182 lines
20 KiB
XML
182 lines
20 KiB
XML
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<mods:title id="1341EA5A9DD7C8B99E9ACE64CB292793">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:namePart id="37D9F0671B23E4D85C1ACFBCE6CF9099">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="CCE9F2A20877763524026B1CC3C6DAA6">Karapunar, Baran</mods:namePart>
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<mods:namePart id="E6A182E467C20C8066CBFAA2BAF0FDF6">Charoentitirat, Thasinee</mods:namePart>
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<mods:namePart id="AB55D34AC360E76648918F167B350FAC">Nützel, Al- Exander</mods:namePart>
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<mods:title id="975F75FE8684D7B808829ADE3E8BD313">Zootaxa</mods:title>
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<mods:date id="812B43122688934E22EB145615DB8BD6">2020</mods:date>
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<mods:number id="B5F26E0ED2A1598CAD62FDF04BFF13E1">2020-04-17</mods:number>
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<mods:number id="5313C6A5061E1CFDABADCC49A8FCE662">4766</mods:number>
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<treatment id="03B587AB4F12155BFF517A13FB44FA3B" ID-DOI="http://doi.org/10.5281/zenodo.3803809" ID-GBIF-Taxon="163737186" ID-Zenodo-Dep="3803809" LSID="urn:lsid:plazi:treatment:03B587AB4F12155BFF517A13FB44FA3B" httpUri="http://treatment.plazi.org/id/03B587AB4F12155BFF517A13FB44FA3B" lastPageNumber="38" pageId="37" pageNumber="38">
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<subSubSection id="C30665364F12155BFF517A13FDB0FD02" box="[151,635,621,648]" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph id="8BA336BD4F12155BFF517A13FDB0FD02" blockId="37.[151,635,621,683]" box="[151,635,621,648]" pageId="37" pageNumber="38">
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<heading id="D0EB81D14F12155BFF517A13FDB0FD02" bold="true" box="[151,635,621,648]" fontSize="10" level="2" pageId="37" pageNumber="38" reason="2">
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<taxonomicName id="4C1C4D3E4F12155BFF517A13FDB0FD02" authority="(Longstaff, 1933)" baseAuthorityName="Longstaff" baseAuthorityYear="1933" box="[151,635,621,648]" class="Gastropoda" family="Palaeostylidae" genus="Knightella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="irregularis">
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<emphasis id="B968EAAF4F12155BFF517A13FDB0FD02" bold="true" box="[151,635,621,648]" pageId="37" pageNumber="38">
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<emphasis id="B968EAAF4F12155BFF517A13FE54FD0D" bold="true" box="[151,415,621,648]" italics="true" pageId="37" pageNumber="38">Knightella irregularis</emphasis>
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(
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<bibRefCitation id="EF8D4B4C4F12155BFE6B7A13FDB8FD02" author="Longstaff" box="[429,627,621,647]" firstAuthor="Longstaff" pageId="37" pageNumber="38" pagination="87 - 124" refId="ref25915" refString="Longstaff, J. D. (1933) A revision of the British Carboniferous Loxonematidae, with descriptions of new forms. Quarterly Journal of the Geological Society of London, 89 (2), 87 - 124. https: // doi. org / 10.1144 / GSL. JGS. 1933.089.01 - 04.05" type="journal article" year="1933">Longstaff, 1933</bibRefCitation>
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)
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30665364F12155BFF517AEFFEE0FD2E" box="[151,299,656,683]" pageId="37" pageNumber="38" type="description">
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<paragraph id="8BA336BD4F12155BFF517AEFFEE0FD2E" blockId="37.[151,635,621,683]" box="[151,299,656,683]" pageId="37" pageNumber="38">
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(
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<figureCitation id="13272A384F12155BFF667AEFFF24FD2E" box="[160,239,656,683]" captionStart="FIGURE 25" captionStartId="38.[151,250,1830,1855]" captionTargetBox="[197,1390,181,1805]" captionTargetId="figure@38.[197,1390,181,1805]" captionTargetPageId="38" captionText="FIGURE 25. A, Donaldospira sp., ESKU-19-LP 85. B–E, Knightella irregularis (Longstaff, 1933), B–C, ESKU-19-LP 41. D–E, ESKU-19-LP 84. F–H, Protostylus sp., F, ESKU-19-LP 39. G–H, ESKU-19-LP 64. I–N, Trypanocochlea lopburiensis sp. nov. I, M, ESKU-19-LP 210, holotype; oblique view of early whorls to show protoconch. J, ESKU-19-LP 201, paratype. K, ESKU-19-LP 203, paratype. L, ESKU-19-LP 206, paratype; lateral view of early whorls to show protoconch. N, ESKU-19-LP 221, paratype. O–R, Streptacis? khaokhadensis sp. nov., O–P, ESKU-19-LP 214, holotype. Q–R, ESKU-19-LP 213, paratype. Scale bars represent: 2 mm (A–H); 200 μm (I–R)." figureDoi="http://doi.org/10.5281/zenodo.3763957" httpUri="https://zenodo.org/record/3763957/files/figure.png" pageId="37" pageNumber="38">Fig. 25</figureCitation>
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B–E)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F12155BFF517AABFB96FD6B" box="[151,1117,725,750]" pageId="37" pageNumber="38" type="reference_group">
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<paragraph id="8BA336BD4F12155BFF517AABFB96FD6B" blockId="37.[151,1117,725,750]" box="[151,1117,725,750]" pageId="37" pageNumber="38">
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<treatmentCitationGroup id="AB0C11934F12155BFF517AABFB96FD6B" box="[151,1117,725,750]" pageId="37" pageNumber="38">
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<treatmentCitation id="0ABD10AC4F12155BFF517AABFDF4FD68" author="Longstaff, J. D." box="[151,575,725,750]" page="118" pageId="37" pageNumber="38" year="1933">
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<taxonomicName id="4C1C4D3E4F12155BFF517AABFDF4FD68" authority="Longstaff, 1933: 118" authorityName="Longstaff" authorityPageNumber="118" authorityYear="1933" box="[151,575,725,750]" class="Magnoliopsida" family="Proteaceae" genus="Knightia" kingdom="Plantae" order="Proteales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="irregularis">
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<emphasis id="B968EAAF4F12155BFF517AABFEA8FD68" box="[151,355,725,749]" italics="true" pageId="37" pageNumber="38">Knightia irregularis</emphasis>
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<bibRefCitation id="EF8D4B4C4F12155BFEAF7AABFDF4FD68" author="Longstaff" box="[361,575,725,750]" firstAuthor="Longstaff" pageId="37" pageNumber="38" pagination="87 - 124" refId="ref25915" refString="Longstaff, J. D. (1933) A revision of the British Carboniferous Loxonematidae, with descriptions of new forms. Quarterly Journal of the Geological Society of London, 89 (2), 87 - 124. https: // doi. org / 10.1144 / GSL. JGS. 1933.089.01 - 04.05" type="journal article" year="1933">Longstaff, 1933: 118</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, pl. 12, figs 4–5;
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<treatmentCitation id="0ABD10AC4F12155BFD367AABFC60FD6B" author="Knight, J. B." box="[752,939,725,750]" page="164" pageId="37" pageNumber="38" year="1941">
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<bibRefCitation id="EF8D4B4C4F12155BFD367AABFC60FD6B" author="Knight" box="[752,939,725,750]" firstAuthor="Knight" pageId="37" pageNumber="38" pagination="1 - 510" refId="ref25231" refString="Knight, J. B. (1941) Paleozoic gastropod genotypes. Geological Society of America Special Paper, 32, 1 - 510. https: // doi. org / 10.1130 / SPE 32 - p 1" type="journal article" year="1941">Knight, 1941: 164</bibRefCitation>
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</treatmentCitation>
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, pl. 49, fig. 6a–b.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30665364F12155BFF517B65FEAFFCFB" pageId="37" pageNumber="38" type="materials_examined">
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<paragraph id="8BA336BD4F12155BFF517B65FD7EFCB3" blockId="37.[151,1437,795,1470]" box="[151,693,795,822]" pageId="37" pageNumber="38">
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<emphasis id="B968EAAF4F12155BFF517B65FECDFCB3" bold="true" box="[151,262,795,822]" pageId="37" pageNumber="38">Material.</emphasis>
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Two specimens: ESKU-19-
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<collectionCode id="ED0DAE784F12155BFDF97B62FDAAFCB3" box="[575,609,796,822]" country="Russia" name="Laboratory of Palaeontology" pageId="37" pageNumber="38">LP</collectionCode>
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41, 84.
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</paragraph>
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<paragraph id="8BA336BD4F12155BFF017B3EFEAFFCFB" blockId="37.[151,1437,795,1470]" pageId="37" pageNumber="38">
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Dimensions (mm): ESKU-19-
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<collectionCode id="ED0DAE784F12155BFDD07B3EFDF3FCDF" box="[534,568,832,858]" country="Russia" name="Laboratory of Palaeontology" pageId="37" pageNumber="38">LP</collectionCode>
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41: height =
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<emphasis id="B968EAAF4F12155BFD0B7B3EFD12FCDC" box="[717,729,832,857]" italics="true" pageId="37" pageNumber="38">c</emphasis>
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. 16.6; width = 6.1. ESKU-19-
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<collectionCode id="ED0DAE784F12155BFBED7B3EFB86FCDF" box="[1067,1101,832,858]" country="Russia" name="Laboratory of Palaeontology" pageId="37" pageNumber="38">LP</collectionCode>
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84: height = 8.9; width = 4.0; apical angle = 42
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<superScript id="7C699BF54F12155BFE917B1CFE97FCF5" attach="none" box="[343,348,866,880]" fontSize="6" pageId="37" pageNumber="38">º</superScript>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F12155BFF017BF9FD1CFB8B" pageId="37" pageNumber="38" type="description">
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<paragraph id="8BA336BD4F12155BFF017BF9FD1CFB8B" blockId="37.[151,1437,795,1470]" pageId="37" pageNumber="38">
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<emphasis id="B968EAAF4F12155BFF017BF9FE92FC27" bold="true" box="[199,345,903,930]" pageId="37" pageNumber="38">Description.</emphasis>
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Shell high-spired, slender; largest specimen consisting of 10 whorls; protoconch smooth, seemingly about two orthostrophic whorls; whorls low, gently increasing; whorls smooth; whorl face evenly convex or somewhat pendent with periphery at mid-whorl or somewhat below; sutures moderately deep, distinct; base evenly convex, rounded; anomphalous; aperture unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F12155BFF017C66FB44FA3B" pageId="37" pageNumber="38" type="discussion">
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<paragraph id="8BA336BD4F12155BFF017C66FB44FA3B" blockId="37.[151,1437,795,1470]" pageId="37" pageNumber="38">
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<emphasis id="B968EAAF4F12155BFF017C66FEF0FBB7" bold="true" box="[199,315,1048,1074]" pageId="37" pageNumber="38">Remarks.</emphasis>
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The present specimens closely resemble the
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<typeStatus id="54A7881F4F12155BFCF87C66FCA5FBB7" box="[830,878,1048,1074]" pageId="37" pageNumber="38">type</typeStatus>
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species,
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<taxonomicName id="4C1C4D3E4F12155BFC107C66FA57FBB7" authority="(Longstaff, 1933)" baseAuthorityName="Longstaff" baseAuthorityYear="1933" box="[982,1436,1048,1074]" class="Gastropoda" family="Palaeostylidae" genus="Knightella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="irregularis">
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<emphasis id="B968EAAF4F12155BFC107C66FB06FBB7" box="[982,1229,1048,1074]" italics="true" pageId="37" pageNumber="38">Knightella irregularis</emphasis>
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||
(
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<bibRefCitation id="EF8D4B4C4F12155BFB1B7C66FA5FFBB7" author="Longstaff" box="[1245,1428,1048,1074]" firstAuthor="Longstaff" pageId="37" pageNumber="38" pagination="87 - 124" refId="ref25915" refString="Longstaff, J. D. (1933) A revision of the British Carboniferous Loxonematidae, with descriptions of new forms. Quarterly Journal of the Geological Society of London, 89 (2), 87 - 124. https: // doi. org / 10.1144 / GSL. JGS. 1933.089.01 - 04.05" type="journal article" year="1933">Longstaff, 1933</bibRefCitation>
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)
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</taxonomicName>
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from the Carboniferous,
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<collectingCountry id="F30B762D4F12155BFE6F7C42FDC0FBD3" box="[425,523,1084,1110]" name="United Kingdom" pageId="37" pageNumber="38">Scotland</collectingCountry>
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as re-described and illustrated by
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<bibRefCitation id="EF8D4B4C4F12155BFC407C42FBD5FBD3" author="Knight" box="[902,1054,1084,1110]" firstAuthor="Knight" pageId="37" pageNumber="38" pagination="1 - 510" refId="ref25231" refString="Knight, J. B. (1941) Paleozoic gastropod genotypes. Geological Society of America Special Paper, 32, 1 - 510. https: // doi. org / 10.1130 / SPE 32 - p 1" type="journal article" year="1941">Knight (1941</bibRefCitation>
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, p.165, pl. 49, fig. 6a–b), although the present specimens are much larger. The general features of this species (i.e., high spire, sharply rounded and somewhat pendent whorl profile, a body whorl which is two-fifths of the total height, rounded base, anomphalous shell and slightly protruding protoconch) are obviously identical with the present specimens. Other similar species are
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<taxonomicName id="4C1C4D3E4F12155BFF797CB2FDDAFB63" authority="Nutzel, 2012" authorityName="Nutzel" authorityYear="2012" box="[191,529,1228,1254]" class="Gastropoda" family="Palaeostylidae" genus="Knightella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="hydrobiformis">
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||
<emphasis id="B968EAAF4F12155BFF797CB2FF19FB60" box="[191,210,1228,1253]" italics="true" pageId="37" pageNumber="38">K</emphasis>
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.
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<emphasis id="B968EAAF4F12155BFF197CB2FEB6FB63" box="[223,381,1228,1254]" italics="true" pageId="37" pageNumber="38">hydrobiformis</emphasis>
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Nützel, 2012
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</taxonomicName>
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and
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<taxonomicName id="4C1C4D3E4F12155BFD837CB2FD70FB63" authorityName="Longstaff" authorityYear="1933" box="[581,699,1228,1254]" class="Gastropoda" family="Palaeostylidae" genus="Knightella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F12155BFD837CB2FD70FB63" box="[581,699,1228,1254]" italics="true" pageId="37" pageNumber="38">Knightella</emphasis>
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</taxonomicName>
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sp. from the Middle Permian Akasaka Limestone of
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<collectingCountry id="F30B762D4F12155BFAC57CB2FA88FB63" box="[1283,1347,1228,1254]" name="Japan" pageId="37" pageNumber="38">Japan</collectingCountry>
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(Nützel &
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<bibRefCitation id="EF8D4B4C4F12155BFF727C8EFEBBFA8F" author="Nakazawa" box="[180,368,1264,1290]" firstAuthor="Nakazawa" pageId="37" pageNumber="38" pagination="103 - 169" refId="ref27024" refString="Nu ¨ tzel, A. & Nakazawa, K. (2012) Permian (Capitanian) gastropods from the Akasaka Limestone (Gifu Prefecture, Japan). Journal of Systematic Palaeontology, 10, 103 - 169. https: // doi. org / 10.1080 / 14772019.2010.549659" type="journal article" year="2012">Nakazawa, 2012</bibRefCitation>
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) and also several
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<taxonomicName id="4C1C4D3E4F12155BFDF07C8EFD67FA8F" authorityName="Longstaff" authorityYear="1933" box="[566,684,1264,1290]" class="Gastropoda" family="Palaeostylidae" genus="Knightella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F12155BFDF07C8EFD67FA8F" box="[566,684,1264,1290]" italics="true" pageId="37" pageNumber="38">Knightella</emphasis>
|
||
</taxonomicName>
|
||
species which have been reported by Nützel (1998) from the Pennsylvanian of the
|
||
<collectingCountry id="F30B762D4F12155BFE927D6AFE5BFAAB" box="[340,400,1300,1326]" name="United States of America" pageId="37" pageNumber="38">USA</collectingCountry>
|
||
. However, these species are smaller and have less convex whorls.
|
||
<taxonomicName id="4C1C4D3E4F12155BFB427D6AFEF4FAD7" authority="Licharev, 1975" authorityName="Licharev" authorityYear="1975" class="Gastropoda" family="Murchisoniidae" genus="Loxonema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="karabolkensis">
|
||
<emphasis id="B968EAAF4F12155BFB427D6AFA56FAAB" box="[1156,1437,1300,1326]" italics="true" pageId="37" pageNumber="38">Loxonema karabolkensis</emphasis>
|
||
<bibRefCitation id="EF8D4B4C4F12155BFF517D46FEF4FAD7" author="Licharev" box="[151,319,1335,1362]" firstAuthor="Licharev" pageId="37" pageNumber="38" pagination="5 - 183" refId="ref25884" refString="Licharev, B. K. (1975) Carboniferous gastropods from the region of the River Karabolka. Trudy Vsesoyuzniei Navcho-Issledovatelbskii Geologicheskii Institut, 206, 5 - 183." type="journal article" year="1975">Licharev, 1975</bibRefCitation>
|
||
</taxonomicName>
|
||
(p. 78, pl. 13, figs 5–8) from the Pennsylvanian of
|
||
<collectingCountry id="F30B762D4F12155BFC427D46FC04FAD7" box="[900,975,1336,1362]" name="Russia" pageId="37" pageNumber="38">Russia</collectingCountry>
|
||
is similar but it differs in having a dense spiral ornament on the whorls. The present specimens are also similar to
|
||
<taxonomicName id="4C1C4D3E4F12155BFC797D22FBF5FAF3" authorityName="Mansuy" authorityYear="1914" box="[959,1086,1372,1398]" class="Gastropoda" family="Orthonematidae" genus="Protostylus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F12155BFC797D22FBF5FAF3" box="[959,1086,1372,1398]" italics="true" pageId="37" pageNumber="38">Protostylus</emphasis>
|
||
</taxonomicName>
|
||
; however, this genus has flatter whorls. This is the first known member of the genus
|
||
<taxonomicName id="4C1C4D3E4F12155BFD237DFEFC90FA1F" authorityName="Longstaff" authorityYear="1933" box="[741,859,1408,1434]" class="Gastropoda" family="Palaeostylidae" genus="Knightella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F12155BFD237DFEFC90FA1F" box="[741,859,1408,1434]" italics="true" pageId="37" pageNumber="38">Knightella</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F30B762D4F12155BFCB87DFEFC29FA1F" box="[894,994,1408,1434]" name="Thailand" pageId="37" pageNumber="38">Thailand</collectingCountry>
|
||
which extends the range of this species and genus to the Middle Permian and expands its distribution to the eastern Palaeo-Tethys.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |