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<document id="501F4593AA08ECEBC00D95DB8DFD102C" ID-CLB-Dataset="28081" ID-DOI="10.11646/zootaxa.4504.2.8" ID-GBIF-Dataset="a435adb8-3cba-4dde-a36c-deba6ea177c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606285" ID-ZooBank="9AC935DB-B027-421A-AB13-5E275F976BAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553584541085" checkinUser="plazi" docAuthor="Huo, Qing-Bo &amp; Du, Yu-Zhou" docDate="2018" docId="03B57A17851AA447A1EC372E6606FF78" docLanguage="en" docName="zootaxa.4504.2.8.pdf" docOrigin="Zootaxa 4504 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Isoperla oncocauda Huo &amp; Du 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="282" masterDocId="FF8C026F851BA441A17B36436420FFAD" masterDocTitle="A new species of the genus Isoperla (Plecoptera: Perlodidae) from TIANMU MOUNTAIN NATURE RESERVE, China" masterLastPageNumber="284" masterPageNumber="276" pageNumber="277" updateTime="1698712743732" updateUser="ExternalLinkService">
<mods:mods id="DC9FE78A99885EDC0ABF34D6F09FDF2A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1FF22C271AD78DC559A7CDE6D7633354">
<mods:title id="11E4B611BFCC3776397B6B0FC79B47A4">A new species of the genus Isoperla (Plecoptera: Perlodidae) from TIANMU MOUNTAIN NATURE RESERVE, China</mods:title>
</mods:titleInfo>
<mods:name id="6FB5CD2D771246A6FE3781916D0C560F" type="personal">
<mods:role id="13D9B91EAD0E6A590034BF051A07DF3B">
<mods:roleTerm id="88BA28A764966AA01003C75B6D2FEE24">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09C98EF103D1CD948EF0609D2398FF68">Huo, Qing-Bo</mods:namePart>
</mods:name>
<mods:name id="21A4C0D92B8F9B6B9AD2283DB6A078D2" type="personal">
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<mods:namePart id="FC89F7283956A1D179A1EE8775F7AD40">Du, Yu-Zhou</mods:namePart>
</mods:name>
<mods:typeOfResource id="78C208559C587A50E44F63292A38B7CF">text</mods:typeOfResource>
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<mods:titleInfo id="FB799AA1B9B6DD6363F24B2E05021E5D">
<mods:title id="F98D3036220FE21A1D21723B0CFD9204">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="261D3A524D34C72A4F503C027EA24E78">
<mods:date id="61BAA312FF7237305EB06CF598F3E96E">2018</mods:date>
<mods:detail id="AB4D9B734B3D7D980DA77099EB9A9292" type="pubDate">
<mods:number id="4A6E943581863415090A9AF72C19B19E">2018-10-24</mods:number>
</mods:detail>
<mods:detail id="DD93348D82FB1504D3AC713D62F17A4A" type="volume">
<mods:number id="B188B36B1085ABB65079B5479E90F75A">4504</mods:number>
</mods:detail>
<mods:detail id="29582AF131CA0067BBA73773577BB085" type="issue">
<mods:number id="D40F2A589BD488D8C630E8A2CCA874F4">2</mods:number>
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<mods:start id="86E0C185305CB83CAC647A741D2C17C5">276</mods:start>
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<mods:classification id="063B89A653C6216DB4532C371D9B32D2">journal article</mods:classification>
<mods:identifier id="E0D1771CD9F2F466E1A1ACCB8F343E5F" type="CLB-Dataset">28081</mods:identifier>
<mods:identifier id="901F5D2DB64F9621C50BB65DE9198B07" type="DOI">10.11646/zootaxa.4504.2.8</mods:identifier>
<mods:identifier id="7E7110C08A7A4F466747D176B2DB603D" type="GBIF-Dataset">a435adb8-3cba-4dde-a36c-deba6ea177c1</mods:identifier>
<mods:identifier id="356819D651D215B0EC0BDF4210914006" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EBA1D7C961B5A9BD3073A57B07B60E3C" type="Zenodo-Dep">2606285</mods:identifier>
<mods:identifier id="A1D058215D0357A1C19ACB46DF176A32" type="ZooBank">9AC935DB-B027-421A-AB13-5E275F976BAD</mods:identifier>
</mods:mods>
<treatment id="03B57A17851AA447A1EC372E6606FF78" ID-DOI="http://doi.org/10.5281/zenodo.5952446" ID-GBIF-Taxon="154495043" ID-Zenodo-Dep="5952446" LSID="urn:lsid:plazi:treatment:03B57A17851AA447A1EC372E6606FF78" httpUri="http://treatment.plazi.org/id/03B57A17851AA447A1EC372E6606FF78" lastPageId="6" lastPageNumber="282" pageId="1" pageNumber="277">
<subSubSection id="C306988A851AA440A1EC372E665BFE2A" box="[151,635,365,391]" pageId="1" pageNumber="277" type="nomenclature">
<paragraph id="8BA3CB01851AA440A1EC372E665BFE2A" blockId="1.[151,635,365,391]" box="[151,635,365,391]" pageId="1" pageNumber="277">
<heading id="D0EB7C6D851AA440A1EC372E665BFE2A" bold="true" box="[151,635,365,391]" fontSize="11" level="1" pageId="1" pageNumber="277" reason="1">
<emphasis id="B9681713851AA440A1EC372E665BFE2A" bold="true" box="[151,635,365,391]" pageId="1" pageNumber="277">
<taxonomicName id="4C1CB082851AA440A1EC372E6630FE2A" ID-CoL="8NH9B" authority="Huo &amp; Du" authorityName="Huo &amp; Du" authorityYear="2018" box="[151,528,365,391]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="oncocauda" status="sp. nov.">
<emphasis id="B9681713851AA440A1EC372E65A5FE2A" bold="true" box="[151,389,365,391]" italics="true" pageId="1" pageNumber="277">Isoperla oncocauda</emphasis>
Huo &amp; Du
</taxonomicName>
,
<taxonomicNameLabel id="A25BAA68851AA440A366372E665BFE2A" box="[541,635,365,391]" pageId="1" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C306988A851AA442A1EC37F16586F84B" lastPageId="3" lastPageNumber="279" pageId="1" pageNumber="277" type="description">
<paragraph id="8BA3CB01851AA440A1EC37F165C0FCD6" blockId="1.[151,1436,434,891]" pageId="1" pageNumber="277">
<emphasis id="B9681713851AA440A1EC37F164FDFE66" bold="true" box="[151,221,434,459]" pageId="1" pageNumber="277">Male:</emphasis>
Forewing length ca. 11.0 mm, hindwing length 10.5 mm, body length 10.0 mm. Head mostly dark brown, anterior ocellus slightly smaller, posterior ocelli closer to compound eyes than to each other; area between posterior ocelli to compound eyes yellow brown (
<figureCitation id="1327D784851AA440A31737B966FBFDBE" box="[620,731,506,531]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1950,1973]" captionTargetBox="[219,1436,946,1927]" captionTargetId="figure@1.[150,1436,946,1927]" captionTargetPageId="1" captionText="FIGURES 1–2. Isoperla oncocauda, sp. nov. 1: Male habitus; 2: Male head and pronotum." httpUri="https://zenodo.org/record/2606287/files/figure.png" pageId="1" pageNumber="277">Figs. 1–2</figureCitation>
). Antennae and palpi dark brown. Pronotum disc wider than long, black, lateral margins yellowbrown, rugosities present medially (
<figureCitation id="1327D784851AA440A2A3345D6069FD9A" box="[984,1097,542,567]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1950,1973]" captionTargetBox="[219,1436,946,1927]" captionTargetId="figure@1.[150,1436,946,1927]" captionTargetPageId="1" captionText="FIGURES 1–2. Isoperla oncocauda, sp. nov. 1: Male habitus; 2: Male head and pronotum." httpUri="https://zenodo.org/record/2606287/files/figure.png" pageId="1" pageNumber="277">Figs. 1–2</figureCitation>
). Legs yellow brown; wings hyaline, veins dark brown (
<figureCitation id="1327D784851AA440A0B53401661DFDF6" box="[462,573,578,603]" captionStart="FIGURES 3–4" captionStartId="2.[151,264,1065,1088]" captionTargetBox="[325,1258,222,1029]" captionTargetId="figure@2.[224,1366,193,1049]" captionTargetPageId="2" captionText="FIGURES 3–4. Isoperla oncocauda, sp. nov., veins of male. 3: Forewing; 4: Hindwing." httpUri="https://zenodo.org/record/2606291/files/figure.png" pageId="1" pageNumber="277">Figs. 3–4</figureCitation>
). Abdomen dark brown and slightly sclerotized. Posterior margin of tergum 10 darkly sclerotized, recessed medially and rearward, extending as a process, with wide lateral grooves on surface, and perpendicular to tergum 10 posteriorly. Paraprocts sclerotized, blunt apically (
<figureCitation id="1327D784851AA440A51634C960E3FD0E" box="[1133,1219,650,675]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1996,2019]" captionTargetBox="[151,1435,1130,1986]" captionTargetId="figure@2.[151,1435,1130,1986]" captionTargetPageId="2" captionText="FIGURES 5–6. Isoperla oncocauda, sp. nov., terminalia of male. 5: Dorsal view; 6: Ventral view." httpUri="https://zenodo.org/record/2606293/files/figure.png" pageId="1" pageNumber="277">Figs. 5</figureCitation>
). Vesicle reduced, slightly projecting beyond anterior margin of sternum 9 (
<figureCitation id="1327D784851AA440A24D34ED675FFD6A" box="[822,895,686,711]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1996,2019]" captionTargetBox="[151,1435,1130,1986]" captionTargetId="figure@2.[151,1435,1130,1986]" captionTargetPageId="2" captionText="FIGURES 5–6. Isoperla oncocauda, sp. nov., terminalia of male. 5: Dorsal view; 6: Ventral view." httpUri="https://zenodo.org/record/2606293/files/figure.png" pageId="1" pageNumber="277">Fig. 6</figureCitation>
). Anteromedially, sternum 9 yellowish-brown, membranous; posterior margin truncate (
<figureCitation id="1327D784851AA440A31D349166C2FD46" box="[614,738,722,747]" captionStart-0="FIGURES 5–6" captionStart-1="FIGURES 7–8" captionStart-2="FIGURES 9–10" captionStartId-0="2.[151,264,1996,2019]" captionStartId-1="3.[151,264,1750,1773]" captionStartId-2="4.[151,264,751,774]" captionTargetBox-0="[151,1435,1130,1986]" captionTargetBox-1="[281,1305,193,1729]" captionTargetBox-2="[355,1263,183,704]" captionTargetId-0="figure@2.[151,1435,1130,1986]" captionTargetId-1="figure@3.[281,1305,193,1729]" captionTargetId-2="figure@4.[267,1327,143,739]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 5–6. Isoperla oncocauda, sp. nov., terminalia of male. 5: Dorsal view; 6: Ventral view." captionText-1="FIGURES 7–8. Isoperla oncocauda, sp. nov., process on T10 of male. 7: Rear view; 8: Lateral view." captionText-2="FIGURES 9–10. Isoperla oncocauda, sp. nov. 9: Terminalia of male, lateral view; 10: Paraproct of male, lateral view." httpUri-0="https://zenodo.org/record/2606293/files/figure.png" httpUri-1="https://zenodo.org/record/2606295/files/figure.png" httpUri-2="https://zenodo.org/record/2606297/files/figure.png" pageId="1" pageNumber="277">Figs. 6–10</figureCitation>
). Aedeagus membranous without conspicuous sclerites (
<figureCitation id="1327D784851AA440A41F349164FDFCA2" captionStart="FIGURES 11–12" captionStartId="4.[151,264,1550,1573]" captionTargetBox="[210,1375,855,1515]" captionTargetId="figure@4.[159,1429,824,1538]" captionTargetPageId="4" captionText="FIGURES 11–12. Isoperla oncocauda, sp. nov. aedeagus. 11: Lateral view; 12: Ventral view." httpUri="https://zenodo.org/record/2606299/files/figure.png" pageId="1" pageNumber="277">Figs. 11–12</figureCitation>
) [partially everted]. In lateral view with one large dorsal lobe bearing few rows of spinules, one anterolateral lobe and three posteroapical lobes. Ventral surface covered with dense scale-like spines increasing in size toward base. A triangular membranous notch near the middle of anterior margin; nail-shaped spines before the notch increasing in size posteriorly.
</paragraph>
<caption id="DF639B89851AA440A1EC31DD6018F818" box="[151,1080,1950,1973]" httpUri="https://zenodo.org/record/2606287/files/figure.png" pageId="1" pageNumber="277" startId="1.[151,264,1950,1973]" targetBox="[219,1436,946,1927]" targetPageId="1">
<paragraph id="8BA3CB01851AA440A1EC31DD6018F818" blockId="1.[151,1080,1950,1973]" box="[151,1080,1950,1973]" pageId="1" pageNumber="277">
<emphasis id="B9681713851AA440A1EC31DD651AF819" bold="true" box="[151,314,1950,1973]" pageId="1" pageNumber="277">FIGURES 1–2.</emphasis>
<taxonomicName id="4C1CB082851AA440A03B31DC662AF818" authority="Huo &amp; Du, 2018" authorityName="Huo &amp; Du" authorityYear="2018" box="[320,522,1951,1973]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="oncocauda" status="sp. nov.">
<emphasis id="B9681713851AA440A03B31DC662AF818" box="[320,522,1951,1973]" italics="true" pageId="1" pageNumber="277">Isoperla oncocauda</emphasis>
</taxonomicName>
,
<emphasis id="B9681713851AA440A36C31DC6647F819" bold="true" box="[535,615,1951,1972]" pageId="1" pageNumber="277">
<taxonomicNameLabel id="A25BAA68851AA440A36C31DC6647F819" box="[535,615,1951,1972]" pageId="1" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1: Male habitus; 2: Male head and pronotum.
</paragraph>
</caption>
<caption id="DF639B898519A443A1EC326A6038FBED" box="[151,1048,1065,1088]" httpUri="https://zenodo.org/record/2606291/files/figure.png" pageId="2" pageNumber="278" startId="2.[151,264,1065,1088]" targetBox="[325,1258,222,1029]" targetPageId="2">
<paragraph id="8BA3CB018519A443A1EC326A6038FBED" blockId="2.[151,1048,1065,1088]" box="[151,1048,1065,1088]" pageId="2" pageNumber="278">
<emphasis id="B96817138519A443A1EC326A651AFB92" bold="true" box="[151,314,1065,1088]" pageId="2" pageNumber="278">FIGURES 3–4.</emphasis>
<taxonomicName id="4C1CB0828519A443A03B3269662AFBED" authority="Huo &amp; Du, 2018" authorityName="Huo &amp; Du" authorityYear="2018" box="[320,522,1066,1088]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="oncocauda" status="sp. nov.">
<emphasis id="B96817138519A443A03B3269662AFBED" box="[320,522,1066,1088]" italics="true" pageId="2" pageNumber="278">Isoperla oncocauda</emphasis>
</taxonomicName>
,
<emphasis id="B96817138519A443A36C32696647FB92" bold="true" box="[535,615,1066,1087]" pageId="2" pageNumber="278">
<taxonomicNameLabel id="A25BAA688519A443A36C32696647FB92" box="[535,615,1066,1087]" pageId="2" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, veins of male. 3: Forewing; 4: Hindwing.
</paragraph>
</caption>
<caption id="DF639B898519A443A1EC318F605DF84E" box="[151,1149,1996,2019]" httpUri="https://zenodo.org/record/2606293/files/figure.png" pageId="2" pageNumber="278" startId="2.[151,264,1996,2019]" targetBox="[151,1435,1130,1986]" targetPageId="2">
<paragraph id="8BA3CB018519A443A1EC318F605DF84E" blockId="2.[151,1149,1996,2019]" box="[151,1149,1996,2019]" pageId="2" pageNumber="278">
<emphasis id="B96817138519A443A1EC318F651AF84F" bold="true" box="[151,314,1996,2019]" pageId="2" pageNumber="278">FIGURES 5–6.</emphasis>
<taxonomicName id="4C1CB0828519A443A03B318E662AF84E" authority="Huo &amp; Du, 2018" authorityName="Huo &amp; Du" authorityYear="2018" box="[320,522,1997,2019]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="oncocauda" status="sp. nov.">
<emphasis id="B96817138519A443A03B318E662AF84E" box="[320,522,1997,2019]" italics="true" pageId="2" pageNumber="278">Isoperla oncocauda</emphasis>
</taxonomicName>
,
<emphasis id="B96817138519A443A36C318D6647F84E" bold="true" box="[535,615,1998,2019]" pageId="2" pageNumber="278">
<taxonomicNameLabel id="A25BAA688519A443A36C318D6647F84E" box="[535,615,1998,2019]" pageId="2" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, terminalia of male. 5: Dorsal view; 6: Ventral view.
</paragraph>
</caption>
<caption id="DF639B898518A442A1EC309560BCF941" box="[151,1180,1750,1773]" httpUri="https://zenodo.org/record/2606295/files/figure.png" pageId="3" pageNumber="279" startId="3.[151,264,1750,1773]" targetBox="[281,1305,193,1729]" targetPageId="3">
<paragraph id="8BA3CB018518A442A1EC309560BCF941" blockId="3.[151,1180,1750,1773]" box="[151,1180,1750,1773]" pageId="3" pageNumber="279">
<emphasis id="B96817138518A442A1EC3095651AF941" bold="true" box="[151,314,1750,1773]" pageId="3" pageNumber="279">FIGURES 7–8.</emphasis>
<taxonomicName id="4C1CB0828518A442A03B3095662AF941" authority="Huo &amp; Du, 2018" authorityName="Huo &amp; Du" authorityYear="2018" box="[320,522,1750,1772]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="279" phylum="Arthropoda" rank="species" species="oncocauda" status="sp. nov.">
<emphasis id="B96817138518A442A03B3095662AF941" box="[320,522,1750,1772]" italics="true" pageId="3" pageNumber="279">Isoperla oncocauda</emphasis>
</taxonomicName>
,
<emphasis id="B96817138518A442A36C30946647F941" bold="true" box="[535,615,1751,1772]" pageId="3" pageNumber="279">
<taxonomicNameLabel id="A25BAA688518A442A36C30946647F941" box="[535,615,1751,1772]" pageId="3" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, process on T10 of male. 7: Rear view; 8: Lateral view.
</paragraph>
</caption>
<paragraph id="8BA3CB018518A442A1BC315A607DF833" blockId="3.[151,1437,1817,2022]" pageId="3" pageNumber="279">
<emphasis id="B96817138518A442A1BC315A6507F89F" bold="true" box="[199,295,1817,1842]" pageId="3" pageNumber="279">Female:</emphasis>
Forewing length 13.0 mm, hindwing length 11.5mm, body length 10.5 mm. Body coloration similar to male (
<figureCitation id="1327D7848518A442A180317E65A5F8FB" box="[251,389,1853,1878]" captionStart="FIGURES 13–14" captionStartId="5.[151,264,1124,1147]" captionTargetBox="[257,1395,193,1108]" captionTargetId="figure@5.[194,1395,193,1108]" captionTargetPageId="5" captionText="FIGURES 13–14. Isoperla oncocauda, sp. nov. 13: Female habitus; 14: Female head and pronotum." httpUri="https://zenodo.org/record/2606301/files/figure.png" pageId="3" pageNumber="279">Figs. 13–14</figureCitation>
). Abdomen brown, weakly sclerotized. Posterior margin of tergum 10 darkly sclerotized and extending as a process similar to male. Subgenital plate with basal sclerites laterally, broadly triangular, covering almost all of sternum 9, truncate apically with an obscure median notch (
<figureCitation id="1327D7848518A442A2BC31C66071F833" box="[967,1105,1925,1950]" captionStart-0="FIGURES 15–16" captionStart-1="FIGURES 17–18" captionStartId-0="5.[151,264,2009,2032]" captionStartId-1="6.[151,264,1822,1845]" captionTargetBox-0="[188,1400,1187,1996]" captionTargetBox-1="[281,1305,265,1801]" captionTargetId-0="figure@5.[188,1400,1187,1996]" captionTargetId-1="figure@6.[281,1305,265,1801]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 15–16. Isoperla oncocauda, sp. nov., terminalia of female. 15: Dorsal view; 16: Ventral view." captionText-1="FIGURES 17–18. Isoperla oncocauda, sp. nov., process on T10 of female. 17: Rear view; 18: Lateral view." httpUri-0="https://zenodo.org/record/2606303/files/figure.png" httpUri-1="https://zenodo.org/record/2606305/files/figure.png" pageId="3" pageNumber="279">Figs. 15–18</figureCitation>
).
</paragraph>
<paragraph id="8BA3CB018518A442A1BC31EA655CF86F" blockId="3.[151,1437,1817,2022]" box="[199,380,1961,1986]" pageId="3" pageNumber="279">
<emphasis id="B96817138518A442A1BC31EA64DEF86F" bold="true" box="[199,254,1961,1986]" pageId="3" pageNumber="279">Egg:</emphasis>
Unknown.
</paragraph>
<paragraph id="8BA3CB018518A442A1BC318E6586F84B" blockId="3.[151,1437,1817,2022]" box="[199,422,1997,2022]" pageId="3" pageNumber="279">
<emphasis id="B96817138518A442A1BC318E6509F84B" bold="true" box="[199,297,1997,2022]" pageId="3" pageNumber="279">Nymph:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="DF639B89851FA445A1EC34AC6169FCAB" box="[151,1353,751,774]" httpUri="https://zenodo.org/record/2606297/files/figure.png" pageId="4" pageNumber="280" startId="4.[151,264,751,774]" targetBox="[355,1263,183,704]" targetPageId="4">
<paragraph id="8BA3CB01851FA445A1EC34AC6169FCAB" blockId="4.[151,1353,751,774]" box="[151,1353,751,774]" pageId="4" pageNumber="280">
<emphasis id="B9681713851FA445A1EC34AC6567FCA8" bold="true" box="[151,327,751,774]" pageId="4" pageNumber="280">FIGURES 9–10.</emphasis>
<taxonomicName id="4C1CB082851FA445A03634B36637FCAB" authority="Huo &amp; Du, 2018" authorityName="Huo &amp; Du" authorityYear="2018" box="[333,535,752,774]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="oncocauda" status="sp. nov.">
<emphasis id="B9681713851FA445A03634B36637FCAB" box="[333,535,752,774]" italics="true" pageId="4" pageNumber="280">Isoperla oncocauda</emphasis>
</taxonomicName>
,
<emphasis id="B9681713851FA445A35834B36654FCA8" bold="true" box="[547,628,752,773]" pageId="4" pageNumber="280">
<taxonomicNameLabel id="A25BAA68851FA445A35834B36654FCA8" box="[547,628,752,773]" pageId="4" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
9: Terminalia of male, lateral view; 10: Paraproct of male, lateral view.
</paragraph>
</caption>
<caption id="DF639B89851FA445A1EC304D606EF988" box="[151,1102,1550,1573]" httpUri="https://zenodo.org/record/2606299/files/figure.png" pageId="4" pageNumber="280" startId="4.[151,264,1550,1573]" targetBox="[210,1375,855,1515]" targetPageId="4">
<paragraph id="8BA3CB01851FA445A1EC304D606EF988" blockId="4.[151,1102,1550,1573]" box="[151,1102,1550,1573]" pageId="4" pageNumber="280">
<emphasis id="B9681713851FA445A1EC304D6572F989" bold="true" box="[151,338,1550,1573]" pageId="4" pageNumber="280">FIGURES 11–12.</emphasis>
<taxonomicName id="4C1CB082851FA445A023304C6602F988" authority="Huo &amp; Du, 2018" authorityName="Huo &amp; Du" authorityYear="2018" box="[344,546,1551,1573]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="oncocauda" status="sp. nov.">
<emphasis id="B9681713851FA445A023304C6602F988" box="[344,546,1551,1573]" italics="true" pageId="4" pageNumber="280">Isoperla oncocauda</emphasis>
</taxonomicName>
,
<emphasis id="B9681713851FA445A355304C665FF989" bold="true" box="[558,639,1551,1572]" pageId="4" pageNumber="280">
<taxonomicNameLabel id="A25BAA68851FA445A355304C665FF989" box="[558,639,1551,1572]" pageId="4" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
aedeagus. 11: Lateral view; 12: Ventral view.
</paragraph>
</caption>
<subSubSection id="C306988A851FA445A1BC300F67C9F900" pageId="4" pageNumber="280" type="materials_examined">
<paragraph id="8BA3CB01851FA445A1BC300F67C9F900" blockId="4.[151,1437,1612,2036]" pageId="4" pageNumber="280">
<emphasis id="B9681713851FA445A1BC300F655CF9C8" bold="true" box="[199,380,1612,1637]" pageId="4" pageNumber="280">Type material:</emphasis>
<materialsCitation id="3B74C15C851FA445A0F2300F60B9F924" ID-GBIF-Occurrence="2028137901" collectingDate="2018-04-22" collectorName="Huo &amp; Gao Peng" country="China" county="Lin'an City" elevation="861" latitude="30.337133" location="Wuliting" longLatPrecision="1" longitude="119.4356" municipality="Tianmu Mountain Nature Reserve" pageId="4" pageNumber="280" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang" typeStatus="holotype">
<typeStatus id="54A775A3851FA445A0F2300F65D6F9C8" box="[393,502,1612,1637]" pageId="4" pageNumber="280" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D1A0088851FA445A37D300E663DF9C9" box="[518,541,1613,1636]" pageId="4" pageNumber="280" type="male">♂</specimenCount>
,
<collectingCountry id="F30B8B91851FA445A355300F6654F9C8" box="[558,628,1612,1637]" name="China" pageId="4" pageNumber="280">China</collectingCountry>
,
<collectingRegion id="49D805E3851FA445A3FE300F677CF9C8" box="[645,860,1612,1637]" country="China" name="Zhejiang" pageId="4" pageNumber="280">Zhejiang Province</collectingRegion>
,
<collectingCounty id="62C2B38D851FA445A217300F67D5F9C8" box="[876,1013,1612,1637]" pageId="4" pageNumber="280">Lin’an City</collectingCounty>
,
<collectingMunicipality id="6BC7517B851FA445A57D300F61B8F9C9" box="[1030,1432,1612,1637]" pageId="4" pageNumber="280">Tianmu Mountain Nature Reserve</collectingMunicipality>
,
<location id="8EC39DDA851FA445A1EC303364DDF924" LSID="urn:lsid:plazi:treatment:03B57A17851AA447A1EC372E6606FF78:8EC39DDA851FA445A1EC303364DDF924" box="[151,253,1648,1673]" country="China" county="Lin'an City" latitude="30.337133" longLatPrecision="1" longitude="119.4356" municipality="Tianmu Mountain Nature Reserve" name="Wuliting" pageId="4" pageNumber="280" stateProvince="Zhejiang">Wuliting</location>
,
<elevation id="00312C32851FA445A0773033656EF924" box="[268,334,1648,1673]" metricMagnitude="2" metricUnit="m" metricValue="8.61" pageId="4" pageNumber="280" unit="m" value="861.0">
<quantity id="4CE466E4851FA445A0773033656EF924" box="[268,334,1648,1673]" metricMagnitude="2" metricUnit="m" metricValue="8.61" pageId="4" pageNumber="280" unit="m" value="861.0">861m</quantity>
</elevation>
,
<geoCoordinate id="EE28ADC6851FA445A025303365CFF924" box="[350,495,1648,1673]" direction="north" orientation="latitude" pageId="4" pageNumber="280" precision="1" value="30.337133">30°20.228′N</geoCoordinate>
,
<geoCoordinate id="EE28ADC6851FA445A084303366BBF924" box="[511,667,1648,1673]" direction="east" orientation="longitude" pageId="4" pageNumber="280" precision="1" value="119.4356">119°26.136′E</geoCoordinate>
,
<date id="FFA2EDC1851FA445A3D13032670EF924" box="[682,814,1649,1673]" pageId="4" pageNumber="280" value="2018-04-22">
<collectingDate id="EFE61429851FA445A3D13032670EF924" box="[682,814,1649,1673]" pageId="4" pageNumber="280" value="2018-04-22">2018-IV-22</collectingDate>
</date>
, leg.
<collectorName id="26E9AED7851FA445A20830326783F925" box="[883,931,1649,1672]" pageId="4" pageNumber="280">Huo</collectorName>
Qing-Bo,
<collectorName id="26E9AED7851FA445A558303260B9F924" box="[1059,1177,1649,1673]" pageId="4" pageNumber="280">Gao Peng</collectorName>
</materialsCitation>
.
<materialsCitation id="3B74C15C851FA445A5D3303267C5F900" ID-GBIF-Occurrence="2028137900" collectingDate="2018-04-22" collectorName="Huo &amp; Gao Peng" country="China" county="Lin'an City" elevation="861" latitude="30.337133" location="Wuliting" longLatPrecision="1" longitude="119.4356" municipality="Tianmu Mountain Nature Reserve" pageId="4" pageNumber="280" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Zhejiang" typeStatus="paratype">
<typeStatus id="54A775A3851FA445A5D330326139F924" box="[1192,1305,1649,1673]" pageId="4" pageNumber="280" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D1A0088851FA445A4533032616CF924" box="[1320,1356,1649,1673]" pageId="4" pageNumber="280" type="male">1♂</specimenCount>
,
<specimenCount id="9D1A0088851FA445A426303261B7F924" box="[1373,1431,1649,1673]" pageId="4" pageNumber="280" type="female">2♀♀</specimenCount>
, similar coloration with the
<typeStatus id="54A775A3851FA445A0BD30D7660BF900" box="[454,555,1684,1709]" pageId="4" pageNumber="280" type="holotype">holotype</typeStatus>
; the same locality and data as
<typeStatus id="54A775A3851FA445A20430D767C5F900" box="[895,997,1684,1709]" pageId="4" pageNumber="280" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C306988A851FA445A1BC30FB6151F97C" box="[199,1393,1720,1745]" pageId="4" pageNumber="280" type="etymology">
<paragraph id="8BA3CB01851FA445A1BC30FB6151F97C" blockId="4.[151,1437,1612,2036]" box="[199,1393,1720,1745]" pageId="4" pageNumber="280">
<emphasis id="B9681713851FA445A1BC30FB6571F97C" bold="true" box="[199,337,1720,1745]" pageId="4" pageNumber="280">Etymology:</emphasis>
The scientific name indicates the sclerotized developed process on the tergum 10 of the adult.
</paragraph>
</subSubSection>
<subSubSection id="C306988A851FA447A1BC309F6606FF78" lastPageId="6" lastPageNumber="282" pageId="4" pageNumber="280" type="discussion">
<paragraph id="8BA3CB01851FA445A1BC309F6087F859" blockId="4.[151,1437,1612,2036]" pageId="4" pageNumber="280">
<emphasis id="B9681713851FA445A1BC309F656EF958" bold="true" box="[199,334,1756,1781]" pageId="4" pageNumber="280">Discussion:</emphasis>
Of the 14 known Chinese
<taxonomicName id="4C1CB082851FA445A304309E66FCF958" authorityName="Banks" authorityYear="1906" box="[639,732,1757,1781]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B9681713851FA445A304309E66FCF958" box="[639,732,1757,1781]" italics="true" pageId="4" pageNumber="280">Isoperla</emphasis>
</taxonomicName>
species, the new species is apparently most similar to
<taxonomicName id="4C1CB082851FA445A43E309E6529F8B4" authority="Wu, 1935" authorityName="Wu" authorityYear="1935" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="B9681713851FA445A43E309E61BBF959" box="[1349,1435,1757,1780]" italics="true" pageId="4" pageNumber="280">I. yangi</emphasis>
<bibRefCitation id="EF8DB6F0851FA445A1EC31426529F8B4" author="Wu, C. F." box="[151,265,1792,1817]" pageId="4" pageNumber="280" pagination="227 - 243" refId="ref1965" refString="Wu, C. F. (1935) New species of stoneflies from east and south China. Peking Natural History Bulletin, 9 (3), 227 - 243." type="journal article" year="1935">Wu, 1935</bibRefCitation>
</taxonomicName>
described from
<collectingRegion id="49D805E3851FA445A0B6314366AEF8B4" box="[461,654,1792,1817]" country="China" name="Jiangxi" pageId="4" pageNumber="280">Jiangxi Province</collectingRegion>
of southeastern
<collectingCountry id="F30B8B91851FA445A22F314367BBF8B4" box="[852,923,1792,1817]" name="China" pageId="4" pageNumber="280">China</collectingCountry>
. Currently, the male of new species can be distinguished by the shape of the vesicle. In
<taxonomicName id="4C1CB082851FA445A3F7316666C2F891" authorityName="Wu" authorityYear="1935" box="[652,738,1829,1852]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="B9681713851FA445A3F7316666C2F891" box="[652,738,1829,1852]" italics="true" pageId="4" pageNumber="280">I. yangi</emphasis>
</taxonomicName>
, the vesicle is well-developed and ovoid in shape (
<bibRefCitation id="EF8DB6F0851FA445A458316661AFF890" author="Wu, C. F." box="[1315,1423,1829,1853]" pageId="4" pageNumber="280" refId="ref1998" refString="Wu, C. F. (1938) Plecopterorum sinensium: A monograph of stoneflies of China (order Plecoptera). Yenching University, Beijing, 225 pp." type="book" year="1938">Wu 1938</bibRefCitation>
), where as in
<taxonomicName id="4C1CB082851FA445A065310F6591F8CE" authorityName="Huo &amp; Du" authorityYear="2018" box="[286,433,1867,1891]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="oncocauda">
<emphasis id="B9681713851FA445A065310F6591F8CE" box="[286,433,1867,1891]" italics="true" pageId="4" pageNumber="280">I. oncocauda</emphasis>
</taxonomicName>
the vesicle is reduced and only slightly projecting beyond anterior margin of 9
<superScript id="7C696649851FA445A45C310B6113F8FB" attach="left" box="[1319,1331,1864,1878]" fontSize="6" pageId="4" pageNumber="280">th</superScript>
sternum. The female apparently can be distinguished from all known Chinese
<taxonomicName id="4C1CB082851FA445A2D0312C6028F82A" authorityName="Banks" authorityYear="1906" box="[939,1032,1903,1927]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B9681713851FA445A2D0312C6028F82A" box="[939,1032,1903,1927]" italics="true" pageId="4" pageNumber="280">Isoperla</emphasis>
</taxonomicName>
females by the posterior margin of tergum 10 extending as a process. Of 14 Chinese species, the male aedeagi of only two northern Chinese species,
<taxonomicName id="4C1CB082851FA445A4F731D76623F87D" authority="Zapekina-Dulkeit, 1975" authorityName="Zapekina-Dulkeit" authorityYear="1975" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="eximia">
<emphasis id="B9681713851FA445A4F731D764C2F862" italics="true" pageId="4" pageNumber="280">I. eximia</emphasis>
Zapekina-Dulkeit, 1975
</taxonomicName>
and
<taxonomicName id="4C1CB082851FA445A33331FB67B9F862" authority="(Klapalek, 1923)" authorityName="Klapalek" authorityYear="1923" baseAuthorityName="Klapalek" baseAuthorityYear="1923" box="[584,921,1975,2000]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="lunigera">
<emphasis id="B9681713851FA445A33331FB66E5F862" box="[584,709,1975,1999]" italics="true" pageId="4" pageNumber="280">I. lunigera</emphasis>
(
<bibRefCitation id="EF8DB6F0851FA445A3A031F467B0F862" author="Klapalek, F." box="[731,912,1975,2000]" pageId="4" pageNumber="280" pagination="25 - 26" refId="ref1734" refString="Klapalek, F. (1923) Plecopteres nouveaux. Annales de la Societe Entomologique de Belgique, 63, 25 - 26." type="journal article" year="1923">Klapálek, 1923</bibRefCitation>
)
</taxonomicName>
have been illustrated in part (
<bibRefCitation id="EF8DB6F0851FA445A46A31F46566F859" author="Teslenko, V. A. &amp; Zhiltzova, L. A." pageId="4" pageNumber="280" refId="ref1914" refString="Teslenko, V. A. &amp; Zhiltzova, L. A. (2009) Key to the stoneflies (Insecta, Plecoptera) of Russia and adjacent countries. Imagines and nymphs. Russian Academy of Sciences, Far Eastern Branch, Dalnauka, Vladivostok, 381 pp." type="book" year="2009">Teslenko &amp; Zhiltzova 2009</bibRefCitation>
). In both of these species, the aedeagus possess distinctive aedeagal sclerites.
</paragraph>
<caption id="DF639B89851EA444A1EC322760B7FBD6" box="[151,1175,1124,1147]" httpUri="https://zenodo.org/record/2606301/files/figure.png" pageId="5" pageNumber="281" startId="5.[151,264,1124,1147]" targetBox="[257,1395,193,1108]" targetPageId="5">
<paragraph id="8BA3CB01851EA444A1EC322760B7FBD6" blockId="5.[151,1175,1124,1147]" box="[151,1175,1124,1147]" pageId="5" pageNumber="281">
<emphasis id="B9681713851EA444A1EC32276573FBD7" bold="true" box="[151,339,1124,1147]" pageId="5" pageNumber="281">FIGURES 13–14.</emphasis>
<taxonomicName id="4C1CB082851EA444A02132266603FBD6" authority="Huo &amp; Du, 2018" authorityName="Huo &amp; Du" authorityYear="2018" box="[346,547,1125,1147]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="oncocauda" status="sp. nov.">
<emphasis id="B9681713851EA444A02132266603FBD6" box="[346,547,1125,1147]" italics="true" pageId="5" pageNumber="281">Isoperla oncocauda</emphasis>
</taxonomicName>
,
<emphasis id="B9681713851EA444A34B322566A0FBD6" bold="true" box="[560,640,1126,1147]" pageId="5" pageNumber="281">
<taxonomicNameLabel id="A25BAA68851EA444A34B322566A0FBD6" box="[560,640,1126,1147]" pageId="5" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
13: Female habitus; 14: Female head and pronotum.
</paragraph>
</caption>
<caption id="DF639B89851EA444A1EC319A60E3F842" box="[151,1219,2009,2032]" httpUri="https://zenodo.org/record/2606303/files/figure.png" pageId="5" pageNumber="281" startId="5.[151,264,2009,2032]" targetBox="[188,1400,1187,1996]" targetPageId="5">
<paragraph id="8BA3CB01851EA444A1EC319A60E3F842" blockId="5.[151,1219,2009,2032]" box="[151,1219,2009,2032]" pageId="5" pageNumber="281">
<emphasis id="B9681713851EA444A1EC319A6573F843" bold="true" box="[151,339,2009,2032]" pageId="5" pageNumber="281">FIGURES 15–16.</emphasis>
<taxonomicName id="4C1CB082851EA444A021319A6603F842" authority="Huo &amp; Du, 2018" authorityName="Huo &amp; Du" authorityYear="2018" box="[346,547,2009,2031]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="oncocauda" status="sp. nov.">
<emphasis id="B9681713851EA444A021319A6603F842" box="[346,547,2009,2031]" italics="true" pageId="5" pageNumber="281">Isoperla oncocauda</emphasis>
</taxonomicName>
,
<emphasis id="B9681713851EA444A34B319966A0F842" bold="true" box="[560,640,2010,2031]" pageId="5" pageNumber="281">
<taxonomicNameLabel id="A25BAA68851EA444A34B319966A0F842" box="[560,640,2010,2031]" pageId="5" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, terminalia of female. 15: Dorsal view; 16: Ventral view.
</paragraph>
</caption>
<paragraph id="8BA3CB01851DA447A1BC36D46606FF78" blockId="6.[151,1436,151,213]" pageId="6" pageNumber="282">
The shallow stream of the
<typeStatus id="54A775A3851DA447A08A36DB6601FF1D" box="[497,545,152,176]" pageId="6" pageNumber="282">type</typeStatus>
locality flows among the large rocks (
<figureCitation id="1327D784851DA447A2AB36D46003FF1D" box="[976,1059,151,176]" captionStart="FIGURE 19" captionStartId="7.[151,250,2000,2022]" captionTargetBox="[150,1436,265,1979]" captionTargetId="figure@7.[150,1436,265,1979]" captionTargetPageId="7" captionText="FIGURE 19. Habitat of Isoperla oncocauda, sp. nov." httpUri="https://zenodo.org/record/2606307/files/figure.png" pageId="6" pageNumber="282">Fig. 19</figureCitation>
). Adults were collected sitting on the rocks and from a nearby bridge.
</paragraph>
</subSubSection>
</treatment>
</document>
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