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<emphasis id="B938845CFFDC916F63E5F9F5FDC8F95F" box="[445,537,1658,1681]" italics="true" pageId="11" pageNumber="220">P. pictus</emphasis>
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</taxonomicName>
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could actually be valid. Until now, three species are considered as valid:
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<taxonomicName id="4C4C23CDFFDC916F6761F9F5FA44F95F" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[1337,1429,1658,1681]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="pictus">
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<emphasis id="B938845CFFDC916F6761F9F5FA44F95F" box="[1337,1429,1658,1681]" italics="true" pageId="11" pageNumber="220">P. pictus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C4C23CDFFDC916F62CFF911FE93F978" authorityName="Purschke, Ding & Muller" authorityYear="1995" box="[151,322,1693,1718]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="qingdaoensis">
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<emphasis id="B938845CFFDC916F62CFF911FE93F978" box="[151,322,1693,1718]" italics="true" pageId="11" pageNumber="220">P. qingdaoensis</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C4C23CDFFDC916F6325F911FC95F978" authority="(Blake & Maciolek 2016 a)" baseAuthorityName="Blake & Maciolek" baseAuthorityYear="2016" box="[381,836,1693,1718]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="translucens">
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<emphasis id="B938845CFFDC916F6325F911FDC4F978" box="[381,533,1693,1718]" italics="true" pageId="11" pageNumber="220">P. translucens</emphasis>
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(
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<bibRefCitation id="EFDD25BFFFDC916F607BF912FCEDF978" author="Blake, J. A. & Maciolek, N. J." box="[547,828,1693,1718]" pageId="11" pageNumber="220" refId="ref12381" refString="Blake, J. A. & Maciolek, N. J. (2016 a) Opheliidae Malmgren, 1867. In: Westheide, W. & Purschke, G. (Eds), Handbook of Zoology. pp. 10, De Gruyter, Available from: https: // www. degruyter. com / view / Zoology / bp _ 029147 - 6 _ 74 (accessed September 2018)" type="url" year="2016">Blake & Maciolek 2016a</bibRefCitation>
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)
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</taxonomicName>
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. A taxonomic revision of the
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<taxonomicName id="4C4C23CDFFDC916F66DEF911FB30F97B" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[1158,1249,1694,1717]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="11" pageNumber="220" phylum="Annelida" rank="species" species="pictus">
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<emphasis id="B938845CFFDC916F66DEF911FB30F97B" box="[1158,1249,1694,1717]" italics="true" pageId="11" pageNumber="220">P. pictus</emphasis>
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</taxonomicName>
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species complex would be desirable including intra- and interspecific variability but also molecular data, whenever possible. There are very few morphological features and body pigmentation has been the main character used to separate species. However, this feature seems to be variable among populations and the pigmentation can also fade in preserved material. The segmental origin of the lateral eyes and its extent in segments, length of capillaries on anterior and posterior end, number and
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<typeStatus id="54F7E6ECFFDC916F63E5F8DDFE26F8A4" box="[445,503,1874,1898]" pageId="11" pageNumber="220">types</typeStatus>
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of capillaries per parapodium, parapodial structures, and number of pygidial papillae should also be comparatively examined.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |