185 lines
22 KiB
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185 lines
22 KiB
XML
<document id="03139DF4A1C69828E1A13BAC335AB950" ID-DOI="10.5281/zenodo.176282" ID-GBIF-Dataset="fe89bad2-db11-4058-a77b-59dbc6714d89" ID-ISSN="1175-5326" ID-Zenodo-Dep="176282" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459830473595" checkinUser="plazi" docAuthor="Mwabvu, Tarombera, Hamer, Michelle L. & Slotow, Robert H." docDate="2007" docId="03B587B06D1CFFE0FF54F88C6699FBC4" docLanguage="en" docName="zt01452p023.pdf" docOrigin="Zootaxa 1452" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Bicoxidens" docType="key" docVersion="9" lastPageNumber="18" masterDocId="FF8CFFC86D0CFFF1FFC3FFEE6305FFB5" masterDocTitle="A taxonomic review of the southern African millipede genus, Bicoxidens Attems, 1928 (Diplopoda: Spirostreptida: Spirostreptidae), with the description of three new species and a tentative phylogeny" masterLastPageNumber="23" masterPageNumber="1" pageNumber="17" updateTime="1698222893347" updateUser="plazi">
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<mods:titleInfo id="D29F73865222663AA27B5CE2658AD253">
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<mods:title id="724F350AA47EAA18921D8F207EC7A26D">A taxonomic review of the southern African millipede genus, Bicoxidens Attems, 1928 (Diplopoda: Spirostreptida: Spirostreptidae), with the description of three new species and a tentative phylogeny</mods:title>
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<mods:name id="A316C3B1A605AEF8431D2E90425A7AB7" type="personal">
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<mods:namePart id="E668EDAE2DB1F01A285DF9048AAC9819">Mwabvu, Tarombera</mods:namePart>
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<mods:namePart id="F12A3131A1CAD31AB56AD577D29CD15F">Hamer, Michelle L.</mods:namePart>
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<mods:name id="0AE13B08370A2181330E7AEB900BD679" type="personal">
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<mods:namePart id="F333EFA3242D761E5DE4313152CD8150">Slotow, Robert H.</mods:namePart>
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<mods:typeOfResource id="2DBC8CACC0464C787C218C5856BE3B39">text</mods:typeOfResource>
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<mods:title id="AB1B54189E31E8966260C0479EC78079">Zootaxa</mods:title>
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<mods:part id="9296849BECAAC641E3E9671F04053693">
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<mods:date id="62B55FADE928C5A2426087909A3ACC6A">2007</mods:date>
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<mods:detail id="116E2E66FDCF818F0ABB97CD2042B2E0" type="volume">
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<mods:number id="2B8BE9BFB3C80EBD1D22025836AA1960">1452</mods:number>
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<mods:start id="5C05B8F60FBCAF0A6F427F1AC914C2E4">1</mods:start>
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<mods:classification id="E65AB1ED5526630164D967C0497E3E0A">journal article</mods:classification>
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<mods:identifier id="36AAAB609FC7EF339DE85DD0BF16833C" type="DOI">10.5281/zenodo.176282</mods:identifier>
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<mods:identifier id="904C59FC24E3F4BF9536E8BFADD25073" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4CA8AB356D73105ED45AF7CC6475A67B" type="Zenodo-Dep">176282</mods:identifier>
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<treatment id="03B587B06D1CFFE0FF54F88C6699FBC4" ID-DOI="http://doi.org/10.5281/zenodo.6251151" ID-GBIF-Taxon="119357832" ID-Zenodo-Dep="6251151" LSID="urn:lsid:plazi:treatment:03B587B06D1CFFE0FF54F88C6699FBC4" httpUri="http://treatment.plazi.org/id/03B587B06D1CFFE0FF54F88C6699FBC4" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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<subSubSection id="C306652D6D1CFFE1FF54F88C6127F8C8" box="[151,546,1890,1917]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="8BA336A66D1CFFE1FF54F88C6127F8C8" blockId="16.[151,546,1890,1917]" box="[151,546,1890,1917]" pageId="16" pageNumber="17">
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<heading id="D0EB81CA6D1CFFE1FF54F88C6127F8C8" bold="true" box="[151,546,1890,1917]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<emphasis id="B968EAB46D1CFFE1FF54F88C6127F8C8" bold="true" box="[151,546,1890,1917]" pageId="16" pageNumber="17">
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Key to the species of
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<taxonomicName id="4C1C4D256D1CFFE1FE5FF88D6127F8C8" ID-CoL="7NT8D" box="[412,546,1891,1917]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAB46D1CFFE1FE5FF88D6127F8C8" bold="true" box="[412,546,1891,1917]" italics="true" pageId="16" pageNumber="17">Bicoxidens</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306652D6D1CFFE0FF54F8406699FBC4" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="key">
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<keyStep id="30E82E036D1CFFE0FF54F8406699FF6F" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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<paragraph id="8BA336A66D1CFFE1FF54F840669EF845" blockId="16.[151,1436,1966,2032]" pageId="16" pageNumber="17">
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<keyLead id="30ED95936D1CFFE1FF54F840669EF845" pageId="16" pageNumber="17">
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1. Apical region of telocoxite with a flattened median process that overlaps the opposite one; median leaf with an apical median axe-shaped process; telopodite with a long femoral lobe (
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<figureCitation id="13272A236D1CFFE1FBADF83867C5F845" box="[1134,1216,2006,2032]" captionStart="FIGURE 2. B" captionStartId="5.[151,255,1582,1606]" captionTargetBox="[300,1288,189,1581]" captionTargetId="figure@5.[264,1324,161,1598]" captionTargetPageId="5" captionText="FIGURE 2. B. flavicollis (NMZ / D 232). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176284/files/figure.png" pageId="16" pageNumber="17">Figs. 2</figureCitation>
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a, 5a)...................2
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</keyLead>
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</paragraph>
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<paragraph id="8BA336A66D1DFFE0FF54FF766699FF6F" blockId="17.[151,1437,152,1138]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FF766699FF6F" pageId="17" pageNumber="18">
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- Apical region of telocoxite without a flattened median process; median leaf without apical axe-shaped process; telopodite with a short femoral lobe (
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<figureCitation id="13272A236D1DFFE0FD1CFF2E6022FF6F" box="[735,807,192,218]" captionStart="FIGURE 1. B" captionStartId="3.[151,255,1731,1755]" captionTargetBox="[271,1304,559,1707]" captionTargetId="figure@3.[264,1324,538,1719]" captionTargetPageId="3" captionText="FIGURE 1. B. brincki (MZLU / L 951 / 3645). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176283/files/figure.png" pageId="17" pageNumber="18">Figs 1</figureCitation>
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a, 3a, 4a, 6a, 7a, 8a)...................................................3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E82E036D1DFFE0FF54FF066699FECC" pageId="17" pageNumber="18">
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<paragraph id="8BA336A66D1DFFE0FF54FF066698FE9C" blockId="17.[151,1437,152,1138]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FF066698FE9C" pageId="17" pageNumber="18">
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2. Apical telocoxite with a horizontal lateral process; apically lateral leaf with a small, transverse, ear-shaped aboral projection (
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<figureCitation id="13272A236D1DFFE0FE5EFEFE62F7FE9F" box="[413,498,272,298]" captionStart="FIGURE 5. B" captionStartId="10.[151,255,1389,1413]" captionTargetBox="[271,1299,205,1353]" captionTargetId="figure@10.[264,1324,197,1362]" captionTargetPageId="10" captionText="FIGURE 5. B. grandis (NMSA 8884). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176287/files/figure.png" pageId="17" pageNumber="18">Fig. 5b</figureCitation>
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) ................................................................................................................
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<taxonomicName id="4C1C4D256D1DFFE0FA82FEE16698FE9C" ID-CoL="LRV4" box="[1345,1437,271,297]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="grandis">
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<emphasis id="B968EAB46D1DFFE0FA82FEE16698FE9C" bold="true" box="[1345,1437,271,297]" italics="true" pageId="17" pageNumber="18">grandis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BA336A66D1DFFE0FF54FED66699FECC" blockId="17.[151,1437,152,1138]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FED66699FECC" pageId="17" pageNumber="18">
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- Apical telocoxite without lateral process; apical lateral leaf with a small rounded aboral lobe on distal margin (
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<figureCitation id="13272A236D1DFFE0FEEAFE8E627AFECF" box="[297,383,352,378]" captionStart="FIGURE 2. B" captionStartId="5.[151,255,1582,1606]" captionTargetBox="[300,1288,189,1581]" captionTargetId="figure@5.[264,1324,161,1598]" captionTargetPageId="5" captionText="FIGURE 2. B. flavicollis (NMZ / D 232). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176284/files/figure.png" pageId="17" pageNumber="18">Fig. 2b</figureCitation>
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) .............................................................................................................................
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<taxonomicName id="4C1C4D256D1DFFE0FAE4FEB16699FECC" ID-CoL="5WJRG" box="[1319,1436,351,377]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="flavicollis">
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<emphasis id="B968EAB46D1DFFE0FAE4FEB16699FECC" bold="true" box="[1319,1436,351,377]" italics="true" pageId="17" pageNumber="18">flavicollis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E82E036D1DFFE0FF54FE666699FE47" pageId="17" pageNumber="18">
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<paragraph id="8BA336A66D1DFFE0FF54FE666699FE7C" blockId="17.[151,1437,152,1138]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FE666699FE7C" pageId="17" pageNumber="18">
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3. Apical telocoxite T-shaped, with three median processes and a long lateral process. Middle median process overlaps with the process from the opposite telocoxite (
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<figureCitation id="13272A236D1DFFE0FC4CFE5E60E5FE7F" box="[911,992,432,458]" captionStart="FIGURE 3. B" captionStartId="6.[151,255,1878,1902]" captionTargetBox="[292,1291,736,1847]" captionTargetId="figure@6.[264,1324,697,1868]" captionTargetPageId="6" captionText="FIGURE 3. B. friendi (MRAC 20557). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176285/files/figure.png" pageId="17" pageNumber="18">Figs. 3</figureCitation>
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a, 3b) .....................................
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<taxonomicName id="4C1C4D256D1DFFE0FA88FE416699FE7C" ID-CoL="LRV2" box="[1355,1436,431,457]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="friendi">
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<emphasis id="B968EAB46D1DFFE0FA88FE416699FE7C" bold="true" box="[1355,1436,431,457]" italics="true" pageId="17" pageNumber="18">friendi</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BA336A66D1DFFE0FF54FE366699FE47" blockId="17.[151,1437,152,1138]" box="[151,1436,472,498]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FE366699FE47" box="[151,1436,472,498]" pageId="17" pageNumber="18">- Apical telocoxite not T-shaped, without three median or a long lateral process..........................................4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E82E036D1DFFE0FF54FDEE6699FD27" pageId="17" pageNumber="18">
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<paragraph id="8BA336A66D1DFFE0FF54FDEE6699FDF7" blockId="17.[151,1437,152,1138]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FDEE6699FDF7" pageId="17" pageNumber="18">4. Apical telocoxite not oval. Aboral lateral leaf lobe basally rounded and not produced into a lateral extension................................................................................................................................................................5</keyLead>
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</paragraph>
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<paragraph id="8BA336A66D1DFFE0FF54FDBE6699FD27" blockId="17.[151,1437,152,1138]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FDBE6699FD27" pageId="17" pageNumber="18">
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- Apical telocoxite oval. Aboral lateral leaf lobe basally not rounded and produced into a lateral extension (
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<figureCitation id="13272A236D1DFFE0FF0CFD966217FD27" box="[207,274,632,658]" captionStart="FIGURE 8. B" captionStartId="14.[151,255,1323,1347]" captionTargetBox="[305,1289,386,1305]" captionTargetId="figure@14.[294,1303,376,1316]" captionTargetPageId="14" captionText="FIGURE 8. B. nyathi sp. nov. (NMZ / D 747). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176290/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
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a).......................................................................................................................................
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<taxonomicName id="4C1C4D256D1DFFE0FAD2FD99665BFD24" box="[1297,1374,631,657]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="nyathi" status="sp. nov.">
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<emphasis id="B968EAB46D1DFFE0FAD2FD99665BFD24" bold="true" box="[1297,1374,631,657]" italics="true" pageId="17" pageNumber="18">nyathi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57CF6D1DFFE0FAA7FD966699FD27" box="[1380,1436,632,658]" pageId="17" pageNumber="18" rank="species">sp. n</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E82E036D1DFFE0FF54FD4E6698FD57" pageId="17" pageNumber="18">
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<paragraph id="8BA336A66D1DFFE0FF54FD4E6699FD0F" blockId="17.[151,1437,152,1138]" box="[151,1436,672,698]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FD4E6699FD0F" box="[151,1436,672,698]" pageId="17" pageNumber="18">5. Apical telocoxite with two broadly rounded median processes that may be in touch................................6</keyLead>
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</paragraph>
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<paragraph id="8BA336A66D1DFFE0FF54FD266698FD57" blockId="17.[151,1437,152,1138]" box="[151,1437,712,738]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FD266698FD57" box="[151,1437,712,738]" pageId="17" pageNumber="18">- Apical telocoxite with three narrowly rounded or overlapping median processes.......................................7</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E82E036D1DFFE0FF54FD1E6699FC67" pageId="17" pageNumber="18">
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<paragraph id="8BA336A66D1DFFE0FF54FD1E6699FCEF" blockId="17.[151,1437,152,1138]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FD1E6699FCEF" pageId="17" pageNumber="18">
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6. Apical telocoxite with an overhanging folded lateral process above and not extending beyond knee bend; median processes do not touch; median leaf constricted subapically but without a basal lateral lobe (
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<figureCitation id="13272A236D1DFFE0FAA2FCF663D2FCEF" captionStart="FIGURE 4" captionStartId="8.[151,255,1628,1652]" captionTargetBox="[270,1310,197,1578]" captionTargetId="figure@8.[264,1324,175,1602]" captionTargetPageId="8" captionText="FIGURE 4. Bicoxidens gokwensis sp. nov. (MRAC 20556). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176286/files/figure.png" pageId="17" pageNumber="18">Figs. 4</figureCitation>
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a, 4b) ................................................................................................................................
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<taxonomicName id="4C1C4D256D1DFFE0FB19FCD16653FCEC" ID-CoL="LRV3" box="[1242,1366,831,857]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gokwensis" status="sp. nov.">
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<emphasis id="B968EAB46D1DFFE0FB19FCD16653FCEC" bold="true" box="[1242,1366,831,857]" italics="true" pageId="17" pageNumber="18">gokwensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57CF6D1DFFE0FA9EFCAE6699FCEF" box="[1373,1436,832,858]" pageId="17" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="8BA336A66D1DFFE0FF54FC866699FC67" blockId="17.[151,1437,152,1138]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FC866699FC67" pageId="17" pageNumber="18">
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- Apical telocoxite with a broadly rounded lateral process above and extending beyond knee bend; median processes reciprocally in touch, leaving a window between them; sub-apically (below setose area), median leaf constricted and with a basal lateral lobe (
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<figureCitation id="13272A236D1DFFE0FCF5FC566079FC67" box="[822,892,952,978]" captionStart="FIGURE 6. B" captionStartId="11.[151,255,1638,1662]" captionTargetBox="[264,1305,329,1595]" captionTargetId="figure@11.[264,1324,310,1614]" captionTargetPageId="11" captionText="FIGURE 6. B. matopoensis sp. nov. (NMZ / D 562). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176288/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
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a) ........................................
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<taxonomicName id="4C1C4D256D1DFFE0FB04FC596658FC64" box="[1223,1373,951,977]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="matopoensis" status="sp. nov.">
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<emphasis id="B968EAB46D1DFFE0FB04FC596658FC64" bold="true" box="[1223,1373,951,977]" italics="true" pageId="17" pageNumber="18">matopoensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57CF6D1DFFE0FAA7FC566699FC67" box="[1380,1436,952,978]" pageId="17" pageNumber="18" rank="species">sp. n</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E82E036D1DFFE0FF54FC0E6699FBC4" pageId="17" pageNumber="18">
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<paragraph id="8BA336A66D1DFFE0FF54FC0E6699FB94" blockId="17.[151,1437,152,1138]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FC0E6699FB94" pageId="17" pageNumber="18">
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7. Apical telocoxite bird head-shaped with three median processes, the middle long, beak-like and overlapping the opposite one (
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<figureCitation id="13272A236D1DFFE0FE12FBE6611AFB97" box="[465,543,1032,1058]" captionStart="FIGURE 1. B" captionStartId="3.[151,255,1731,1755]" captionTargetBox="[271,1304,559,1707]" captionTargetId="figure@3.[264,1324,538,1719]" captionTargetPageId="3" captionText="FIGURE 1. B. brincki (MZLU / L 951 / 3645). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176283/files/figure.png" pageId="17" pageNumber="18">Figs. 1</figureCitation>
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a, 1b)...................................................................................................
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<taxonomicName id="4C1C4D256D1DFFE0FA86FBE96699FB94" ID-CoL="68K3H" box="[1349,1436,1031,1057]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brincki">
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<emphasis id="B968EAB46D1DFFE0FA86FBE96699FB94" bold="true" box="[1349,1436,1031,1057]" italics="true" pageId="17" pageNumber="18">brincki</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BA336A66D1DFFE0FF54FBDE6699FBC4" blockId="17.[151,1437,152,1138]" pageId="17" pageNumber="18">
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<keyLead id="30ED95936D1DFFE0FF54FBDE6699FBC4" pageId="17" pageNumber="18">
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- Apical telocoxite not bird head-shaped, with two median processes that do not overlap. (
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<figureCitation id="13272A236D1DFFE0FB11FBDE6612FBFF" box="[1234,1303,1072,1098]" captionStart="FIGURE 7. B" captionStartId="13.[151,255,1629,1653]" captionTargetBox="[280,1313,214,1595]" captionTargetId="figure@13.[264,1324,197,1602]" captionTargetPageId="13" captionText="FIGURE 7. B. nigerrimus (SAM B 3361). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176289/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
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a) ............. ....................................................................................................................................................
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<taxonomicName id="4C1C4D256D1DFFE0FAD5FBB96699FBC4" ID-CoL="LRV8" box="[1302,1436,1111,1137]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="nigerrimus">
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<emphasis id="B968EAB46D1DFFE0FAD5FBB96699FBC4" bold="true" box="[1302,1436,1111,1137]" italics="true" pageId="17" pageNumber="18">nigerrimus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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</document> |