512 lines
68 KiB
XML
512 lines
68 KiB
XML
<document id="E17E5B85C9CBA91A2C11DAC1F9BDFEC0" ID-CLB-Dataset="32365" ID-DOI="10.11646/zootaxa.4306.4.5" ID-GBIF-Dataset="4554fbae-1865-4f67-8250-ea2df5324672" ID-ISSN="1175-5326" ID-Zenodo-Dep="845161" ID-ZooBank="AC76C958-CA96-48CB-AB4A-5591F20070DF" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503039247289" checkinUser="plazi" docAuthor="Marshall, David C. & Hill, Kathy B. R." docDate="2017" docId="03B94A72EA65FFA2FF5EFF43FCE7592B" docLanguage="en" docName="zootaxa.4306.4.5.pdf" docOrigin="Zootaxa 4306 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cicadettana Marshall & Hill 2017, gen. n." docType="treatment" docUuid="A8B153EA-D5AD-4D58-BA52-D597F0FDAAFC" docUuidSource="ZooBank" docVersion="8" lastPageNumber="547" masterDocId="FF80320AEA63FFA8FFC9FF9EFFCE5D73" masterDocTitle="A new genus for North American Cicadetta species (Hemiptera: Cicadidae)" masterLastPageNumber="550" masterPageNumber="537" pageNumber="543" updateTime="1698466899604" updateUser="ExternalLinkService">
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<mods:mods id="2A554A8E77C470B9A74C75A33D6E45A0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="939B1B6B5292B302A00BD7771610A13F">
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<mods:title id="3E820E78D0627775E30EBD2210BBDDF3">A new genus for North American Cicadetta species (Hemiptera: Cicadidae)</mods:title>
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<mods:namePart id="0688305F52AE88D111EAE487E789CCCB">Marshall, David C.</mods:namePart>
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<mods:name id="1451C761EB1E97D30035252518EF6924" type="personal">
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<mods:roleTerm id="4BDE6169468E5D8865B357D596142191">Author</mods:roleTerm>
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<mods:namePart id="6A7B624A3500F9FD999C847A3710E1C6">Hill, Kathy B. R.</mods:namePart>
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<mods:title id="C2E81AD63BC7DBB43C2BA6916F941231">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="21A6FEA34767073FDAFB17369E653363">
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<mods:date id="4A102065295164A413923CFA9FDC56DD">2017</mods:date>
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<mods:detail id="35BA0AADD49E1365F5C66796AE1180D9" type="volume">
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<mods:number id="8714A20E43A2DB81FBC23C1A33150586">4306</mods:number>
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<mods:number id="5B3A7526A8B2DDBA959FAD7F205ACD52">4</mods:number>
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<mods:identifier id="E41ED5FA61DB95EB70224B5085404200" type="CLB-Dataset">32365</mods:identifier>
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<mods:identifier id="03D9D7000A86272ED4D5084220F1170C" type="DOI">10.11646/zootaxa.4306.4.5</mods:identifier>
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<treatment id="03B94A72EA65FFA2FF5EFF43FCE7592B" ID-DOI="http://doi.org/10.5281/zenodo.6024493" ID-GBIF-Taxon="132785613" ID-Zenodo-Dep="6024493" LSID="urn:lsid:zoobank.org:act:A8B153EA-D5AD-4D58-BA52-D597F0FDAAFC" httpUri="http://treatment.plazi.org/id/03B94A72EA65FFA2FF5EFF43FCE7592B" lastPageId="10" lastPageNumber="547" pageId="6" pageNumber="543">
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<subSubSection id="C30AA8EFEA65FFAEFF5EFF43FE165D84" box="[151,472,221,247]" pageId="6" pageNumber="543" type="nomenclature">
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<paragraph id="8BAFFB64EA65FFAEFF5EFF43FE165D84" blockId="6.[151,472,221,247]" box="[151,472,221,247]" pageId="6" pageNumber="543">
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<heading id="D0E74C08EA65FFAEFF5EFF43FE165D84" bold="true" box="[151,472,221,247]" fontSize="11" level="1" pageId="6" pageNumber="543" reason="1">
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<emphasis id="B9642776EA65FFAEFF5EFF43FE165D84" bold="true" box="[151,472,221,247]" pageId="6" pageNumber="543">
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Genus
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<taxonomicName id="4C1080E7EA65FFAEFF27FF43FEB15D84" ID-CoL="8HGBM" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[238,383,221,247]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9642776EA65FFAEFF27FF43FEB15D84" bold="true" box="[238,383,221,247]" italics="true" pageId="6" pageNumber="543">Cicadettana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2579A0DEA65FFAEFE4FFF43FE165D84" box="[390,472,221,247]" pageId="6" pageNumber="543" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30AA8EFEA65FFAEFF5EFEBAFDD45CB4" pageId="6" pageNumber="543" type="materials_examined">
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<paragraph id="8BAFFB64EA65FFAEFF5EFEBAFD5A5C49" blockId="6.[151,660,292,314]" box="[151,660,292,314]" pageId="6" pageNumber="543">
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<emphasis id="B9642776EA65FFAEFF5EFEBAFEEB5C49" bold="true" box="[151,293,292,314]" pageId="6" pageNumber="543">Type species:</emphasis>
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<taxonomicName id="4C1080E7EA65FFAEFEE2FEBAFD415C49" authority="Walker, 1850: 212" authorityName="Walker" authorityPageNumber="212" authorityYear="1850" box="[299,655,292,314]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="calliope">
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<emphasis id="B9642776EA65FFAEFEE2FEBAFE025C49" box="[299,460,292,314]" italics="true" pageId="6" pageNumber="543">Cicada calliope</emphasis>
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<bibRefCitation id="EF818695EA65FFAEFE1BFEBAFD925C49" author="Walker" box="[466,604,292,314]" pageId="6" pageNumber="543" refString="Walker, F. (1850) List of specimens of Homopterous insects in the collection of the British Museum. British Museum Trustees, London, 260 pp." type="book" year="1850">Walker, 1850</bibRefCitation>
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: 212
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BAFFB64EA65FFAEFF5EFEF8FDD45CB4" blockId="6.[151,1436,358,779]" pageId="6" pageNumber="543">
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<emphasis id="B9642776EA65FFAEFF5EFEF8FEAA5C0C" bold="true" box="[151,356,358,383]" pageId="6" pageNumber="543">Included species:</emphasis>
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<taxonomicName id="4C1080E7EA65FFAEFEA4FEF8FCC95C0C" authority="calliope (Walker 1850: 212)" authorityName="Walker" authorityPageNumber="212" authorityYear="1850" box="[365,775,358,383]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="variety" species="calliope" status="comb. nov." variety="calliope">
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<emphasis id="B9642776EA65FFAEFEA4FEF8FDE75C0C" box="[365,553,358,383]" italics="true" pageId="6" pageNumber="543">calliope calliope</emphasis>
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(
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<bibRefCitation id="EF818695EA65FFAEFDF0FEF8FD305C0C" author="Walker" box="[569,766,358,383]" pageId="6" pageNumber="543" refString="Walker, F. (1850) List of specimens of Homopterous insects in the collection of the British Museum. British Museum Trustees, London, 260 pp." type="book" year="1850">Walker 1850:212</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A2579A0DEA65FFAEFCC6FEF8FCBE5C0C" box="[783,880,358,383]" pageId="6" pageNumber="543">
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<emphasis id="B9642776EA65FFAEFCC6FEF8FCBE5C0C" bold="true" box="[783,880,358,383]" pageId="6" pageNumber="543">comb n.</emphasis>
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</taxonomicNameLabel>
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,
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<emphasis id="B9642776EA65FFAEFCB7FEF8FB995C0C" box="[894,1111,358,383]" italics="true" pageId="6" pageNumber="543">
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<taxonomicName id="4C1080E7EA65FFAEFCB7FEF8FC165C0C" authorityName="Walker" authorityYear="1850" box="[894,984,358,383]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="calliope">calliope</taxonomicName>
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floridensis
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</emphasis>
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||
(
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<bibRefCitation id="EF818695EA65FFAEFBAEFEF8FAD35C0C" author="Davis" box="[1127,1309,358,383]" pageId="6" pageNumber="543" refString="Davis, W. T. (1920) North American cicadas belonging to the genera Platypedia and Melampsalta. Journal of the New York Entomological Society, 28, 95 - 135." type="journal article" year="1920">Davis 1920:131</bibRefCitation>
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)
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<taxonomicNameLabel id="A2579A0DEA65FFAEFAE7FEF8FA5B5C0C" box="[1326,1429,358,383]" pageId="6" pageNumber="543">
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<emphasis id="B9642776EA65FFAEFAE7FEF8FA5B5C0C" bold="true" box="[1326,1429,358,383]" pageId="6" pageNumber="543">comb. n.</emphasis>
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||
</taxonomicNameLabel>
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||
,
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||
<taxonomicName id="4C1080E7EA65FFAEFF5EFE12FE175CD0" authority="(Davis 1920: 134)" authorityName="Marshall & Hill" authorityPageNumber="134" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1920" box="[151,473,394,419]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="camerona" status="comb. nov.">
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<emphasis id="B9642776EA65FFAEFF5EFE12FEC85CD0" box="[151,262,396,419]" italics="true" pageId="6" pageNumber="543">camerona</emphasis>
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||
(
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||
<bibRefCitation id="EF818695EA65FFAEFED0FE14FE1C5CD0" author="Davis" box="[281,466,394,419]" pageId="6" pageNumber="543" refString="Davis, W. T. (1920) North American cicadas belonging to the genera Platypedia and Melampsalta. Journal of the New York Entomological Society, 28, 95 - 135." type="journal article" year="1920">Davis 1920:134</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A2579A0DEA65FFAEFE2CFE14FD815CD0" box="[485,591,394,419]" pageId="6" pageNumber="543">
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||
<emphasis id="B9642776EA65FFAEFE2CFE14FD815CD0" bold="true" box="[485,591,394,419]" pageId="6" pageNumber="543">comb. n.</emphasis>
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||
</taxonomicNameLabel>
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||
,
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||
<taxonomicName id="4C1080E7EA65FFAEFDA8FE14FCB85CD0" authority="(Davis 1919: 340)" authorityName="Marshall & Hill" authorityPageNumber="340" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1919" box="[609,886,394,419]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="kansa" status="comb. nov.">
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<emphasis id="B9642776EA65FFAEFDA8FE14FD6C5CD0" box="[609,674,394,419]" italics="true" pageId="6" pageNumber="543">kansa</emphasis>
|
||
(
|
||
<bibRefCitation id="EF818695EA65FFAEFD7CFE14FCA05CD0" author="Davis" box="[693,878,394,419]" pageId="6" pageNumber="543" refString="Davis, W. T. (1919) A new cicada of the genus Melampsalta. Journal of the New York Entomological Society, 27, 340 - 341." type="journal article" year="1919">Davis 1919:340</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A2579A0DEA65FFAEFC4BFE14FC225CD0" box="[898,1004,394,419]" pageId="6" pageNumber="543">
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||
<emphasis id="B9642776EA65FFAEFC4BFE14FC225CD0" bold="true" box="[898,1004,394,419]" pageId="6" pageNumber="543">comb. n.</emphasis>
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||
</taxonomicNameLabel>
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,
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<taxonomicName id="4C1080E7EA65FFAEFC37FE12FAEE5CD0" authority="(Sanborn 2009: 1)" authorityName="Marshall & Hill" authorityPageNumber="1" authorityYear="2017" baseAuthorityName="Sanborn" baseAuthorityYear="2009" box="[1022,1312,394,419]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="ramosi" status="comb. nov.">
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||
<emphasis id="B9642776EA65FFAEFC37FE12FB825CD0" box="[1022,1100,396,419]" italics="true" pageId="6" pageNumber="543">ramosi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF818695EA65FFAEFB97FE14FAD95CD0" author="Sanborn" box="[1118,1303,394,419]" pageId="6" pageNumber="543" refString="Sanborn, A. F. (2009) A new species of Cicadetta (Hemiptera: Cicadomorpha: Cicadidae) from Hispaniola. Carribean Journal of Science, 45, 1 - 7. https: // doi. org / 10.18475 / cjos. v 45 i 1. a 2" type="journal article" year="2009">Sanborn 2009:1</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A2579A0DEA65FFAEFAE2FE14FA5B5CD0" box="[1323,1429,394,419]" pageId="6" pageNumber="543">
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||
<emphasis id="B9642776EA65FFAEFAE2FE14FA5B5CD0" bold="true" box="[1323,1429,394,419]" pageId="6" pageNumber="543">comb. n.</emphasis>
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||
</taxonomicNameLabel>
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||
,
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||
<taxonomicName id="4C1080E7EA65FFAEFF5EFE31FE635CB4" authority="(Davis 1936: 105)" authorityName="Marshall & Hill" authorityPageNumber="105" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1936" box="[151,429,430,455]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="texana" status="comb. nov.">
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||
<emphasis id="B9642776EA65FFAEFF5EFE31FF2C5CB5" box="[151,226,431,454]" italics="true" pageId="6" pageNumber="543">texana</emphasis>
|
||
(
|
||
<bibRefCitation id="EF818695EA65FFAEFF39FE30FE6B5CB4" author="Davis" box="[240,421,430,455]" pageId="6" pageNumber="543" refString="Davis, W. T. (1936) A remarkable cicada from Mexico and other North American species. Journal of the New York Entomological Society, 44, 101 - 123." type="journal article" year="1936">Davis 1936:105</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2579A0DEA65FFAEFE7DFE30FDD45CB4" box="[436,538,430,455]" pageId="6" pageNumber="543">
|
||
<emphasis id="B9642776EA65FFAEFE7DFE30FDD45CB4" bold="true" box="[436,538,430,455]" pageId="6" pageNumber="543">comb. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30AA8EFEA65FFAEFF0EFE4CFB2D5F40" pageId="6" pageNumber="543" type="etymology">
|
||
<paragraph id="8BAFFB64EA65FFAEFF0EFE4CFECC5F7C" blockId="6.[151,1436,358,779]" pageId="6" pageNumber="543">
|
||
<emphasis id="B9642776EA65FFAEFF0EFE4CFE805C98" bold="true" box="[199,334,466,491]" pageId="6" pageNumber="543">Etymology.</emphasis>
|
||
The stem "
|
||
<taxonomicName id="4C1080E7EA65FFAEFE1EFE4CFD885C98" authorityName="Kolenati" authorityYear="1857" box="[471,582,466,491]" class="Insecta" family="Cicadidae" genus="Cicadetta" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="genus">Cicadetta</taxonomicName>
|
||
" followed by the letters "n" and "a" referring to North America. Gender feminine.
|
||
</paragraph>
|
||
<paragraph id="8BAFFB64EA65FFAEFF0EFD84FB2D5F40" blockId="6.[151,1436,358,779]" box="[199,1251,538,563]" pageId="6" pageNumber="543">
|
||
<emphasis id="B9642776EA65FFAEFF0EFD84FE4B5F40" bold="true" box="[199,389,538,563]" pageId="6" pageNumber="543">Zoobank LSID:</emphasis>
|
||
urn:lsid:zoobank.org:act:A8B153EA-D5AD-4D58-BA52-D597F0FDAAFC
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30AA8EFEA65FFAEFF0EFDA0FDEC5FB0" pageId="6" pageNumber="543" type="distribution">
|
||
<paragraph id="8BAFFB64EA65FFAEFF0EFDA0FDEC5FB0" blockId="6.[151,1436,358,779]" pageId="6" pageNumber="543">
|
||
<emphasis id="B9642776EA65FFAEFF0EFDA0FEAE5F24" bold="true" box="[199,352,574,599]" pageId="6" pageNumber="543">Distribution.</emphasis>
|
||
The eastern and central
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<collectingCountry id="F307BBF4EA65FFAEFDBDFDA1FD615F24" box="[628,687,575,599]" name="United States of America" pageId="6" pageNumber="543">USA</collectingCountry>
|
||
, and Hispaniola in the Greater Antilles (
|
||
<bibRefCitation id="EF818695EA65FFAEFBBDFDA0FAD85F24" author="Sanborn" box="[1140,1302,574,599]" pageId="6" pageNumber="543" refString="Sanborn, A. F. (2009) A new species of Cicadetta (Hemiptera: Cicadomorpha: Cicadidae) from Hispaniola. Carribean Journal of Science, 45, 1 - 7. https: // doi. org / 10.18475 / cjos. v 45 i 1. a 2" type="journal article" year="2009">Sanborn 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF818695EA65FFAEFAEBFDA0FEFE5F08" author="Sanborn" pageId="6" pageNumber="543" refString="Sanborn, A. F. & Phillips, P. (2013) Biogeography of the cicadas (Hemiptera: Cicadidae) of North America, north of Mexico. Diversity, 5, 166 - 239. https: // doi. org / 10.3390 / d 5020166" type="journal article" year="2013">Sanborn & Phillips 2013</bibRefCitation>
|
||
).
|
||
<collectingCountry id="F307BBF4EA65FFAEFE8BFDFDFEB45F08" box="[322,378,611,635]" name="United States of America" pageId="6" pageNumber="543">USA</collectingCountry>
|
||
records extend to the foothills of the Rocky Mountains but exclude New
|
||
<collectingCountry id="F307BBF4EA65FFAEFB63FDFCFAC45F08" box="[1194,1290,610,635]" name="United Kingdom" pageId="6" pageNumber="543">England</collectingCountry>
|
||
, the northern Great Lakes region, and the northern Great Plains.
|
||
<taxonomicName id="4C1080E7EA65FFAEFD13FD18FC145FED" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1920" box="[730,986,646,671]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="camerona">
|
||
<emphasis id="B9642776EA65FFAEFD13FD18FC145FED" box="[730,986,646,671]" italics="true" pageId="6" pageNumber="543">Cicadettana camerona</emphasis>
|
||
</taxonomicName>
|
||
has a restricted distribution in southern
|
||
<collectingRegion id="49D43586EA65FFAEFF5EFD35FF165FB1" box="[151,216,683,706]" country="United States of America" name="Texas" pageId="6" pageNumber="543">Texas</collectingRegion>
|
||
but may extend into
|
||
<collectingRegion id="49D43586EA65FFAEFE0CFD34FDD15FB0" box="[453,543,682,707]" country="Mexico" name="Mexico" pageId="6" pageNumber="543">
|
||
<collectingCountry id="F307BBF4EA65FFAEFE0CFD34FDD15FB0" box="[453,543,682,707]" name="Mexico" pageId="6" pageNumber="543">Mexico</collectingCountry>
|
||
</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30AA8EFEA65FFA0FF0EFD50FE1E5F80" lastPageId="8" lastPageNumber="545" pageId="6" pageNumber="543" type="diagnosis">
|
||
<paragraph id="8BAFFB64EA65FFAEFF0EFD50FB765E78" blockId="6.[151,1436,358,779]" pageId="6" pageNumber="543">
|
||
<emphasis id="B9642776EA65FFAEFF0EFD50FD8C5F94" bold="true" box="[199,578,718,743]" pageId="6" pageNumber="543">
|
||
Diagnosis (see also
|
||
<figureCitation id="132BE7E1EA65FFAEFE75FD50FDDE5F94" box="[444,528,718,743]" captionStart="FIGURE 2" captionStartId="7.[151,250,1535,1557]" captionTargetBox="[150,1436,193,1514]" captionTargetId="figure@7.[150,1436,193,1514]" captionTargetPageId="7" captionText="FIGURE 2. Cicadettana calliope calliope (Walker) gen. n., comb. n. male (above) and female (below) specimens collected on 14 June 2007 in Cass Co., Illinois, E. of Chandlerville (Lat. 40.0467, Lon. - 90.1084, WGS 84). Scale bars = 4 mm." httpUri="https://zenodo.org/record/845165/files/figure.png" pageId="6" pageNumber="543">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="132BE7E1EA65FFAEFDEAFD50FDFC5F94" box="[547,562,718,743]" captionStart="FIGURE 3" captionStartId="9.[151,250,1690,1712]" captionTargetBox="[152,1434,193,1668]" captionTargetId="figure@9.[152,1434,193,1668]" captionTargetPageId="9" captionText="FIGURE 3. Cicadettana calliope calliope (Walker) gen. n., comb. n. Upper left, male pygofer in lateral view. Upper right, same image labelled and with dotted lines emphasizing margins of key features including one clasper, both pseudoparameres, upper margin of one upper pygofer lobe, upper margin of one basal pygofer lobe, and thecal ventral support. The endotheca is obscured but lies between the ventral support and the pseudoparameres and does not extend beyond the claspers. Lower left, pygofer in ventral view, labelled; thecal structures are not in focus. Lower right, male timbal. BL = basal pygofer lobe, CL = clasper, DB = dorsal beak, PS = pseudoparameres, UL = upper pygofer lobe, UN = uncus (median lobe), VS = ventral support. Scale bars = 0.5 mm. Specimen is G 133 genitalia prep. from US. KS. GYP site (see Table 1)." httpUri="https://zenodo.org/record/845167/files/figure.png" pageId="6" pageNumber="543">3</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Small cicadas (
|
||
<emphasis id="B9642776EA65FFAEFCCDFD51FCD15F95" box="[772,799,719,742]" italics="true" pageId="6" pageNumber="543">ca</emphasis>
|
||
.
|
||
<quantity id="4CE85681EA65FFAEFCFAFD51FC7F5F95" box="[819,945,719,743]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="6" pageNumber="543" unit="mm" value="16.0" valueMax="20.0" valueMin="12.0">12–20 mm</quantity>
|
||
length, excluding wings) with narrow to moderate-width bodies and primarily brown and/or green color with only limited black marks.
|
||
</paragraph>
|
||
<caption id="DF6FABECEA65FFAEFF5EF8B6FB6A5AA4" httpUri="https://zenodo.org/record/845163/files/figure.png" pageId="6" pageNumber="543" targetBox="[151,1434,865,1805]" targetPageId="6">
|
||
<paragraph id="8BAFFB64EA65FFAEFF5EF8B6FB6A5AA4" blockId="6.[151,1436,1832,2007]" pageId="6" pageNumber="543">
|
||
<emphasis id="B9642776EA65FFAEFF5EF8B6FEDB5A4D" bold="true" box="[151,277,1832,1854]" pageId="6" pageNumber="543">FIGURE 1.</emphasis>
|
||
A section of the molecular phylogeny of the tribe
|
||
<taxonomicName id="4C1080E7EA65FFAEFCE4F8B6FC525A4D" authorityName="Buckton" authorityYear="1889" box="[813,924,1832,1854]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
|
||
published by Marshall
|
||
<emphasis id="B9642776EA65FFAEFB5DF8B7FB045A4D" box="[1172,1226,1832,1854]" italics="true" pageId="6" pageNumber="543">et al.</emphasis>
|
||
(2016) showing the North American taxa (formerly in
|
||
<taxonomicName id="4C1080E7EA65FFAEFE3CF8D9FD965A2E" authorityName="Kolenati" authorityYear="1857" box="[501,600,1863,1885]" class="Insecta" family="Cicadidae" genus="Cicadetta" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9642776EA65FFAEFE3CF8D9FD965A2E" box="[501,600,1863,1885]" italics="true" pageId="6" pageNumber="543">Cicadetta</emphasis>
|
||
</taxonomicName>
|
||
, here newly combined in the new genus
|
||
<taxonomicName id="4C1080E7EA65FFAEFC31F8D9FBBA5A2E" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[1016,1140,1863,1885]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA65FFAEFC31F8D9FBBA5A2E" box="[1016,1140,1863,1885]" italics="true" pageId="6" pageNumber="543">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
) and their closest allies from Eurasia and Oceania forming a clade nested within genera from Australia. This clade, termed the R2 clade by Marshall
|
||
<emphasis id="B9642776EA65FFAEFA85F8F9FAB15A08" box="[1356,1407,1893,1916]" italics="true" pageId="6" pageNumber="543">et al.</emphasis>
|
||
, is one of two inferred to have radiated from Australasia within the tribe. The type species of the genus
|
||
<taxonomicName id="4C1080E7EA65FFAEFB60F81AFAC25AE9" authorityName="Kolenati" authorityYear="1857" box="[1193,1292,1924,1946]" class="Insecta" family="Cicadidae" genus="Cicadetta" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9642776EA65FFAEFB60F81AFAC25AE9" box="[1193,1292,1924,1946]" italics="true" pageId="6" pageNumber="543">Cicadetta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C1080E7EA65FFAEFAD5F81BFA5A5AE9" authorityName="Scopoli (Scopoli" authorityYear="1772" box="[1308,1428,1925,1946]" class="Insecta" family="Cicadidae" genus="Cicadetta" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="543" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="B9642776EA65FFAEFAD5F81BFA5A5AE9" box="[1308,1428,1925,1946]" italics="true" pageId="6" pageNumber="543">C. montana</emphasis>
|
||
</taxonomicName>
|
||
) was found to cluster in a distantly related group (the R1 clade, not shown) representing a separate, earlier out-of-Australasia radiation. See Marshall
|
||
<emphasis id="B9642776EA65FFAEFE40F85CFE725AA4" box="[393,444,1985,2007]" italics="true" pageId="6" pageNumber="543">et al.</emphasis>
|
||
for the full tree and for details on described and undescribed taxa shown.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6FABECEA64FFAFFF5EFA61FAD35B47" httpUri="https://zenodo.org/record/845165/files/figure.png" pageId="7" pageNumber="544" targetBox="[150,1436,193,1514]" targetPageId="7">
|
||
<paragraph id="8BAFFB64EA64FFAFFF5EFA61FAD35B47" blockId="7.[151,1435,1535,1588]" pageId="7" pageNumber="544">
|
||
<emphasis id="B9642776EA64FFAFFF5EFA61FEDC5B66" bold="true" box="[151,274,1535,1557]" pageId="7" pageNumber="544">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C1080E7EA64FFAFFED1FA61FD6D5B66" authority="(Walker)" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Walker" box="[280,675,1535,1557]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="544" phylum="Arthropoda" rank="subSpecies" species="calliope" status="gen. nov." subSpecies="calliope">
|
||
<emphasis id="B9642776EA64FFAFFED1FA61FD8C5B66" box="[280,578,1535,1557]" italics="true" pageId="7" pageNumber="544">Cicadettana calliope calliope</emphasis>
|
||
(Walker)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2579A0DEA64FFAFFD61FA61FC995B66" box="[680,855,1535,1557]" pageId="7" pageNumber="544">
|
||
<emphasis id="B9642776EA64FFAFFD61FA61FC995B66" bold="true" box="[680,855,1535,1557]" pageId="7" pageNumber="544">
|
||
<taxonomicNameLabel id="A2579A0DEA64FFAFFD61FA61FD3E5B66" box="[680,752,1535,1557]" pageId="7" pageNumber="544" rank="genus">gen. n.</taxonomicNameLabel>
|
||
, comb. n.
|
||
</emphasis>
|
||
</taxonomicNameLabel>
|
||
male (above) and female (below) specimens collected on 14 June 2007 in Cass Co., Illinois, E. of Chandlerville (Lat. 40.0467, Lon. -90.1084, WGS84). Scale bars = 4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAFFB64EA64FFA0FF0EF9FEFE1E5F80" blockId="7.[151,1437,1632,2021]" lastBlockId="8.[151,1437,151,2018]" lastPageId="8" lastPageNumber="545" pageId="7" pageNumber="544">
|
||
<emphasis id="B9642776EA64FFAFFF0EF9FEFECA5B0A" box="[199,260,1632,1657]" italics="true" pageId="7" pageNumber="544">Head</emphasis>
|
||
including eyes as wide as mesonotum or narrower; supra-antennal plate nearly meeting eye; postclypeus broadly rounded transversely across ventral midline (dorsal view), in lateral profile angular at most anterior part.
|
||
<emphasis id="B9642776EA64FFAFFF5EF936FF265BB2" box="[151,232,1704,1729]" italics="true" pageId="7" pageNumber="544">Thorax</emphasis>
|
||
: pronotum in dorsal view parallel-sided or widening towards posterior; pronotal collar width at midline less than diameter of eyes; paranota confluent with adjoining pronotal sclerites (sometimes subtle ampliation in
|
||
<taxonomicName id="4C1080E7EA64FFAFFF5EF96EFE955A7A" authority="calliope" box="[151,347,1776,1801]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="544" phylum="Arthropoda" rank="variety" species="calliope" variety="calliope">
|
||
<emphasis id="B9642776EA64FFAFFF5EF96EFE955A7A" box="[151,347,1776,1801]" italics="true" pageId="7" pageNumber="544">calliope calliope</emphasis>
|
||
</taxonomicName>
|
||
), no mid lateral tooth; cruciform elevation wider than long; epimeral lobe not reaching operculum.
|
||
<emphasis id="B9642776EA64FFAFFED7F888FE515A5F" box="[286,415,1813,1837]" italics="true" pageId="7" pageNumber="544">Fore Wings</emphasis>
|
||
hyaline, with 8 apical cells (rarely 7); subapical cells absent; ulnar cell 3 angled to radial cell; apical cell 8 (or 7 if only seven apical cells) with its maximum height at least 1/2 of its width along the ambient vein; basal cell longer than wide, narrowing distally; costal vein not clearly higher than R+Sc; costa parallel-sided to node or gently widening; costa of male gently and evenly curved; pterostigma absent to partial (incomplete distally); vein CuA bowed so that cubital cell wider than medial cell; veins M and CuA meeting basal cell with their stems completely fused as one; vein RA1 aligned closely with Sc for its length and not diverging in subapical region; vein CuA1 divided by crossvein m-cu so that proximal portion shortest; veins CuP and 1A fused in part; distance between cross veins
|
||
<taxonomicName id="4C1080E7EA6BFFA0FDF7FF5EFDB95DA4" box="[574,631,190,215]" pageId="8" pageNumber="545" rank="subSpecies" subSpecies="and">r and</taxonomicName>
|
||
r-m most often about equal to distance between r-m and m (rarely less than); apical cells 3–6 about equal to or longer than ulnar cells; radial cell length somewhat less than from its apex to wing tip; wing outer margin developed for its total length, never reduced to be confluent with ambient vein.
|
||
<emphasis id="B9642776EA6BFFA0FAADFE98FF175C30" italics="true" pageId="8" pageNumber="545">Hind wings</emphasis>
|
||
with 5–6 apical cells (most commonly six in
|
||
<taxonomicName id="4C1080E7EA6BFFA0FD16FEB4FCF75C30" box="[735,825,298,323]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="calliope">
|
||
<emphasis id="B9642776EA6BFFA0FD16FEB4FCF75C30" box="[735,825,298,323]" italics="true" pageId="8" pageNumber="545">calliope</emphasis>
|
||
</taxonomicName>
|
||
); no infuscation on ambient vein; width of 1
|
||
<taxonomicName id="4C1080E7EA6BFFA0FAF8FEB5FA525C30" box="[1329,1436,298,323]" pageId="8" pageNumber="545" rank="subSpecies" subSpecies="cubital">st cubital</taxonomicName>
|
||
cell at distal end at least twice that of 2nd cubital cell; anal lobe broad with vein 3A curved, long, separated from wing margin; veins RP and M fixed basally.
|
||
<emphasis id="B9642776EA6BFFA0FD63FEEAFCC05CF8" box="[682,782,370,395]" italics="true" pageId="8" pageNumber="545">Fore leg</emphasis>
|
||
usually with four femoral spines, three well-developed; femoral primary spine erect.
|
||
<emphasis id="B9642776EA6BFFA0FE29FE08FD4B5CDC" box="[480,645,406,431]" italics="true" pageId="8" pageNumber="545">Male opercula</emphasis>
|
||
of variable length across species, only sometimes reaching margin of tympanal cavity or beyond, apically broadly rounded, not meeting along medial margins, raised above level of tympanal cavity on their outer halves or so (less so in
|
||
<taxonomicName id="4C1080E7EA6BFFA0FD36FE7EFCA35C84" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1920" box="[767,877,480,503]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="camerona">
|
||
<emphasis id="B9642776EA6BFFA0FD36FE7EFCA35C84" box="[767,877,480,503]" italics="true" pageId="8" pageNumber="545">camerona</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<emphasis id="B9642776EA6BFFA0FC4FFE40FBE15C84" box="[902,1071,478,503]" italics="true" pageId="8" pageNumber="545">Male abdomen</emphasis>
|
||
as wide as or a little wider than thorax; tergites in cross-section with sides straight or weakly convex; epipleurites reflexed ventrally from junction with tergites; tergite 1 narrow along dorsal midline; tergite 2 sometimes wider than tergites 3–7 (but not in
|
||
<taxonomicName id="4C1080E7EA6BFFA0FF5EFDD4FF3F5F10" box="[151,241,586,611]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="calliope">
|
||
<emphasis id="B9642776EA6BFFA0FF5EFDD4FF3F5F10" box="[151,241,586,611]" italics="true" pageId="8" pageNumber="545">calliope</emphasis>
|
||
</taxonomicName>
|
||
); sternites somewhat swollen so as to be visible in lateral view; sternites III–VI in cross-section convex, gently and evenly rounded; sternite VIII about as long as sternite VII (but sternite VIII much longer in
|
||
<taxonomicName id="4C1080E7EA6BFFA0FAD5FDEEFA455FF4" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1920" box="[1308,1419,624,647]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="camerona">
|
||
<emphasis id="B9642776EA6BFFA0FAD5FDEEFA455FF4" box="[1308,1419,624,647]" italics="true" pageId="8" pageNumber="545">camerona</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<emphasis id="B9642776EA6BFFA0FF5EFD0CFF205FD8" box="[151,238,658,683]" italics="true" pageId="8" pageNumber="545">Timbals</emphasis>
|
||
with two long ribs spanning the full height of the timbal and two shorter ribs, with no intermediate ribs; basal dome large; anterior part of timbal mostly occupied by ribs; timbals rarely extending slightly below wing bases; timbal covers absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30AA8EFEA6BFFA2FF0EFD60FAD05C87" lastPageId="10" lastPageNumber="547" pageId="8" pageNumber="545" type="description">
|
||
<paragraph id="8BAFFB64EA6BFFA0FF0EFD60FC215824" blockId="8.[151,1437,151,2018]" pageId="8" pageNumber="545">
|
||
<emphasis id="B9642776EA6BFFA0FF0EFD60FEBD5E64" box="[199,371,766,791]" italics="true" pageId="8" pageNumber="545">Male genitalia.</emphasis>
|
||
Pygofer in ventral view ovoid to sub ovoid in shape, distal portion of upper pygofer lobes not the widest point, not strongly tapered from upper pygofer lobes to base; distal shoulders not developed; upper lobes flat, moderately developed, set well away from dorsal beak, rounded; basal lobes undivided, moderately to weakly developed, rounded in lateral view, abutted against or partly tucked behind pygofer margin; dorsal beak present as a developed apical spine or pointed apex (visible in dorsal view) and a part of chitinized pygofer, more weakly present in
|
||
<taxonomicName id="4C1080E7EA6BFFA0FEDEFC2AFEAC5EB8" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1936" box="[279,354,948,971]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="texana">
|
||
<emphasis id="B9642776EA6BFFA0FEDEFC2AFEAC5EB8" box="[279,354,948,971]" italics="true" pageId="8" pageNumber="545">texana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1080E7EA6BFFA0FE6AFC2AFDDC5EB8" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1920" box="[419,530,948,971]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="camerona">
|
||
<emphasis id="B9642776EA6BFFA0FE6AFC2AFDDC5EB8" box="[419,530,948,971]" italics="true" pageId="8" pageNumber="545">camerona</emphasis>
|
||
</taxonomicName>
|
||
. Uncus small, short, flattened, more or less duck-bill shaped. Claspers well developed, large, dominant, restraining aedeagus; claw-like with cavity lateroventrally, unfused, lacking a rounded, inward-facing swelling on proximal half or so of inner margin, diverging towards distal ends (parallel in
|
||
<taxonomicName id="4C1080E7EA6BFFA0FAEBFC62FAA35960" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1936" box="[1314,1389,1020,1043]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="texana">
|
||
<emphasis id="B9642776EA6BFFA0FAEBFC62FAA35960" box="[1314,1389,1020,1043]" italics="true" pageId="8" pageNumber="545">texana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1080E7EA6BFFA0FF5EFBBEFEC85944" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1920" box="[151,262,1056,1079]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="camerona">
|
||
<emphasis id="B9642776EA6BFFA0FF5EFBBEFEC85944" box="[151,262,1056,1079]" italics="true" pageId="8" pageNumber="545">camerona</emphasis>
|
||
</taxonomicName>
|
||
), their apices not forming the widest dimension of the claspers. Aedeagus with basal plate in lateral view undulated, weakly depressed on dorsal midline, in dorsal view as long as or longer than broad, apically broadened with 'ears', basal portion of basal plate directed forwards away from thecal shaft; junction between theca and basal plate with a functional 'hinge' with a partly chitinous back; thecal shaft nearly straight; pseudoparameres present, dorsal of theca and originating distal of thecal base, in dorsal view parallel or turning in then gradually diverging, in lateral view aligned with thecal shaft for much of their length with proximal half or so in line with ventral support; endotheca exposed, soft, entirely fleshy; endothecal ventral support present, of medium length (no more than about half the length of pseudoparameres); thecal subapical cerci absent; flabellum absent; conjunctival claws absent; vesical opening apical on theca.
|
||
<emphasis id="B9642776EA6BFFA0FD90FAA0FCB65824" box="[601,888,1342,1367]" italics="true" pageId="8" pageNumber="545">Male reproductive system</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
<paragraph id="8BAFFB64EA6BFFA0FF0EFAFCFE2258EC" blockId="8.[151,1437,151,2018]" pageId="8" pageNumber="545">
|
||
<emphasis id="B9642776EA6BFFA0FF0EFAFCFE795808" box="[199,439,1378,1403]" italics="true" pageId="8" pageNumber="545">Female dorsal beak</emphasis>
|
||
with a developed apical spine or pointed apex (visible in dorsal view).
|
||
<emphasis id="B9642776EA6BFFA0FA81FAFCFEB858EC" italics="true" pageId="8" pageNumber="545">Female reproductive system</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
<paragraph id="8BAFFB64EA6BFFA0FF0EFA34FE155B41" blockId="8.[151,1437,151,2018]" pageId="8" pageNumber="545">
|
||
<emphasis id="B9642776EA6BFFA0FF0EFA34FE2958B0" bold="true" box="[199,487,1450,1475]" pageId="8" pageNumber="545">Distinguishing features.</emphasis>
|
||
<taxonomicName id="4C1080E7EA6BFFA0FE3AFA34FDB258B0" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[499,636,1450,1475]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA6BFFA0FE3AFA34FDB258B0" box="[499,636,1450,1475]" italics="true" pageId="8" pageNumber="545">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
species possess the following attributes that place them within the subfamily
|
||
<taxonomicName id="4C1080E7EA6BFFA0FEC4FA50FE525894" authorityName="Buckton" authorityYear="1889" box="[269,412,1486,1511]" class="Insecta" family="Cicadettinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="family">Cicadettinae</taxonomicName>
|
||
, tribe
|
||
<taxonomicName id="4C1080E7EA6BFFA0FE28FA50FD915894" authorityName="Buckton" authorityYear="1889" box="[481,607,1486,1511]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
|
||
, following
|
||
<bibRefCitation id="EF818695EA6BFFA0FD14FA50FC495894" author="Moulds" box="[733,903,1486,1511]" pageId="8" pageNumber="545" refString="Moulds, M. S. (2005) An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum, 57, 375 - 446. https: // doi. org / 10.3853 / j. 0067 - 1975.57.2005.1447" type="journal article" year="2005">Moulds (2005)</bibRefCitation>
|
||
: Pygofer with large ventral claspers dominating the 10th abdominal segment; pygofer upper lobe flat; aedeagus with paired pseudoparameres; timbals not much extending below wing bases.
|
||
</paragraph>
|
||
<paragraph id="8BAFFB64EA6BFFA0FF0EF9A3FEF25A21" blockId="8.[151,1437,151,2018]" pageId="8" pageNumber="545">
|
||
The initial comparison of the six
|
||
<taxonomicName id="4C1080E7EA6BFFA0FD9CF9A3FD105B25" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[597,734,1597,1622]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA6BFFA0FD9CF9A3FD105B25" box="[597,734,1597,1622]" italics="true" pageId="8" pageNumber="545">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
taxa to the
|
||
<typeStatus id="54AB45C6EA6BFFA0FCBBF9A0FC6C5B25" box="[882,930,1598,1622]" pageId="8" pageNumber="545">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C1080E7EA6BFFA0FBE5F9A3FB575B25" authorityName="Kolenati" authorityYear="1857" box="[1068,1177,1597,1622]" class="Insecta" family="Cicadidae" genus="Cicadetta" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9642776EA6BFFA0FBE5F9A3FB575B25" box="[1068,1177,1597,1622]" italics="true" pageId="8" pageNumber="545">Cicadetta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C1080E7EA6BFFA0FB67F9A0FAFB5B26" authorityName="Scopoli (Scopoli" authorityYear="1772" box="[1198,1333,1598,1621]" class="Insecta" family="Cicadidae" genus="Cicadetta" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="B9642776EA6BFFA0FB67F9A0FAFB5B26" box="[1198,1333,1598,1621]" italics="true" pageId="8" pageNumber="545">C. montana</emphasis>
|
||
</taxonomicName>
|
||
) yielded several differences in generic-level characters, confirming the distant phylogenetic relationship indicated by the molecular data. Unlike the
|
||
<taxonomicName id="4C1080E7EA6BFFA0FE21F91BFDBF5BED" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[488,625,1669,1694]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA6BFFA0FE21F91BFDBF5BED" box="[488,625,1669,1694]" italics="true" pageId="8" pageNumber="545">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
species,
|
||
<taxonomicName id="4C1080E7EA6BFFA0FD2EF91BFC085BEE" authorityName="Scopoli (Scopoli" authorityYear="1772" box="[743,966,1669,1694]" class="Insecta" family="Cicadidae" genus="Cicadetta" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="B9642776EA6BFFA0FD2EF91BFC085BEE" box="[743,966,1669,1694]" italics="true" pageId="8" pageNumber="545">Cicadetta montana</emphasis>
|
||
</taxonomicName>
|
||
possesses the following distinguishing attributes: fore wing veins M and Cu with stems meeting at basal cell but commonly not fused (but sometimes briefly fused as in
|
||
<taxonomicName id="4C1080E7EA6BFFA0FEA1F950FDC05B95" box="[360,526,1741,1766]" class="Insecta" family="Cicadidae" genus="Cicadetta" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="species" species="brevipennis">C. brevipennis</taxonomicName>
|
||
, see Fig.
|
||
<quantity id="4CE85681EA6BFFA0FDBEF950FD6D5B95" box="[631,675,1741,1766]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="545" unit="in" value="8.0">8 in</quantity>
|
||
<bibRefCitation id="EF818695EA6BFFA0FD60F953FC4B5B95" author="Hertach" box="[681,901,1741,1766]" pageId="8" pageNumber="545" refString="Hertach, T., Puissant, S., Gogala, M., Trilar, T., Hagmann, R., Baur, H., Kunz, G., Wade, E. J., Loader, S. P., Simon, C. & Nagel, P. (2016) Complex within a complex: Integrative taxonomy reveals hidden diversity in Cicadetta brevipennis (Hemiptera: Cicadidae) and unexpected relationships with a song divergent relative. PLoS ONE, 11, e 0165562. https: // doi. org / 10.1371 / journal. pone. 0165562" type="journal volume" year="2016">
|
||
Hertach
|
||
<emphasis id="B9642776EA6BFFA0FCC0F950FC8F5B95" box="[777,833,1741,1766]" italics="true" pageId="8" pageNumber="545">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
); hind wing with infuscation at the distal end of vein 2A; claspers with their apices widely separated and clearly forming the widest dimension of the claspers; pseudoparameres much longer than theca; and thecal ventral support absent (see
|
||
<bibRefCitation id="EF818695EA6BFFA0FB8FF88BFB105A5D" author="Moulds" box="[1094,1246,1813,1838]" pageId="8" pageNumber="545" refString="Moulds, M. S. (2005) An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum, 57, 375 - 446. https: // doi. org / 10.3853 / j. 0067 - 1975.57.2005.1447" type="journal article" year="2005">Moulds 2005</bibRefCitation>
|
||
and Fig.
|
||
<quantity id="4CE85681EA6BFFA0FA9BF888FA525A5D" box="[1362,1436,1813,1838]" metricMagnitude="0" metricUnit="m" metricValue="3.2512" pageId="8" pageNumber="545" unit="in" value="128.0">128 in</quantity>
|
||
<bibRefCitation id="EF818695EA6BFFA0FF5EF8A7FEFE5A21" author="Moulds" box="[151,304,1849,1874]" pageId="8" pageNumber="545" refString="Moulds, M. S. (2012) A review of the genera of Australian cicadas (Hemiptera: Cicadoidea). Zootaxa, 3287, 1 - 262." type="journal article" year="2012">Moulds 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAFFB64EA6BFFA2FF0EF8C3FAD05C87" blockId="8.[151,1437,151,2018]" lastBlockId="10.[151,1437,151,1112]" lastPageId="10" lastPageNumber="547" pageId="8" pageNumber="545">
|
||
Compared to the remaining
|
||
<taxonomicName id="4C1080E7EA6BFFA0FDC3F8C3FD4B5A05" authorityName="Buckton" authorityYear="1889" box="[522,645,1885,1910]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
|
||
genera studied here,
|
||
<taxonomicName id="4C1080E7EA6BFFA0FCB4F8C3FBC85A05" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[893,1030,1885,1910]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA6BFFA0FCB4F8C3FBC85A05" box="[893,1030,1885,1910]" italics="true" pageId="8" pageNumber="545">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
is distinguishable by the following combination of characters: (1) paranota confluent with adjoining pronotal sclerites (sometimes subtle ampliation in
|
||
<taxonomicName id="4C1080E7EA6BFFA0FF5EF83BFE9D5ACD" authority="calliope" box="[151,339,1957,1982]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="545" phylum="Arthropoda" rank="variety" species="calliope" variety="calliope">
|
||
<emphasis id="B9642776EA6BFFA0FF5EF83BFE9D5ACD" box="[151,339,1957,1982]" italics="true" pageId="8" pageNumber="545">calliope calliope</emphasis>
|
||
</taxonomicName>
|
||
), with no mid-lateral tooth; (2) fore wing radial cell length somewhat less than the distance from its apex to wing tip; (3) timbals with two long ribs spanning the full height of the timbal, usually with two shorter ribs and no intervening ribs; (4) male sternites somewhat swollen so that each is visible in lateral profile (not scored for
|
||
<taxonomicName id="4C1080E7EA69FFA2FF08FF23FEC15DA7" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Sanborn" baseAuthorityYear="2009" box="[193,271,189,212]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="species" species="ramosi">
|
||
<emphasis id="B9642776EA69FFA2FF08FF23FEC15DA7" box="[193,271,189,212]" italics="true" pageId="10" pageNumber="547">ramosi</emphasis>
|
||
</taxonomicName>
|
||
); (5) male sternite VIII about as long as sternite VII (except
|
||
<taxonomicName id="4C1080E7EA69FFA2FC15FF23FB855DA7" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1920" box="[988,1099,189,212]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="species" species="camerona">
|
||
<emphasis id="B9642776EA69FFA2FC15FF23FB855DA7" box="[988,1099,189,212]" italics="true" pageId="10" pageNumber="547">camerona</emphasis>
|
||
</taxonomicName>
|
||
); and (6) claspers curved (in lateral view) and claw-like with cavity lateroventrally (more fang-like in
|
||
<taxonomicName id="4C1080E7EA69FFA2FBDFFF7FFB4B5D8B" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1920" box="[1046,1157,225,248]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="species" species="camerona">
|
||
<emphasis id="B9642776EA69FFA2FBDFFF7FFB4B5D8B" box="[1046,1157,225,248]" italics="true" pageId="10" pageNumber="547">camerona</emphasis>
|
||
</taxonomicName>
|
||
but still curved). This combination of characters is sufficient to distinguish
|
||
<taxonomicName id="4C1080E7EA69FFA2FD25FE9AFCBB5C6E" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[748,885,260,285]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA69FFA2FD25FE9AFCBB5C6E" box="[748,885,260,285]" italics="true" pageId="10" pageNumber="547">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
from all described
|
||
<taxonomicName id="4C1080E7EA69FFA2FB99FE9AFB055C6E" authorityName="Buckton" authorityYear="1889" box="[1104,1227,260,285]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
|
||
genera possessing a trifid aedeagus. To these, however, can be added the following additional external characters examined for the most closely related genera (the R2 clade of Marshall
|
||
<emphasis id="B9642776EA69FFA2FD31FED3FCFE5C16" box="[760,816,332,357]" italics="true" pageId="10" pageNumber="547">et al.</emphasis>
|
||
2016): (7) body color brown and/or green with limited black marks dorsally (as opposed to a dominant black background color in many other genera); (8) wings hyaline with pterostigma absent to partial (sometimes almost complete in
|
||
<taxonomicName id="4C1080E7EA69FFA2FCB1FE0BFC0D5CDF" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1936" box="[888,963,405,428]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="species" species="texana">
|
||
<emphasis id="B9642776EA69FFA2FCB1FE0BFC0D5CDF" box="[888,963,405,428]" italics="true" pageId="10" pageNumber="547">texana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1080E7EA69FFA2FC30FE0AFBF45CDE" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1919" box="[1017,1082,404,429]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="species" species="kansa">
|
||
<emphasis id="B9642776EA69FFA2FC30FE0AFBF45CDE" box="[1017,1082,404,429]" italics="true" pageId="10" pageNumber="547">kansa</emphasis>
|
||
</taxonomicName>
|
||
); (9) fore wing apical cell 8 (or 7 if only seven apical cells) with its maximum height at least 1/2 of its width along the ambient vein (but not in
|
||
<taxonomicName id="4C1080E7EA69FFA2FF5EFE43FF2B5C87" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Sanborn" baseAuthorityYear="2009" box="[151,229,477,500]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="species" species="ramosi">
|
||
<emphasis id="B9642776EA69FFA2FF5EFE43FF2B5C87" box="[151,229,477,500]" italics="true" pageId="10" pageNumber="547">ramosi</emphasis>
|
||
</taxonomicName>
|
||
); (10) male sternites III–VI in cross-section convex and evenly rounded (not scored for
|
||
<taxonomicName id="4C1080E7EA69FFA2FB09FE43FAC05C87" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Sanborn" baseAuthorityYear="2009" box="[1216,1294,477,500]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="species" species="ramosi">
|
||
<emphasis id="B9642776EA69FFA2FB09FE43FAC05C87" box="[1216,1294,477,500]" italics="true" pageId="10" pageNumber="547">ramosi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6FABECEA6AFFA1FF5EF904FBF45A1B" httpUri="https://zenodo.org/record/845167/files/figure.png" pageId="9" pageNumber="546" targetBox="[152,1434,193,1668]" targetPageId="9">
|
||
<paragraph id="8BAFFB64EA6AFFA1FF5EF904FBF45A1B" blockId="9.[151,1437,1690,1896]" pageId="9" pageNumber="546">
|
||
<emphasis id="B9642776EA6AFFA1FF5EF904FEDA5BC3" bold="true" box="[151,276,1690,1712]" pageId="9" pageNumber="546">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C1080E7EA6AFFA1FED4F904FD605BC3" authority="(Walker)" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Walker" box="[285,686,1690,1712]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="546" phylum="Arthropoda" rank="subSpecies" species="calliope" status="gen. nov." subSpecies="calliope">
|
||
<emphasis id="B9642776EA6AFFA1FED4F904FD835BC3" box="[285,589,1690,1712]" italics="true" pageId="9" pageNumber="546">Cicadettana calliope calliope</emphasis>
|
||
(Walker)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2579A0DEA6AFFA1FD7FF904FCA25BC3" box="[694,876,1690,1712]" pageId="9" pageNumber="546">
|
||
<emphasis id="B9642776EA6AFFA1FD7FF904FCA25BC3" bold="true" box="[694,876,1690,1712]" pageId="9" pageNumber="546">
|
||
<taxonomicNameLabel id="A2579A0DEA6AFFA1FD7FF904FCCE5BC3" box="[694,768,1690,1712]" pageId="9" pageNumber="546" rank="genus">gen. n.</taxonomicNameLabel>
|
||
, comb. n.
|
||
</emphasis>
|
||
</taxonomicNameLabel>
|
||
Upper left, male pygofer in lateral view. Upper right, same image labelled and with dotted lines emphasizing margins of key features including one clasper, both pseudoparameres, upper margin of one upper pygofer lobe, upper margin of one basal pygofer lobe, and thecal ventral support. The endotheca is obscured but lies between the ventral support and the pseudoparameres and does not extend beyond the claspers. Lower left, pygofer in ventral view, labelled; thecal structures are not in focus. Lower right, male timbal. BL=basal pygofer lobe, CL=clasper, DB=dorsal beak, PS=pseudoparameres, UL=upper pygofer lobe, UN=uncus (median lobe), VS=ventral support. Scale bars = 0.5 mm. Specimen is G133 genitalia prep. from US.KS.GYP site (see Table 1).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30AA8EFEA69FFA2FF0EFE61FCE7592B" pageId="10" pageNumber="547" type="reference_group">
|
||
<paragraph id="8BAFFB64EA69FFA2FF0EFE61FCE7592B" blockId="10.[151,1437,151,1112]" pageId="10" pageNumber="547">
|
||
<taxonomicName id="4C1080E7EA69FFA2FF0EFE61FE9E5F6B" ID-CoL="8HGBM" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[199,336,511,536]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA69FFA2FF0EFE61FE9E5F6B" box="[199,336,511,536]" italics="true" pageId="10" pageNumber="547">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C1080E7EA69FFA2FE28FE61FCD15F6B" ID-CoL="8HPDV" authority="(Lee et al. 2016)" authorityName="Lee et al." authorityYear="2016" box="[481,799,511,536]" class="Insecta" family="Cicadidae" genus="Philipsalta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9642776EA69FFA2FE28FE61FD925F6B" box="[481,604,511,536]" italics="true" pageId="10" pageNumber="547">Philipsalta</emphasis>
|
||
(
|
||
<bibRefCitation id="EF818695EA69FFA2FDA3FD9EFCD95F6B" author="Lee" box="[618,791,511,536]" pageId="10" pageNumber="547" refString="Lee, Y. J., Marshall, D. C., Mohagan, A. & Hill, K. B. R. (2016) Cicadas (Hemiptera: Cicadidae) from Camiguin of Mindanao Province and Dinagat Island in the Philippines, with a new genus and three new species. Zootaxa, 4097 (2), 187 - 202. https: // doi. org / 10.11646 / zootaxa. 4097.2.2" type="journal article" year="2016">
|
||
Lee
|
||
<emphasis id="B9642776EA69FFA2FD55FD9FFD1A5F6B" box="[668,724,511,536]" italics="true" pageId="10" pageNumber="547">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, a closely related genus in the molecular tree of Marshall
|
||
<emphasis id="B9642776EA69FFA2FF5EFDBBFF1C5F4E" box="[151,210,548,573]" italics="true" pageId="10" pageNumber="547">et al.</emphasis>
|
||
(2016). However,
|
||
<taxonomicName id="4C1080E7EA69FFA2FE64FDBAFDE65F4E" ID-CoL="8HPDV" box="[429,552,548,573]" class="Insecta" family="Cicadidae" genus="Philipsalta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9642776EA69FFA2FE64FDBAFDE65F4E" box="[429,552,548,573]" italics="true" pageId="10" pageNumber="547">Philipsalta</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C1080E7EA69FFA2FD0CFDBAFC805F4E" ID-CoL="8HGBM" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[709,846,548,573]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA69FFA2FD0CFDBAFC805F4E" box="[709,846,548,573]" italics="true" pageId="10" pageNumber="547">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
in having (1) a black body color dorsally, (2) fore wing apical cell 8 (or 7 if only seven) longer than wide, (4) male sternites III–VI sharply convex forming a parabolically bowed V shape; (5) fang-like claspers that are angled at nearly 90 degrees. (
|
||
<taxonomicName id="4C1080E7EA69FFA2FB52FDF2FA525FF7" ID-CoL="V799" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1920" box="[1179,1436,620,645]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="species" species="camerona">
|
||
<emphasis id="B9642776EA69FFA2FB52FDF2FA525FF7" box="[1179,1436,620,645]" italics="true" pageId="10" pageNumber="547">Cicadettana camerona</emphasis>
|
||
</taxonomicName>
|
||
has somewhat fang-like claspers but they are not sharply angled.)
|
||
<taxonomicName id="4C1080E7EA69FFA2FCBFFD11FBCC5FDB" ID-CoL="8HRZ8" authorityName="Puissant" authorityYear="2010" box="[886,1026,655,680]" class="Insecta" family="Cicadidae" genus="Tettigettalna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9642776EA69FFA2FCBFFD11FBCC5FDB" box="[886,1026,655,680]" italics="true" pageId="10" pageNumber="547">Tettigettalna</emphasis>
|
||
</taxonomicName>
|
||
, a European genus closely related to
|
||
<taxonomicName id="4C1080E7EA69FFA2FF5EFD2AFEEE5FBE" ID-CoL="8HGBM" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[151,288,692,717]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA69FFA2FF5EFD2AFEEE5FBE" box="[151,288,692,717]" italics="true" pageId="10" pageNumber="547">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
in the molecular tree, is also similar but can be distinguished by a slightly to substantially ampliate paranotal margin (sometimes with a mid lateral tooth) and three long ribs on the timbal (see also
|
||
<bibRefCitation id="EF818695EA69FFA2FB10FD49FF1D5E67" author="Puissant" pageId="10" pageNumber="547" refString="Puissant, S. & Sueur, J. (2010) A hotspot for Mediterranean cicadas (Insecta: Hemiptera: Cicadidae): new genera, species and songs from southern Spain. Systematics and Biodiversity, 8, 555 - 574. https: // doi. org / 10.1080 / 14772000.2010.532832" type="journal article" year="2010">Puissant & Sueur 2010</bibRefCitation>
|
||
), in addition to its usually black dorsal background color, while
|
||
<taxonomicName id="4C1080E7EA69FFA2FC7EFD62FB8E5E66" ID-CoL="8HGBM" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[951,1088,764,789]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA69FFA2FC7EFD62FB8E5E66" box="[951,1088,764,789]" italics="true" pageId="10" pageNumber="547">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
specimens have at most weak paranotal expansion and four timbal ribs (one
|
||
<taxonomicName id="4C1080E7EA69FFA2FD6CFCBEFCC75E4B" ID-CoL="V79B" authorityName="Marshall & Hill" authorityYear="2017" baseAuthorityName="Davis" baseAuthorityYear="1919" box="[677,777,799,824]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="species" species="kansa">
|
||
<emphasis id="B9642776EA69FFA2FD6CFCBEFCC75E4B" box="[677,777,799,824]" italics="true" pageId="10" pageNumber="547">C. kansa</emphasis>
|
||
</taxonomicName>
|
||
had a tiny fifth rib).
|
||
<taxonomicName id="4C1080E7EA69FFA2FC33FC81FB945E4B" ID-CoL="8HKNR" box="[1018,1114,799,824]" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9642776EA69FFA2FC33FC81FB945E4B" box="[1018,1114,799,824]" italics="true" pageId="10" pageNumber="547">Huechys</emphasis>
|
||
</taxonomicName>
|
||
is easily distinguished from
|
||
<taxonomicName id="4C1080E7EA69FFA2FF5EFCDAFEEE5E2E" ID-CoL="8HGBM" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[151,288,836,861]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA69FFA2FF5EFCDAFEEE5E2E" box="[151,288,836,861]" italics="true" pageId="10" pageNumber="547">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
by its fully colored wings, its fore wing radial cell which is only about half as long as the distance from its apex to the wing tip, and by its long, curving, beak-like tubular claspers on the pygofer.
|
||
<taxonomicName id="4C1080E7EA69FFA2FB0FFCF6FAF55E0C" ID-CoL="8HD7M" authorityName="Distant" authorityYear="1882" box="[1222,1339,872,895]" class="Insecta" family="Cicadidae" genus="Arcystasia" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9642776EA69FFA2FB0FFCF6FAF55E0C" box="[1222,1339,872,895]" italics="true" pageId="10" pageNumber="547">Arcystasia</emphasis>
|
||
</taxonomicName>
|
||
is easily distinguished by its complex and apically reticulated wing venation (
|
||
<bibRefCitation id="EF818695EA69FFA2FC69FC12FBFA5ED7" author="Distant" box="[928,1076,908,933]" pageId="10" pageNumber="547" refString="Distant, W. L. (1882) On some undescribed Cicadidae from the Australian and Pacific Regions. Proceedings of the Zoological Society of London, 1882, 125 - 134." type="proceedings paper" year="1882">Distant 1882</bibRefCitation>
|
||
) in addition to other characters. Some Australian genera related to
|
||
<taxonomicName id="4C1080E7EA69FFA2FDE5FC31FD7B5EBB" ID-CoL="8HGBM" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[556,693,943,968]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA69FFA2FDE5FC31FD7B5EBB" box="[556,693,943,968]" italics="true" pageId="10" pageNumber="547">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
are superficially similar morphologically (e.g.,
|
||
<taxonomicName id="4C1080E7EA69FFA2FB2BFC31FAA75EBB" ID-CoL="8HG4C" box="[1250,1385,943,968]" class="Insecta" family="Cicadidae" genus="Chelapsalta" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9642776EA69FFA2FB2BFC31FAA75EBB" box="[1250,1385,943,968]" italics="true" pageId="10" pageNumber="547">Chelapsalta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1080E7EA69FFA2FF5EFC4BFF255E9F" ID-CoL="8HR4J" box="[151,235,981,1004]" class="Insecta" family="Cicadidae" genus="Simona" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="547" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9642776EA69FFA2FF5EFC4BFF255E9F" box="[151,235,981,1004]" italics="true" pageId="10" pageNumber="547">Simona</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<bibRefCitation id="EF818695EA69FFA2FEC2FC4BFE195E9E" author="Ewart" box="[267,471,980,1005]" pageId="10" pageNumber="547" refString="Ewart, A., Popple, L. W. & Marshall, D. C. (2015) New species of Simona Moulds, 2012 and Chelapsalta Moulds, 2012 cicadas (Cicadidae: Cicadettinae: Cicadettini) from Australia: comparative morphology, songs, behaviour and distributions. Zootaxa, 4001 (1), 1 - 65. https: // doi. org / 10.11646 / zootaxa. 4001.1.1" type="journal article" year="2015">
|
||
Ewart
|
||
<emphasis id="B9642776EA69FFA2FE9EFC4BFE5C5E9E" box="[343,402,980,1005]" italics="true" pageId="10" pageNumber="547">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
), but all can be distinguished with the characters shown in
|
||
<tableCitation id="C692CEDFEA69FFA2FB4AFC4AFB145E9F" box="[1155,1242,980,1005]" captionStart="TABLE 3" captionStartId="11.[125,204,572,593]" captionText="TABLE 3. Character matrix from Moulds (2012) for Cicadettini with a ̎ trifid aedeagus ̎ combined with data obtained here for the new genus Cicadettana (all Cicadettana taxa are comb." pageId="10" pageNumber="547">Table 3</tableCitation>
|
||
. Considering the matrix as a whole, the most important characters are the paranota, the timbal rib number, the dorsal body color, and the shape of the claspers on the pygofer, which in combination most consistently distinguish the
|
||
<taxonomicName id="4C1080E7EA69FFA2FADAFB82FA525946" ID-CoL="8HGBM" authority="Marshall & Hill, 2017" authorityName="Marshall & Hill" authorityYear="2017" box="[1299,1436,1052,1077]" class="Insecta" family="Cicadidae" genus="Cicadettana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9642776EA69FFA2FADAFB82FA525946" box="[1299,1436,1052,1077]" italics="true" pageId="10" pageNumber="547">Cicadettana</emphasis>
|
||
</taxonomicName>
|
||
species, and which were scored for all species in the study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |