245 lines
31 KiB
XML
245 lines
31 KiB
XML
<document id="0FD2F83D99C529AE0AAB772A962BF6B4" ID-CLB-Dataset="25611" ID-DOI="10.11646/zootaxa.4664.2.8" ID-GBIF-Dataset="18d1060a-bb56-4396-87d0-c66979d5059f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3385943" ID-ZooBank="37B4B170-153D-4309-9976-F8D64D80C097" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1567592196170" checkinUser="plazi" docAuthor="Bukejs, Andris, Alekseev, Vitalii I. & Pollock, Darren A." docDate="2019" docId="03B587B51E18FFBAFF747760FA8A5BF1" docLanguage="en" docName="zootaxa.4664.2.8.pdf.imf" docOrigin="Zootaxa 4664 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Waidelotinae Bukejs, Alekseev & Pollock 2019, subfam. nov." docType="treatment" docUuid="F0102E1F-BFED-4809-8C42-E91C8117522E" docUuidSource="ZooBank" docVersion="14" lastPageNumber="264" masterDocId="FF8CFFCD1E19FFB9FFE37055FFE75A06" masterDocTitle="Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location" masterLastPageNumber="273" masterPageNumber="261" pageNumber="262" updateTime="1698754542217" updateUser="ExternalLinkService">
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<mods:mods id="2BBE3B1F4044AD996B8B4783945580AA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0D353D129F16AC116E5E34A00B58D2CE">
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<mods:title id="6FD9A558A0804BBEE94F02AC35AF1D74">Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location</mods:title>
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<mods:name id="835BB0761687405BBD21C36005954848" type="personal">
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<mods:roleTerm id="6CB90BF00A4A81C9AF806903B75E4571">Author</mods:roleTerm>
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<mods:namePart id="6AE881207AE22A748C806AFFF6ADF2DD">Bukejs, Andris</mods:namePart>
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<mods:name id="0CA3B15EAD297F28003A62FC35D883B0" type="personal">
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</mods:role>
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<mods:namePart id="A50F6B5F74D5222AD24A454C359D8B32">Alekseev, Vitalii I.</mods:namePart>
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</mods:name>
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<mods:name id="223502422BB7CD24EBC0162E0351F5DE" type="personal">
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<mods:roleTerm id="6ADA3C2A40B317D8F85CC8A406F9CF1B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="566EA8E427788BA3552C0837F029B64D">Pollock, Darren A.</mods:namePart>
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<mods:typeOfResource id="14AB252ED7986A488C51B562711CB725">text</mods:typeOfResource>
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<mods:title id="B8FCBFE2E8B8096ACD76DE1D402DBCE4">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="29062B42C0777918095AF8B300A3ECFC">
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<mods:date id="2E17678F5C6C9341F8641F545DA718CE">2019</mods:date>
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<mods:number id="6EA64D06AFFEA0397F0C7949A0FE5D06">2019-09-03</mods:number>
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<mods:detail id="E80959FFD02D0146500BE05B962214BA" type="volume">
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<mods:number id="5E792E7ED2FD397AB36ECD1C963975EA">4664</mods:number>
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<mods:detail id="6FD9975FEA31CD75195B15F0F515EFC2" type="issue">
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<mods:number id="FC66C98597D9A85A0C1CE922A1B52F9F">2</mods:number>
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<mods:classification id="DA5AC2307CA618052CD5F060847AF6FD">journal article</mods:classification>
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<mods:identifier id="2AF52462D8DD753870EEDFE88865E651" type="CLB-Dataset">25611</mods:identifier>
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<mods:identifier id="12D24DCF16E92902EAD671C6A17F3330" type="DOI">10.11646/zootaxa.4664.2.8</mods:identifier>
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<mods:identifier id="EFDBDEA68C42E2214164195320CE680F" type="GBIF-Dataset">18d1060a-bb56-4396-87d0-c66979d5059f</mods:identifier>
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<mods:identifier id="98E4FF85383BA6C8CC295C3B4ACF564E" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03B587B51E18FFBAFF747760FA8A5BF1" ID-DOI="http://doi.org/10.5281/zenodo.6462520" ID-GBIF-Taxon="159374699" ID-Zenodo-Dep="6462520" LSID="urn:lsid:zoobank.org:act:F0102E1F-BFED-4809-8C42-E91C8117522E" httpUri="http://treatment.plazi.org/id/03B587B51E18FFBAFF747760FA8A5BF1" lastPageId="3" lastPageNumber="264" pageId="1" pageNumber="262">
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<subSubSection id="C30665281E18FFB8FF747760FC5E5D75" pageId="1" pageNumber="262" type="nomenclature">
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<paragraph id="8BA336A31E18FFB8FF747760FD8E5D49" blockId="1.[151,953,1845,1907]" box="[151,617,1845,1871]" pageId="1" pageNumber="262">
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<heading id="D0EB81CF1E18FFB8FF747760FD8E5D49" bold="true" box="[151,617,1845,1871]" fontSize="11" level="1" pageId="1" pageNumber="262" reason="1">
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<emphasis id="B968EAB11E18FFB8FF747760FD8E5D49" bold="true" box="[151,617,1845,1871]" pageId="1" pageNumber="262">
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Subfamily
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<taxonomicName id="4C1C4D201E18FFB8FEFC7760FE235D49" authority="Bukejs, Alekseev & Pollock, 2019" authorityName="Bukejs, Alekseev & Pollock" authorityYear="2019" box="[287,452,1845,1871]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Waidelotinae">Waidelotinae</taxonomicName>
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<taxonomicNameLabel id="A25B57CA1E18FFB8FE287760FD8E5D49" box="[459,617,1845,1871]" pageId="1" pageNumber="262">subfam. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA336A31E18FFB8FF74770DFC5E5D75" blockId="1.[151,953,1845,1907]" box="[151,953,1880,1907]" pageId="1" pageNumber="262">
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<uri id="FF8D3AA11E18FFB8FF74770DFC5E5D75" box="[151,953,1880,1907]" pageId="1" pageNumber="262">
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urn:lsid:zoobank.org:act:
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<uuid id="FFBA0C761E18FFB8FE44770DFC5E5D75" box="[423,953,1880,1907]" pageId="1" pageNumber="262">F0102E1F-BFED-4809-8C42-E91C8117522E</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665281E18FFB8FF7477CDFCA55DB4" box="[151,834,1944,1970]" pageId="1" pageNumber="262" type="type_taxon">
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<paragraph id="8BA336A31E18FFB8FF7477CDFCA55DB4" blockId="1.[151,834,1944,1970]" box="[151,834,1944,1970]" pageId="1" pageNumber="262">
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<emphasis id="B968EAB11E18FFB8FF7477CDFECA5DB4" bold="true" box="[151,301,1944,1970]" pageId="1" pageNumber="262">
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<typeStatus id="54A788011E18FFB8FF7477CDFF335DB4" box="[151,212,1944,1970]" pageId="1" pageNumber="262">Type</typeStatus>
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genus:
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</emphasis>
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<taxonomicName id="4C1C4D201E18FFB8FED677CDFE515DB4" authority="Bukejs & Alekseev & Pollock, 2019" authorityName="Bukejs & Alekseev & Pollock" authorityYear="2019" box="[309,438,1944,1970]" class="Insecta" family="Pyrochroidae" genus="Waidelotus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B968EAB11E18FFB8FED677CDFE515DB4" box="[309,438,1944,1970]" italics="true" pageId="1" pageNumber="262">Waidelotus</emphasis>
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</taxonomicName>
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<emphasis id="B968EAB11E18FFB8FE5E77CDFDCC5DB4" bold="true" box="[445,555,1944,1970]" pageId="1" pageNumber="262">
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<taxonomicNameLabel id="A25B57CA1E18FFB8FE5E77CDFDCC5DB4" box="[445,555,1944,1970]" pageId="1" pageNumber="262" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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, by present designation
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665281E18FFBBFF74778AFDF65B61" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262" type="diagnosis">
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<paragraph id="8BA336A31E18FFBBFF74778AFDF65B61" blockId="1.[151,1436,2015,2042]" lastBlockId="2.[151,1437,153,2015]" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262">
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<emphasis id="B968EAB11E18FFB8FF74778AFE785DFC" bold="true" box="[151,415,2015,2042]" pageId="1" pageNumber="262">Differential diagnosis.</emphasis>
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Waidelotinae
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<emphasis id="B968EAB11E18FFB8FDA2778AFD3D5DFC" bold="true" box="[577,730,2015,2042]" pageId="1" pageNumber="262">
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<taxonomicNameLabel id="A25B57CA1E18FFB8FDA2778AFD3D5DFC" box="[577,730,2015,2042]" pageId="1" pageNumber="262">subfam. nov.</taxonomicNameLabel>
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</emphasis>
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differs from the other five subfamilies of
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<taxonomicName id="4C1C4D201E18FFB8FB5A778AFAA95DFF" authorityName="Latreille" authorityYear="1806" box="[1209,1358,2015,2041]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="family">Pyrochroidae</taxonomicName>
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by the following combination of features: (1) penultimate tarsomere of all tarsi deeply bilobed, antepenultimate tarsomere of pro- and mesothoracic tarsi slightly bilobed, antepenultimate tarsomere of each metathoracic tarsus slightly widened apically; (2) pronotum laterally margined in basal half; (3) eyes emarginate; (4) pretarsal claws appendiculate; (5) prosternal intercoxal process incompletely separating prothoracic coxae; (6) pronotum with fine posterior submarginal groove; (7) head without distinct constriction behind eyes; (8) posterior pronotal pits absent; and (9) elytral pubescence homogeneous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665281E1BFFBBFF247124FA7F58AA" pageId="2" pageNumber="263" type="discussion">
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<paragraph id="8BA336A31E1BFFBBFF247124FB775BF1" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="263">
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Waidelotinae
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<emphasis id="B968EAB11E1BFFBBFE877125FE195B8D" bold="true" box="[356,510,368,395]" pageId="2" pageNumber="263">
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<taxonomicNameLabel id="A25B57CA1E1BFFBBFE877125FE195B8D" box="[356,510,368,395]" pageId="2" pageNumber="263">subfam. nov.</taxonomicNameLabel>
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</emphasis>
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shares several characters with the extant subfamilies
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<taxonomicName id="4C1C4D201E1BFFBBFB8F7124FB125B8D" authorityName="Abdullah" authorityYear="1964" box="[1132,1269,369,395]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pilipalpinae">Pilipalpinae</taxonomicName>
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,
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<taxonomicName id="4C1C4D201E1BFFBBFAE17124FA8D5B8D" authorityName="Lacordaire" authorityYear="1859" box="[1282,1386,369,395]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pedilinae">Pedilinae</taxonomicName>
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and
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<taxonomicName id="4C1C4D201E1BFFBBFF7471C0FEF45BA9" authorityName="Nikitsky" authorityYear="1986" box="[151,275,405,431]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Tydessinae">Tydessinae</taxonomicName>
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but cannot be placed in any of them as they are presently defined. The full suite of characters exhibited by this adult specimen renders it distinct from any of the extant subfamilies. Therefore, we propose a new subfamily for inclusion of the single Eocene genus. The reasons for such decision are outlined below.
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</paragraph>
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<paragraph id="8BA336A31E1BFFBBFF247254FA7F58AA" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="263">
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<taxonomicName id="4C1C4D201E1BFFBBFF247254FEAA581D" authorityName="Abdullah" authorityYear="1964" box="[199,333,513,539]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pilipalpinae">Pilipalpinae</taxonomicName>
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and the new subfamily share the following characters: terminal maxillary palpomere cultriform, head without distinct constriction behind eyes, pretarsal claws appendiculate. Meanwhile, Waidelotinae
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<emphasis id="B968EAB11E1BFFBBFADE7271FF2F5865" bold="true" pageId="2" pageNumber="263">
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<taxonomicNameLabel id="A25B57CA1E1BFFBBFADE7271FF2F5865" pageId="2" pageNumber="263">subfam. nov.</taxonomicNameLabel>
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</emphasis>
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differs from
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<taxonomicName id="4C1C4D201E1BFFBBFEBF721CFE055865" authorityName="Abdullah" authorityYear="1964" box="[348,482,585,611]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pilipalpinae">Pilipalpinae</taxonomicName>
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in havingthe penultimate tarsomere of each metathoracic tarsus not lobed (bilobed in
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<taxonomicName id="4C1C4D201E1BFFBBFF747238FEC6588E" authorityName="Abdullah" authorityYear="1964" box="[151,289,621,648]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pilipalpinae">Pilipalpinae</taxonomicName>
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), elytral pubescence short and homogenous (mostly dimorphic in
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<taxonomicName id="4C1C4D201E1BFFBBFC1B7238FB65588E" authorityName="Abdullah" authorityYear="1964" box="[1016,1154,621,648]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pilipalpinae">Pilipalpinae</taxonomicName>
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), pronotum laterally margined in basal half (entirely smooth in
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<taxonomicName id="4C1C4D201E1BFFBBFDAB72C4FD3558AA" authorityName="Abdullah" authorityYear="1964" box="[584,722,657,684]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pilipalpinae">Pilipalpinae</taxonomicName>
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), and emarginated compound eyes (eyes entire in
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<taxonomicName id="4C1C4D201E1BFFBBFAE272C4FA6B58AA" authorityName="Abdullah" authorityYear="1964" box="[1281,1420,657,684]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pilipalpinae">Pilipalpinae</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665281E1BFFBAFF2472E0FC685BD5" lastPageId="3" lastPageNumber="264" pageId="2" pageNumber="263" type="description">
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<paragraph id="8BA336A31E1BFFBBFF2472E0FA9C5966" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="263">
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<taxonomicName id="4C1C4D201E1BFFBBFF2472E0FEC858C9" authorityName="Lacordaire" authorityYear="1859" box="[199,303,693,719]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pedilinae">Pedilinae</taxonomicName>
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and the new subfamily share following characters: eyes emarginate, pretarsal claws appendiculate, intercoxal prosternal process incompletely separating prothoracic coxae. Waidelotinae
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<taxonomicNameLabel id="A25B57CA1E1BFFBBFB9B728CFAEC58F5" box="[1144,1291,729,755]" pageId="2" pageNumber="263">subfam. nov.</taxonomicNameLabel>
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differs from
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<taxonomicName id="4C1C4D201E1BFFBBFF7472A8FF185911" authorityName="Lacordaire" authorityYear="1859" box="[151,255,765,791]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pedilinae">Pedilinae</taxonomicName>
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in having its antepenultimate tarsomere of each metathoracic tarsus not lobed (bilobed in
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<taxonomicName id="4C1C4D201E1BFFBBFB0D72A8FABC591E" authorityName="Lacordaire" authorityYear="1859" box="[1262,1371,765,792]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pedilinae">Pedilinae</taxonomicName>
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), pronotum with fine posterior submarginal groove (deep in
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<taxonomicName id="4C1C4D201E1BFFBBFCEC7374FC9B593A" authorityName="Lacordaire" authorityYear="1859" box="[783,892,801,828]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pedilinae">Pedilinae</taxonomicName>
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), pronotum laterally margined in basal half (not margined in
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<taxonomicName id="4C1C4D201E1BFFBBFEC77310FE765966" authorityName="Lacordaire" authorityYear="1859" box="[292,401,837,864]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pedilinae">Pedilinae</taxonomicName>
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), and head without distinct constriction behind eyes (head with constriction in
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<taxonomicName id="4C1C4D201E1BFFBBFAE17310FA975966" authorityName="Lacordaire" authorityYear="1859" box="[1282,1392,837,864]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pedilinae">Pedilinae</taxonomicName>
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).
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</paragraph>
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<paragraph id="8BA336A31E1BFFBBFF24733CFCE45E3E" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="263">
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Waidelotinae
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<emphasis id="B968EAB11E1BFFBBFE82733DFE1E5985" bold="true" box="[353,505,872,899]" pageId="2" pageNumber="263">
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<taxonomicNameLabel id="A25B57CA1E1BFFBBFE82733DFE1E5985" box="[353,505,872,899]" pageId="2" pageNumber="263">subfam. nov.</taxonomicNameLabel>
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</emphasis>
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shares with
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<taxonomicName id="4C1C4D201E1BFFBBFD65733CFCE55985" authorityName="Nikitsky" authorityYear="1986" box="[646,770,873,899]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Tydessinae">Tydessinae</taxonomicName>
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following characters: pronotum laterally margined in basal half, pretarsal claws appendiculate, elytral pubescence homogenous. The newly proposed subfamily differs from
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<taxonomicName id="4C1C4D201E1BFFBBFF7473E4FEF459CD" authorityName="Nikitsky" authorityYear="1986" box="[151,275,945,971]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Tydessinae">Tydessinae</taxonomicName>
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in possessing afine posterior submarginal pronotal groove (absent in
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<taxonomicName id="4C1C4D201E1BFFBBFBCA73E4FB4D59CA" authorityName="Nikitsky" authorityYear="1986" box="[1065,1194,945,972]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Tydessinae">Tydessinae</taxonomicName>
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), mesothoracic coxae conical, head without distinct constriction behind eyes (constricted in
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<taxonomicName id="4C1C4D201E1BFFBBFC597380FBDC59F6" authorityName="Nikitsky" authorityYear="1986" box="[954,1083,981,1008]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Tydessinae">Tydessinae</taxonomicName>
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), posterior pronotal pits absent (present in
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|
<taxonomicName id="4C1C4D201E1BFFBBFEF773ACFE725E12" authorityName="Nikitsky" authorityYear="1986" box="[276,405,1017,1044]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Tydessinae">Tydessinae</taxonomicName>
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), and intercoxal prosternal process incompletely separating prothoracic coxae (in contrast to completely separated prothoracic coxae in
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<taxonomicName id="4C1C4D201E1BFFBBFD957448FD105E3E" authorityName="Nikitsky" authorityYear="1986" box="[630,759,1053,1080]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Tydessinae">Tydessinae</taxonomicName>
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).
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</paragraph>
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<paragraph id="8BA336A31E1BFFBBFF247414FA875EA5" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="263">
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Waidelotinae
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<emphasis id="B968EAB11E1BFFBBFE827415FE1F5E5D" bold="true" box="[353,504,1088,1115]" pageId="2" pageNumber="263">
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<taxonomicNameLabel id="A25B57CA1E1BFFBBFE827415FE1F5E5D" box="[353,504,1088,1115]" pageId="2" pageNumber="263">subfam. nov.</taxonomicNameLabel>
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</emphasis>
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is distinguished from
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<taxonomicName id="4C1C4D201E1BFFBBFD137414FC625E5D" authorityName="Latreille" authorityYear="1806" box="[752,901,1089,1115]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Pyrochroinae">Pyrochroinae</taxonomicName>
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in the appendiculate pretarsal claws, antepenultimate tarsomere of each pro- and mesothoracic tarsus slightly bilobed, pronotum laterally margined in basal half, pronotum with fine posterior submarginal groove, and posterior pronotal margin without narrow, raised bead.
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</paragraph>
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<paragraph id="8BA336A31E1BFFBBFF2474F8FBCD5F35" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="263">
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The new subfamily differs from
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<taxonomicName id="4C1C4D201E1BFFBBFDD974F8FD5C5EC1" authorityName="Lacordaire" authorityYear="1859" box="[570,699,1197,1223]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Agnathinae">Agnathinae</taxonomicName>
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by its emarginate eyes, exposed antennal insertions, filiform antennae, cultriform terminal maxillary palpomere (securiform and slightly expanded in
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<taxonomicName id="4C1C4D201E1BFFBBFBBD7484FB035EEA" authorityName="Lacordaire" authorityYear="1859" box="[1118,1252,1233,1260]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Agnathinae">Agnathinae</taxonomicName>
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), pronotum with fine posterior submarginal groove, prothoracic coxal cavities open posteriorly, trochantins exposed, penultimate and antepenultimate tarsomeres of each pro- and mesothoracic tarsus slightly bilobed.
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</paragraph>
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<paragraph id="8BA336A31E1BFFBBFF247568FC585C71" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="263">
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Additionally,
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<taxonomicName id="4C1C4D201E1BFFBBFE837568FE3C5F51" authority="Bukejs & Alekseev & Pollock, 2019" authorityName="Bukejs & Alekseev & Pollock" authorityYear="2019" box="[352,475,1341,1367]" class="Insecta" family="Pyrochroidae" genus="Waidelotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B968EAB11E1BFFBBFE837568FE3C5F51" box="[352,475,1341,1367]" italics="true" pageId="2" pageNumber="263">Waidelotus</emphasis>
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</taxonomicName>
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<emphasis id="B968EAB11E1BFFBBFE027568FDAD5F51" bold="true" box="[481,586,1341,1367]" pageId="2" pageNumber="263">
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<taxonomicNameLabel id="A25B57CA1E1BFFBBFE027568FDAD5F51" box="[481,586,1341,1367]" pageId="2" pageNumber="263" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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can be distinguished from the fossil genus
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<taxonomicName id="4C1C4D201E1BFFBBFBC47568FA7B5F51" authority="Abdullah, 1965" authorityName="Abdullah" authorityYear="1965" box="[1063,1436,1340,1367]" class="Insecta" family="Pyrochroidae" genus="Palaeopyrochroa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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|
<emphasis id="B968EAB11E1BFFBBFBC47568FB0E5F51" box="[1063,1257,1341,1367]" italics="true" pageId="2" pageNumber="263">Palaeopyrochroa</emphasis>
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|
<bibRefCitation id="EF8D4B521E1BFFBBFB0C7568FA7B5F51" author="Abdullah, M." box="[1263,1436,1340,1367]" pageId="2" pageNumber="263" pagination="38 - 42" refId="ref7114" refString="Abdullah, M. (1965) New Anthicidae and Pyrochroidae (Coleoptera) from the Baltic amber (Oligocene). The Entomologist, 98, 38 - 42." type="journal article" year="1965">Abdullah, 1965</bibRefCitation>
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</taxonomicName>
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(described from Baltic amber) as follows: (1) elytra with homogenous short, semierect setae (in contrast to dimorphic in
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<taxonomicName id="4C1C4D201E1BFFBBFF1175D0FE535F99" authorityName="Abdullah" authorityYear="1965" box="[242,436,1413,1439]" class="Insecta" family="Pyrochroidae" genus="Palaeopyrochroa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAB11E1BFFBBFF1175D0FE535F99" box="[242,436,1413,1439]" italics="true" pageId="2" pageNumber="263">Palaeopyrochroa</emphasis>
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</taxonomicName>
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); (2) filiform antennae (serrate in female of
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|
<taxonomicName id="4C1C4D201E1BFFBBFC2075D0FB625F99" authorityName="Abdullah" authorityYear="1965" box="[963,1157,1413,1439]" class="Insecta" family="Pyrochroidae" genus="Palaeopyrochroa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAB11E1BFFBBFC2075D0FB625F99" box="[963,1157,1413,1439]" italics="true" pageId="2" pageNumber="263">Palaeopyrochroa</emphasis>
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</taxonomicName>
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, and probably pectinate or flabellate in male (
|
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<bibRefCitation id="EF8D4B521E1BFFBBFE6D75FCFDCE5FC5" author="Pollock, D. A." box="[398,553,1448,1475]" pageId="2" pageNumber="263" pagination="563 - 708" refId="ref8524" refString="Pollock, D. A. (1995) Classification, reconstructed pylogeny and geographical history of genera of Pilipalpinae (Coleoptera: Tenebrionoidea: Pyrochroidae). Invertebrate Taxonomy, 9 (4), 563 - 708. https: // doi. org / 10.1071 / IT 9950563" type="journal article" year="1995">Pollock 1995</bibRefCitation>
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)); (3) pronotum with fine posterior submarginal groove (pronotum with deep posterior submarginal groove in
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|
<taxonomicName id="4C1C4D201E1BFFBBFDE37598FD255FE1" authorityName="Abdullah" authorityYear="1965" box="[512,706,1485,1511]" class="Insecta" family="Pyrochroidae" genus="Palaeopyrochroa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAB11E1BFFBBFDE37598FD255FE1" box="[512,706,1485,1511]" italics="true" pageId="2" pageNumber="263">Palaeopyrochroa</emphasis>
|
|
</taxonomicName>
|
|
); (4) pronotum laterally margined in basal half (
|
|
<taxonomicName id="4C1C4D201E1BFFBBFB397598FA7B5FE1" authorityName="Abdullah" authorityYear="1965" box="[1242,1436,1485,1511]" class="Insecta" family="Pyrochroidae" genus="Palaeopyrochroa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B968EAB11E1BFFBBFB397598FA7B5FE1" box="[1242,1436,1485,1511]" italics="true" pageId="2" pageNumber="263">Palaeopyrochroa</emphasis>
|
|
</taxonomicName>
|
|
lacking lateral pronotal margins); (5) penultimate tarsomeres and antepenultimate tarsomeres of each pro- and mesothoracic tarsus slightly bilobed (only penultimate tarsomeres bilobed in
|
|
<taxonomicName id="4C1C4D201E1BFFBBFC287640FB6A5C29" authorityName="Abdullah" authorityYear="1965" box="[971,1165,1557,1583]" class="Insecta" family="Pyrochroidae" genus="Palaeopyrochroa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B968EAB11E1BFFBBFC287640FB6A5C29" box="[971,1165,1557,1583]" italics="true" pageId="2" pageNumber="263">Palaeopyrochroa</emphasis>
|
|
</taxonomicName>
|
|
) (6) pronotal punctation small and elytral punctation finer (pronotal punctation fine and elytral punctation coarser in
|
|
<taxonomicName id="4C1C4D201E1BFFBBFB78766CFABA5C55" authorityName="Abdullah" authorityYear="1965" box="[1179,1373,1593,1619]" class="Insecta" family="Pyrochroidae" genus="Palaeopyrochroa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B968EAB11E1BFFBBFB78766CFABA5C55" box="[1179,1373,1593,1619]" italics="true" pageId="2" pageNumber="263">Palaeopyrochroa</emphasis>
|
|
</taxonomicName>
|
|
), and (7) smaller body size (
|
|
<quantity id="4CE49B461E1BFFBBFE717608FE0E5C71" box="[402,489,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="2" pageNumber="263" unit="mm" value="4.7">4.7 mm</quantity>
|
|
in contrast to
|
|
<quantity id="4CE49B461E1BFFBBFD6A7608FD2D5C71" box="[649,714,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="263" unit="mm" value="6.0">6 mm</quantity>
|
|
in
|
|
<taxonomicName id="4C1C4D201E1BFFBBFD0E7608FC485C71" authorityName="Abdullah" authorityYear="1965" box="[749,943,1629,1655]" class="Insecta" family="Pyrochroidae" genus="Palaeopyrochroa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B968EAB11E1BFFBBFD0E7608FC485C71" box="[749,943,1629,1655]" italics="true" pageId="2" pageNumber="263">Palaeopyrochroa</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BA336A31E1BFFBAFF2476D5FC685BD5" blockId="2.[151,1437,153,2015]" lastBlockId="3.[151,1437,153,503]" lastPageId="3" lastPageNumber="264" pageId="2" pageNumber="263">
|
|
<emphasis id="B968EAB11E1BFFBBFF2476D5FEBE5C9D" bold="true" box="[199,345,1664,1691]" pageId="2" pageNumber="263">Description.</emphasis>
|
|
Body medium-sized, elongate, subparallel-sided, flattened dorsally and slightly convex ventrally; covered with homogeneous setae; punctation dense, small to fine. Head prognathous, without distinct constriction behind eyes; eyes large, prominent, emarginate, widely separated; clypeus transverse, convex; frontoclypeal suture distinct; labrum free, transverse; mandibles with bifid apex; maxillary palpi 4-segmented, with terminal palpomere cultriform; antennae 11-segmented, filiform, moderately long, terminal antennomere short. Antennal insertions exposed. Pronotum transverse, with maximum width in posterior one-fourth, slightly narrower than basal width of elytra, without posterior pits or antebasal sulci; pronotum laterally margined in basal half; posterior margin slightly trisinuate, with fine submarginal groove; posterior angles orthogonal. Scutellar shield trapezoidal, moderately large. Elytra elongate, subparallel-sided, widest in posterior one-third; humeri prominent. Epipleura narrow, extending about to posterior margin of abdominal ventrite 3. Prosternum with anterior margin not produced anteriorly; prosternal process narrow, incompletely separating prothoracic coxae. Prothoracic coxal cavities open posteriorly. Trochantins exposed. Mesothoracic ventrite with longitudinal sharp carina medio-posteriorly, and posterior margin with narrow process, separated from mesothoracic episterna by complete suture. Metathoracic ventrite wide and long; discrimen well developed in posterior 1/2. Legs long and slender. Prothoracic coxae large, conical, projecting. Trochanters heteromeroid. Tibiae long, slender, almost straight, with two small, simple apical spurs of equal length. Penultimate tarsomere of all tarsi deeply bilobed; antepenultimate tarsomere of each prothoracic tarsus slightly bilobed, antepenultimate tarsomere of each mesothoracic tarsus slightly bilobed ventrally, antepenultimate tarsomere of each metathoracic tarsus slightly widened apically. Pretarsal claws symmetrical, appendiculate, with small denticle basally. Abdomen with five visible, freely articulated ventrites.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C30665281E1AFFBAFF247189FA8A5BF1" box="[199,1389,476,503]" pageId="3" pageNumber="264" type="discussion">
|
|
<paragraph id="8BA336A31E1AFFBAFF247189FA8A5BF1" blockId="3.[151,1437,153,503]" box="[199,1389,476,503]" pageId="3" pageNumber="264">
|
|
<emphasis id="B968EAB11E1AFFBAFF247189FE805BF1" bold="true" box="[199,359,476,503]" pageId="3" pageNumber="264">Composition.</emphasis>
|
|
The new subfamily is proposed for a single genus consisting of one extinct Eocene species.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |