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<document id="DC10544243A1D6965B261A4C18B3C4A2" ID-CLB-Dataset="6634" ID-DOI="10.5852/ejt.2021.749.1355" ID-GBIF-Dataset="83109e54-596a-4be0-ac2c-c4e1702cfb8a" ID-ISSN="2118-9773" ID-Zenodo-Dep="4770293" ID-ZooBank="25FD5744-861D-45E4-B6AD-350716AA29BF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621349695117" checkinUser="felipe" docAuthor="Gao, Chuan-bu &amp; Coca-Abia, María Milagro" docDate="2021" docId="03B56743A67C0666FDE3F96BFCF3E656" docLanguage="en" docName="ejt-749_1-94.pdf" docOrigin="European Journal of Taxonomy 749" docSource="http://dx.doi.org/10.5852/ejt.2021.749.1355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Miridiba scutata Gao &amp; Coca-Abia 2021, comb. nov." docType="treatment" docVersion="9" lastPageNumber="77" masterDocId="FF8C1F3BA635062AFFFBFFA9FFE2E431" masterDocTitle="Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data" masterLastPageNumber="94" masterPageNumber="1" pageNumber="74" updateTime="1698921317801" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5AEC8BAC25A79089ED6E2D74BB88CC08">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:namePart id="7E8639A7826291FD855B836FA9C21FED">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="87330E07CE5B2BE8034665CBB0AD1A4D">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
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<mods:namePart id="F14CB10CE09F3CA02D116CDDDAC1B7B1">Coca-Abia, María Milagro</mods:namePart>
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<mods:date id="78AB0AF50E11C74B8E8D6675FEF6EE0C">2021</mods:date>
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<treatment id="03B56743A67C0666FDE3F96BFCF3E656" ID-DOI="http://doi.org/10.5281/zenodo.4773336" ID-GBIF-Taxon="182237204" ID-Zenodo-Dep="4773336" LSID="urn:lsid:plazi:treatment:03B56743A67C0666FDE3F96BFCF3E656" httpUri="http://treatment.plazi.org/id/03B56743A67C0666FDE3F96BFCF3E656" lastPageId="76" lastPageNumber="77" pageId="73" pageNumber="74">
<subSubSection id="C30685DEA67C0663FDE3F96BFBFEE2EC" box="[536,1052,1730,1757]" pageId="73" pageNumber="74" type="nomenclature">
<paragraph id="8BA3D655A67C0663FDE3F96BFBFEE2EC" blockId="73.[536,1052,1730,1792]" box="[536,1052,1730,1757]" pageId="73" pageNumber="74">
<heading id="D0EB6139A67C0663FDE3F96BFBFEE2EC" box="[536,1052,1730,1757]" centered="true" fontSize="11" level="2" pageId="73" pageNumber="74" reason="2">
<taxonomicName id="4C1CADD6A67C0663FDE3F96BFC70E2EC" authority="(Reitter, 1902)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Reitter" baseAuthorityYear="1902" box="[536,914,1730,1757]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="scutata" status="comb. nov.">
<emphasis id="B9680A47A67C0663FDE3F96BFD3DE2EC" bold="true" box="[536,735,1730,1757]" italics="true" pageId="73" pageNumber="74">Miridiba scutata</emphasis>
(
<bibRefCitation id="EF8DABA4A67C0663FD14F96AFC68E2EC" author="Reitter E." box="[751,906,1731,1757]" pageId="73" pageNumber="74" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25BB73CA67C0663FC61F96AFBFEE2EC" box="[922,1052,1731,1757]" pageId="73" pageNumber="74" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA67C0663FD39F94FFC93E331" box="[706,881,1766,1792]" pageId="73" pageNumber="74" type="description">
<paragraph id="8BA3D655A67C0663FD39F94FFC93E331" blockId="73.[536,1052,1730,1792]" box="[706,881,1766,1792]" pageId="73" pageNumber="74">
<figureCitation id="1327CAD0A67C0663FD39F94FFCAAE331" box="[706,840,1766,1792]" captionStart="Figs 20–22" captionStartId="14.[189,241,1251,1277]" captionTargetBox="[291,1324,285,1192]" captionTargetId="figure-142@14.[278,1337,265,1205]" captionTargetPageId="14" captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="73" pageNumber="74">Figs 20–22</figureCitation>
,
<figureCitation id="1327CAD0A67C0663FCAFF94FFC93E331" box="[852,881,1766,1792]" captionStart="Figs 94–97. 94" captionStartId="75.[189,241,1887,1913]" captionTargetBox="[277,1301,581,1832]" captionTargetId="figure-102@75.[264,1324,560,1851]" captionTargetPageId="75" captionText="Figs 94–97. 94. Miridiba scutata (Reitter, 1902), lectotype, ♂ (MFNB). Habitus in drosal view. 95– 97. M. schoolmeesteri Keith, 2010, holotype, ♂ (DKC). 95. Habitus in dorsal view. 96. Parameres in dorsal (A) and lateral (B) views. 97. Endophallus and temones in lateral view. Scale bars: 94–95 = 5.0 mm; 96–97 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770413" httpUri="https://zenodo.org/record/4770413/files/figure.png" pageId="73" pageNumber="74">94</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA67C0660FF46F88EFE49E5BC" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="reference_group">
<paragraph id="8BA3D655A67C0663FF46F88EFC10E373" blockId="73.[189,1399,1831,1964]" box="[189,1010,1831,1858]" pageId="73" pageNumber="74">
<treatmentCitationGroup id="AB0CF17BA67C0663FF46F88EFC10E373" box="[189,1010,1831,1858]" pageId="73" pageNumber="74">
<treatmentCitation id="0ABDF044A67C0663FF46F88EFD9FE373" author="Reitter E." box="[189,637,1831,1858]" page="175" pageId="73" pageNumber="74" year="1902">
<taxonomicName id="4C1CADD6A67C0663FF46F88EFD9FE373" authority="Reitter, 1902: 175" authorityName="Reitter" authorityPageNumber="175" authorityYear="1902" box="[189,637,1831,1858]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67C0663FF46F88EFE40E370" box="[189,418,1831,1857]" italics="true" pageId="73" pageNumber="74">Holotrichia scutata</emphasis>
<bibRefCitation id="EF8DABA4A67C0663FE52F88EFD9FE373" author="Reitter E." box="[425,637,1831,1858]" pageId="73" pageNumber="74" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902: 175</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54A768F7A67C0663FD77F88EFD22E370" box="[652,704,1831,1857]" pageId="73" pageNumber="74">type</typeStatus>
loc.:
<collectingCountry id="F30B96C5A67C0663FCF9F881FC6DE373" box="[770,911,1832,1858]" name="Hong Kong" pageId="73" pageNumber="74">Hong Kong</collectingCountry>
,
<collectingCountry id="F30B96C5A67C0663FC60F88EFC04E373" box="[923,998,1831,1858]" name="China" pageId="73" pageNumber="74">China</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A67C0663FF46F8E3FB5CE39A" blockId="73.[189,1399,1831,1964]" pageId="73" pageNumber="74">
<treatmentCitationGroup id="AB0CF17BA67C0663FF46F8E3FB5CE39A" pageId="73" pageNumber="74">
<treatmentCitation id="0ABDF044A67C0663FF46F8E3FD9AE354" author="Moser J." box="[189,632,1866,1893]" page="471" pageId="73" pageNumber="74" year="1909">
<taxonomicName id="4C1CADD6A67C0663FF46F8E3FD9AE354" authority="Moser, 1909: 471" authorityName="Moser" authorityPageNumber="471" authorityYear="1909" box="[189,632,1866,1893]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67C0663FF46F8E3FE40E355" box="[189,418,1866,1892]" italics="true" pageId="73" pageNumber="74">Holotrichia scutata</emphasis>
<bibRefCitation id="EF8DABA4A67C0663FE52F8E2FD9AE354" author="Moser J." box="[425,632,1867,1893]" pageId="73" pageNumber="74" pagination="468 - 488" refId="ref45295" refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488." type="journal article" year="1909">Moser, 1909: 471</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(primary homonym of
<taxonomicName id="4C1CADD6A67C0663FC70F8E2FB45E354" authority="Reitter, 1902" authorityName="Reitter" authorityYear="1902" box="[907,1191,1866,1893]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67C0663FC70F8E2FBE6E355" box="[907,1028,1867,1892]" italics="true" pageId="73" pageNumber="74">H. scutata</emphasis>
<bibRefCitation id="EF8DABA4A67C0663FBF0F8E3FB45E354" author="Reitter E." box="[1035,1191,1866,1893]" pageId="73" pageNumber="74" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
</taxonomicName>
). syn. nov.
<treatmentCitation id="0ABDF044A67C0663FF46F8C7FD2BE3B9" author="Dalla Torre K. W. von" box="[189,713,1902,1928]" page="206" pageId="73" pageNumber="74" year="1912">
<taxonomicName id="4C1CADD6A67C0663FF46F8C7FD2BE3B9" authority="Dalla Torre, 1912: 206" authorityName="Dalla Torre" authorityPageNumber="206" authorityYear="1912" box="[189,713,1902,1928]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="scutulata">
<emphasis id="B9680A47A67C0663FF46F8C7FE5BE3B9" box="[189,441,1902,1928]" italics="true" pageId="73" pageNumber="74">Holotrichia scutulata</emphasis>
<bibRefCitation id="EF8DABA4A67C0663FE44F8C7FD2BE3B9" author="Dalla Torre K. W. von" box="[447,713,1902,1928]" pageId="73" pageNumber="74" pagination="135 - 290" refId="ref43620" refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany." type="book chapter" year="1912">Dalla Torre, 1912: 206</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(new name of
<taxonomicName id="4C1CADD6A67C0663FC8CF8C7FB18E3B9" authority="Moser, 1909" authorityName="Moser" authorityYear="1909" box="[887,1274,1902,1928]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67C0663FC8CF8C7FBBEE3B9" box="[887,1116,1902,1928]" italics="true" pageId="73" pageNumber="74">Holotrichia scutata</emphasis>
<bibRefCitation id="EF8DABA4A67C0663FB98F8C7FB18E3B9" author="Moser J." box="[1123,1274,1902,1928]" pageId="73" pageNumber="74" pagination="468 - 488" refId="ref45295" refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488." type="journal article" year="1909">Moser, 1909</bibRefCitation>
</taxonomicName>
). syn. nov.
<taxonomicName id="4C1CADD6A67C0663FF46F838FDBDE39A" authority="Keith, 2010: 237" authorityName="Keith" authorityPageNumber="237" authorityYear="2010" box="[189,607,1937,1963]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="B9680A47A67C0663FF46F838FE6DE39A" box="[189,399,1937,1963]" italics="true" pageId="73" pageNumber="74">Miridiba lassallei</emphasis>
Keith, 2010: 237
</taxonomicName>
, figs 9–11 (
<typeStatus id="54A768F7A67C0663FD1EF838FCFBE39A" box="[741,793,1937,1963]" pageId="73" pageNumber="74">type</typeStatus>
loc.: Nghia Lo,
<collectingCountry id="F30B96C5A67C0663FC22F838FBA0E39D" box="[985,1090,1937,1964]" name="Vietnam" pageId="73" pageNumber="74">Vietnam</collectingCountry>
). syn. nov.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A67F0660FF46FEA0FC39E515" blockId="74.[189,1399,265,397]" box="[189,987,265,292]" pageId="74" pageNumber="75">
<treatmentCitationGroup id="AB0CF17BA67F0660FF46FEA0FC39E515" box="[189,987,265,292]" pageId="74" pageNumber="75">
<taxonomicName id="4C1CADD6A67F0660FF46FEA0FE1BE512" authority="Reitter" authorityName="Reitter" authorityYear="1902" box="[189,505,265,291]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FF46FEA0FE40E512" box="[189,418,265,291]" italics="true" pageId="74" pageNumber="75">Holotrichia scutata</emphasis>
Reitter
</taxonomicName>
–
<treatmentCitation id="0ABDF044A67F0660FDEDFEA0FD03E512" author="Chang Y. - W." box="[534,737,265,291]" page="149" pageId="74" pageNumber="75" year="1964">
<bibRefCitation id="EF8DABA4A67F0660FDEDFEA0FD03E512" author="Chang Y. - W." box="[534,737,265,291]" pageId="74" pageNumber="75" pagination="139 - 152" refId="ref43461" refString="Chang Y. - W. 1964. Revision of Chinese may beetles of the genus Holotrichia Hope I. (Coleoptera, Scarabaeidae). Acta Zootaxonomica Sinica 1 (1): 139 - 152." type="journal article" year="1964">Chang 1964: 149</bibRefCitation>
</treatmentCitation>
(species list; in key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A67F0660FF46FE85FC03E576" blockId="74.[189,1399,265,397]" box="[189,993,300,327]" pageId="74" pageNumber="75">
<treatmentCitationGroup id="AB0CF17BA67F0660FF46FE85FC03E576" box="[189,993,300,327]" pageId="74" pageNumber="75">
<taxonomicName id="4C1CADD6A67F0660FF46FE85FE16E576" authority="Moser" authorityName="Moser" authorityYear="1909" box="[189,500,300,327]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FF46FE85FE40E577" box="[189,418,300,326]" italics="true" pageId="74" pageNumber="75">Holotrichia scutata</emphasis>
Moser
</taxonomicName>
–
<treatmentCitation id="0ABDF044A67F0660FDE9FE85FCACE577" author="Smetana A. &amp; Kral D." box="[530,846,300,326]" page="220" pageId="74" pageNumber="75" year="2006">
<bibRefCitation id="EF8DABA4A67F0660FDE9FE85FCACE577" author="Smetana A. &amp; Kral D." box="[530,846,300,326]" pageId="74" pageNumber="75" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 220</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A67F0660FF46FEE6FE49E5BC" blockId="74.[189,1399,265,397]" pageId="74" pageNumber="75">
<treatmentCitationGroup id="AB0CF17BA67F0660FF46FEE6FE49E5BC" pageId="74" pageNumber="75">
<taxonomicName id="4C1CADD6A67F0660FF46FEE6FDA8E55B" authority="Dalla Torre" authorityName="Dalla Torre" authorityYear="1912" box="[189,586,335,362]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutulata">
<emphasis id="B9680A47A67F0660FF46FEE6FE59E558" box="[189,443,335,361]" italics="true" pageId="74" pageNumber="75">Holotrichia scutulata</emphasis>
Dalla Torre
</taxonomicName>
–
<treatmentCitation id="0ABDF044A67F0660FD90FEE6FCDBE55B" author="Chang Y. - W." box="[619,825,335,362]" page="149" pageId="74" pageNumber="75" year="1964">
<bibRefCitation id="EF8DABA4A67F0660FD90FEE6FCDBE55B" author="Chang Y. - W." box="[619,825,335,362]" pageId="74" pageNumber="75" pagination="139 - 152" refId="ref43461" refString="Chang Y. - W. 1964. Revision of Chinese may beetles of the genus Holotrichia Hope I. (Coleoptera, Scarabaeidae). Acta Zootaxonomica Sinica 1 (1): 139 - 152." type="journal article" year="1964">Chang 1964: 149</bibRefCitation>
</treatmentCitation>
(species list; in key). —
<treatmentCitation id="0ABDF044A67F0660FB92FEF9FEFAE5BC" author="Smetana A. &amp; Kral D." page="220" pageId="74" pageNumber="75" year="2006">
<bibRefCitation id="EF8DABA4A67F0660FB92FEF9FEFAE5BC" author="Smetana A. &amp; Kral D." pageId="74" pageNumber="75" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 220</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA67F0660FF46FE1DFC3AE047" pageId="74" pageNumber="75" type="diagnosis">
<paragraph id="8BA3D655A67F0660FF46FE1DFED4E5FF" blockId="74.[189,310,436,462]" box="[189,310,436,462]" pageId="74" pageNumber="75">
<heading id="D0EB6139A67F0660FF46FE1DFED4E5FF" bold="true" box="[189,310,436,462]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="B9680A47A67F0660FF46FE1DFED4E5FF" bold="true" box="[189,310,436,462]" pageId="74" pageNumber="75">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A67F0660FF46FE4BFC3AE047" blockId="74.[188,1401,482,1143]" pageId="74" pageNumber="75">
External morphology of adult (
<figureCitation id="1327CAD0A67F0660FDCEFE4BFD72E5CC" box="[565,656,482,509]" captionStart="Figs 94–97. 94" captionStartId="75.[189,241,1887,1913]" captionTargetBox="[277,1301,581,1832]" captionTargetId="figure-102@75.[264,1324,560,1851]" captionTargetPageId="75" captionText="Figs 94–97. 94. Miridiba scutata (Reitter, 1902), lectotype, ♂ (MFNB). Habitus in drosal view. 95– 97. M. schoolmeesteri Keith, 2010, holotype, ♂ (DKC). 95. Habitus in dorsal view. 96. Parameres in dorsal (A) and lateral (B) views. 97. Endophallus and temones in lateral view. Scale bars: 94–95 = 5.0 mm; 96–97 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770413" httpUri="https://zenodo.org/record/4770413/files/figure.png" pageId="74" pageNumber="75">Fig. 94</figureCitation>
). Body size 14.0–16.0 mm. Dorsal surface glabrous. Clypeus rounded laterally, with conspicuous punctures, anterior edge emarginate at middle. Frons with carina moderately developed and blunt. Pronotal surface shiny, regularly punctate, with punctures closer together just behind anterior margin, glabrous, at most with a tiny seta on each puncture, anterior pronotal margin flanged, glabrous; posterior margin glabrous; lateral margins serrated and pubescent; anterior angles almost right not projected forward, posterior angles obtuse and marked. Scutellum without punctures and glabrous. Elytral surface glabrous, at most with a tiny seta on each puncture, with striations weakly defined, punctures distributed between striae; epipleuron glabrous, at most with few setae at base. Foretibia with dorsal carina moderately developed; insertion of inner spur closer third outer tooth than second one. Meso- and metatibia with transverse carina complete, inner margin of dorsal surface without spines. Pygidium with deep punctuation, surface with inconspicuous pubescence, margin pubescent. Ventrites with conspicuous punctures; ventrites 2, 3 and 4 glabrous, at most with a tiny seta in each puncture; ventrite 5 with long pubescence irregularly distributed at posterior half; ventrite 6 moderately bulging at anterior margin, with long pubescence on posterior margin. Male genitalia: parameres (
<figureCitation id="1327CAD0A67F0660FE46FC66FDF6E7D8" box="[445,532,975,1001]" captionStart="Figs 20–22" captionStartId="14.[189,241,1251,1277]" captionTargetBox="[291,1324,285,1192]" captionTargetId="figure-142@14.[278,1337,265,1205]" captionTargetPageId="14" captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="74" pageNumber="75">Fig. 20</figureCitation>
) with dorsal and ventral branches joined laterally. Dorsal branches fused with apex asymmetrical, elevated and twisted. Ventral branches reduced, lightly sclerotized and fused by a ventral membrane. Endophallus (
<figureCitation id="1327CAD0A67F0660FDA3FBBFFD4CE001" box="[600,686,1046,1072]" captionStart="Figs 20–22" captionStartId="14.[189,241,1251,1277]" captionTargetBox="[291,1324,285,1192]" captionTargetId="figure-142@14.[278,1337,265,1205]" captionTargetPageId="14" captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="74" pageNumber="75">Fig. 21</figureCitation>
) with a raspula with soft setae and spines at distal end; with spines isolated at middle. Apophysis of temones (
<figureCitation id="1327CAD0A67F0660FCF2FB90FCBDE062" box="[777,863,1081,1107]" captionStart="Figs 20–22" captionStartId="14.[189,241,1251,1277]" captionTargetBox="[291,1324,285,1192]" captionTargetId="figure-142@14.[278,1337,265,1205]" captionTargetPageId="14" captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="74" pageNumber="75">Fig. 21</figureCitation>
) long and separated from each other. Female genitalia (
<figureCitation id="1327CAD0A67F0660FEC8FBF5FE6BE047" box="[307,393,1116,1142]" captionStart="Figs 20–22" captionStartId="14.[189,241,1251,1277]" captionTargetBox="[291,1324,285,1192]" captionTargetId="figure-142@14.[278,1337,265,1205]" captionTargetPageId="14" captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="74" pageNumber="75">Fig. 22</figureCitation>
) as described above (morphotype VIII “
<emphasis id="B9680A47A67F0660FC9AFBF4FC5BE046" box="[865,953,1117,1143]" italics="true" pageId="74" pageNumber="75">Scutata</emphasis>
”).
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA67F0660FF46FB34FCD8E2F4" pageId="74" pageNumber="75" type="materials_examined">
<paragraph id="8BA3D655A67F0660FF46FB34FE48E086" blockId="74.[189,426,1181,1207]" box="[189,426,1181,1207]" pageId="74" pageNumber="75">
<heading id="D0EB6139A67F0660FF46FB34FE48E086" bold="true" box="[189,426,1181,1207]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="B9680A47A67F0660FF46FB34FE48E086" bold="true" box="[189,426,1181,1207]" pageId="74" pageNumber="75">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A67F0660FF11FB65FCB3E0D6" blockId="74.[189,1399,1228,1360]" box="[234,849,1228,1255]" pageId="74" pageNumber="75">
<emphasis id="B9680A47A67F0660FF11FB65FD9DE0D7" bold="true" box="[234,639,1228,1254]" pageId="74" pageNumber="75">
<typeStatus id="54A768F7A67F0660FF11FB65FE84E0D7" box="[234,358,1228,1254]" pageId="74" pageNumber="75" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C1CADD6A67F0660FE76FB65FD9DE0D7" authority="(Reitter)" baseAuthorityName="Reitter" baseAuthorityYear="1902" box="[397,639,1228,1254]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FE76FB65FDEEE0D7" bold="true" box="[397,524,1228,1254]" italics="true" pageId="74" pageNumber="75">M. scutata</emphasis>
(Reitter)
</taxonomicName>
</emphasis>
(here designated)
</paragraph>
<paragraph id="8BA3D655A67F0660FF46FB46FEFAE161" blockId="74.[189,1399,1228,1360]" pageId="74" pageNumber="75">
<collectingCountry id="F30B96C5A67F0660FF46FB46FEFAE138" box="[189,280,1263,1289]" name="China" pageId="74" pageNumber="75">CHINA</collectingCountry>
•
<specimenCount id="9D1A1DDCA67F0660FEC8FB59FEAEE13B" box="[307,332,1264,1290]" pageId="74" pageNumber="75" type="male">♂</specimenCount>
; “Hong Kong”; “
<taxonomicName id="4C1CADD6A67F0660FDE1FB59FD42E138" authority="Rtt" authorityName="Reitter" authorityYear="1902" box="[538,672,1263,1289]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FDE1FB59FD8DE138" box="[538,623,1264,1289]" italics="true" pageId="74" pageNumber="75">scutata</emphasis>
Rtt
</taxonomicName>
”; “?
<typeStatus id="54A768F7A67F0660FD21FB46FCBCE138" box="[730,862,1263,1289]" pageId="74" pageNumber="75" type="syntype">SYNTYPE</typeStatus>
<taxonomicName id="4C1CADD6A67F0660FC9CFB46FB0FE138" authority="Reitter, 1902" authorityName="Reitter" authorityYear="1902" box="[871,1261,1263,1289]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FC9CFB46FBAFE138" box="[871,1101,1263,1289]" italics="true" pageId="74" pageNumber="75">Holotrichia scutata</emphasis>
<bibRefCitation id="EF8DABA4A67F0660FBAFFB46FB0FE138" author="Reitter E." box="[1108,1261,1263,1289]" pageId="74" pageNumber="75" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
</taxonomicName>
labelled by
<collectionCode id="ED0D4E90A67F0660FF45FABAFEF3E11C" box="[190,273,1299,1325]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="74" pageNumber="75">MFNB</collectionCode>
2018”; “
<typeStatus id="54A768F7A67F0660FE84FABAFE15E11C" box="[383,503,1299,1325]" pageId="74" pageNumber="75" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C1CADD6A67F0660FDF9FABBFC60E11C" authority="(Reitter, 1902)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Reitter" baseAuthorityYear="1902" box="[514,898,1298,1325]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FDF9FABBFD2BE11D" box="[514,713,1298,1324]" italics="true" pageId="74" pageNumber="75">Miridiba scutata</emphasis>
(
<bibRefCitation id="EF8DABA4A67F0660FD20FABBFC98E11C" author="Reitter E." box="[731,890,1298,1325]" pageId="74" pageNumber="75" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
)
</taxonomicName>
Chuan-bu Gao &amp; Coca-Abia det. 2019”;
<collectionCode id="ED0D4E90A67F0660FF46FA9FFEF6E161" box="[189,276,1334,1360]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="74" pageNumber="75">MFNB</collectionCode>
.
</paragraph>
<paragraph id="8BA3D655A67F0660FF11FADEFCABE1A3" blockId="74.[187,1398,1399,1496]" box="[234,841,1399,1426]" pageId="74" pageNumber="75">
<emphasis id="B9680A47A67F0660FF11FADEFD95E1A0" bold="true" box="[234,631,1399,1425]" pageId="74" pageNumber="75">
<typeStatus id="54A768F7A67F0660FF11FADEFE84E1A0" box="[234,358,1399,1425]" pageId="74" pageNumber="75" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C1CADD6A67F0660FE76FADEFD95E1A0" authority="(Moser)" baseAuthorityName="Moser" box="[397,631,1399,1425]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FE76FADEFDEEE1A0" bold="true" box="[397,524,1399,1425]" italics="true" pageId="74" pageNumber="75">M. scutata</emphasis>
(Moser)
</taxonomicName>
</emphasis>
(here designated)
</paragraph>
<paragraph id="8BA3D655A67F0660FF46FA33FB24E1E9" blockId="74.[187,1398,1399,1496]" pageId="74" pageNumber="75">
<materialsCitation id="3B74DC08A67F0660FF46FA33FB20E1E9" ID-GBIF-Occurrence="3114121385" collectionCode="MFNB" collectorName="Brenske" country="China" determinerName="Chuan-bu Gao &amp; Coca-Abia" location="CHINA" pageId="74" pageNumber="75" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<collectingCountry id="F30B96C5A67F0660FF46FA33FEFAE185" box="[189,280,1434,1460]" name="China" pageId="74" pageNumber="75">CHINA</collectingCountry>
•
<specimenCount id="9D1A1DDCA67F0660FED6FA32FEA4E184" box="[301,326,1435,1461]" pageId="74" pageNumber="75" type="male">♂</specimenCount>
; “
<collectingCountry id="F30B96C5A67F0660FEA1FA33FE08E185" box="[346,490,1434,1460]" name="Hong Kong" pageId="74" pageNumber="75">Hong Kong</collectingCountry>
”; “Coll
<collectorName id="26E9B383A67F0660FDB8FA33FD48E185" box="[579,682,1434,1460]" pageId="74" pageNumber="75">Brenske</collectorName>
”; “
<taxonomicName id="4C1CADD6A67F0660FD31FA32FCA3E185" box="[714,833,1435,1460]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FD31FA32FCA3E185" box="[714,833,1435,1460]" italics="true" pageId="74" pageNumber="75">H. scutata</emphasis>
</taxonomicName>
Type Mos”, “
<taxonomicName id="4C1CADD6A67F0660FC24FA32FBD6E185" box="[991,1076,1435,1460]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FC24FA32FBD6E185" box="[991,1076,1435,1460]" italics="true" pageId="74" pageNumber="75">scutata</emphasis>
</taxonomicName>
” [label in pink]; “Lectotype
<taxonomicName id="4C1CADD6A67F0660FF40FA14FD60E1E9" authority="(Dalla Torre, 1912)" baseAuthorityName="Dalla Torre" baseAuthorityYear="1912" box="[187,642,1469,1496]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutulata">
<emphasis id="B9680A47A67F0660FF40FA14FE74E1E6" box="[187,406,1469,1495]" italics="true" pageId="74" pageNumber="75">Miridiba scutulata</emphasis>
(
<bibRefCitation id="EF8DABA4A67F0660FE53FA14FD9BE1E9" author="Dalla Torre K. W. von" box="[424,633,1469,1496]" pageId="74" pageNumber="75" pagination="135 - 290" refId="ref43620" refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany." type="book chapter" year="1912">Dalla Torre, 1912</bibRefCitation>
)
</taxonomicName>
<determinerName id="7E7F3A91A67F0660FD72FA14FCD7E1E9" box="[649,821,1469,1496]" pageId="74" pageNumber="75">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A67F0660FCA1FA14FC3BE1E6" box="[858,985,1469,1495]" pageId="74" pageNumber="75">Coca-Abia</determinerName>
det. 2019”;
<collectionCode id="ED0D4E90A67F0660FB90FA17FB20E1E9" box="[1131,1218,1470,1496]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="74" pageNumber="75">MFNB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA3D655A67F0660FF11FA56FDECE228" blockId="74.[189,1399,1534,1632]" box="[234,526,1534,1561]" pageId="74" pageNumber="75">
<emphasis id="B9680A47A67F0660FF11FA56FDECE228" bold="true" box="[234,526,1534,1561]" pageId="74" pageNumber="75">
<typeStatus id="54A768F7A67F0660FF11FA56FEBEE228" box="[234,348,1535,1561]" pageId="74" pageNumber="75" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C1CADD6A67F0660FE7FFA56FDECE228" authorityName="Keith" authorityYear="2010" box="[388,526,1534,1561]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="B9680A47A67F0660FE7FFA56FDECE228" bold="true" box="[388,526,1534,1561]" italics="true" pageId="74" pageNumber="75">M. lassallei</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA3D655A67F0660FF46F98BFD64E251" blockId="74.[189,1399,1534,1632]" pageId="74" pageNumber="75">
<materialsCitation id="3B74DC08A67F0660FF46F98BFD64E251" ID-GBIF-Occurrence="3114121351" collectingDate="2004-05" collectionCode="DKC" collectorName="Lassalle &amp; Vietnam" country="Vietnam" determinerName="D. Keith" location="VIETNAM" pageId="74" pageNumber="75" specimenCount="1" specimenCount-male="1" stateProvince="Yen Bai" typeStatus="holotype">
<collectingCountry id="F30B96C5A67F0660FF46F98BFEA6E20D" box="[189,324,1570,1596]" name="Vietnam" pageId="74" pageNumber="75">VIETNAM</collectingCountry>
•
<specimenCount id="9D1A1DDCA67F0660FEA4F98AFE9AE20C" box="[351,376,1571,1597]" pageId="74" pageNumber="75" type="male">♂</specimenCount>
; “
<locationDeviation id="9A11CF77A67F0660FE6BF98BFD35E20D" box="[400,727,1570,1597]" pageId="74" pageNumber="75">Nghia Lo ouest de Yen Bai</locationDeviation>
” [
<locationDeviation id="9A11CF77A67F0660FD0AF98BFBD5E20C" box="[753,1079,1570,1597]" pageId="74" pageNumber="75">Nghia Lo, west of Yen Bai</locationDeviation>
]; “
<date id="FFA2F095A67F0660FBADF98BFB37E20D" box="[1110,1237,1570,1596]" pageId="74" pageNumber="75" value="2004-05">
<collectingDate id="EFE6097DA67F0660FBADF98BFB37E20D" box="[1110,1237,1570,1596]" pageId="74" pageNumber="75" value="2004-05">May 2004</collectingDate>
</date>
;
<collectorName id="26E9B383A67F0660FB18F98BFAA1E20D" box="[1251,1347,1570,1596]" pageId="74" pageNumber="75">Lassalle</collectorName>
leg.
<collectorName id="26E9B383A67F0660FF46F9ECFEC5E26E" box="[189,295,1605,1631]" pageId="74" pageNumber="75">Vietnam</collectorName>
”; “
<determinerName id="7E7F3A91A67F0660FEB2F9EFFE4DE26E" box="[329,431,1605,1632]" pageId="74" pageNumber="75">D. Keith</determinerName>
det. 2009”;
<collectionCode id="ED0D4E90A67F0660FDBBF9EFFD60E251" box="[576,642,1606,1632]" pageId="74" pageNumber="75">DKC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA3D655A67F0660FF11F92EFE44E290" blockId="74.[189,826,1671,1733]" box="[234,422,1671,1697]" pageId="74" pageNumber="75">
<emphasis id="B9680A47A67F0660FF11F92EFE44E290" bold="true" box="[234,422,1671,1697]" pageId="74" pageNumber="75">Other material</emphasis>
</paragraph>
<paragraph id="8BA3D655A67F0660FF46F903FCD8E2F4" blockId="74.[189,826,1671,1733]" box="[189,826,1706,1733]" pageId="74" pageNumber="75">
<materialsCitation id="3B74DC08A67F0660FF46F903FCD4E2F4" ID-GBIF-Occurrence="3114121373" box="[189,822,1706,1733]" collectionCode="MFNB" collectorName="Pipitz &amp; Coll. Brenske" country="China" location="CHINA" pageId="74" pageNumber="75" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCountry id="F30B96C5A67F0660FF46F903FEFAE2F5" box="[189,280,1706,1732]" name="China" pageId="74" pageNumber="75">CHINA</collectingCountry>
•
<specimenCount id="9D1A1DDCA67F0660FECAF903FEBDE2F4" box="[305,351,1706,1733]" pageId="74" pageNumber="75" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1A1DDCA67F0660FE97F903FE7BE2F4" box="[364,409,1706,1733]" pageId="74" pageNumber="75" type="female">1 ♀</specimenCount>
;
<collectorName id="26E9B383A67F0660FE5CF903FE09E2F5" box="[423,491,1706,1732]" pageId="74" pageNumber="75">Pipitz</collectorName>
leg.;
<collectorName id="26E9B383A67F0660FDD5F903FD31E2F5" box="[558,723,1706,1732]" pageId="74" pageNumber="75">Coll. Brenske</collectorName>
;
<collectionCode id="ED0D4E90A67F0660FD24F902FCD4E2F4" box="[735,822,1707,1733]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="74" pageNumber="75">MFNB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA67F0666FF46F945FD4FE5C6" lastPageId="76" lastPageNumber="77" pageId="74" pageNumber="75" type="discussion">
<paragraph id="8BA3D655A67F0660FF46F945FECDE337" blockId="74.[189,303,1772,1798]" box="[189,303,1772,1798]" pageId="74" pageNumber="75">
<heading id="D0EB6139A67F0660FF46F945FECDE337" bold="true" box="[189,303,1772,1798]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="B9680A47A67F0660FF46F945FECDE337" bold="true" box="[189,303,1772,1798]" pageId="74" pageNumber="75">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A67F0666FF46F8B3FD4FE5C6" blockId="74.[189,1399,1818,2021]" lastBlockId="76.[188,1399,265,503]" lastPageId="76" lastPageNumber="77" pageId="74" pageNumber="75">
Nonfried (1892) cited
<taxonomicName id="4C1CADD6A67F0660FE2EF8B3FC89E305" authority="Reitter, 1892" authorityName="Reitter" authorityYear="1892" box="[469,875,1818,1844]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FE2EF8B3FD22E305" box="[469,704,1818,1844]" italics="true" pageId="74" pageNumber="75">Holotrichia scutata</emphasis>
Reitter, 1892
</taxonomicName>
collected in Nienghali (
<collectingRegion id="49D818B7A67F0660FB67F8B3FA95E305" box="[1180,1399,1818,1844]" country="China" name="Guangxi" pageId="74" pageNumber="75">Guangxi province</collectingRegion>
of
<collectingCountry id="F30B96C5A67F0660FF1AF894FECEE369" box="[225,300,1853,1880]" name="China" pageId="74" pageNumber="75">China</collectingCountry>
), mentioning a publication, which, supposedly, contained the original species description. Unfortunately, we have not found this paper.
<bibRefCitation id="EF8DABA4A67F0660FD1EF8C9FC6FE34A" author="Reitter E." box="[741,909,1888,1915]" pageId="74" pageNumber="75" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter (1902)</bibRefCitation>
described this species from
<collectingCountry id="F30B96C5A67F0660FB11F8C8FA94E34A" box="[1258,1398,1889,1915]" name="Hong Kong" pageId="74" pageNumber="75">Hong Kong</collectingCountry>
without specifying the origin or number of specimens described. Later,
<bibRefCitation id="EF8DABA4A67F0660FBD6F82DFB36E3AF" author="Moser J." box="[1069,1236,1924,1950]" pageId="74" pageNumber="75" pagination="468 - 488" refId="ref45295" refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488." type="journal article" year="1909">Moser (1909)</bibRefCitation>
, ignoring the existence of Brenske’s species and Reitter’s description, published another taxon named
<taxonomicName id="4C1CADD6A67F0660FB15F80EFE45E3D5" authority="Moser, 1909" authorityName="Moser" authorityYear="1909" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67F0660FB15F80EFEF0E3D5" italics="true" pageId="74" pageNumber="75">Holotrichia scutata</emphasis>
<bibRefCitation id="EF8DABA4A67F0660FEEEF862FE45E3D5" author="Moser J." box="[277,423,1994,2021]" pageId="74" pageNumber="75" pagination="468 - 488" refId="ref45295" refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488." type="journal article" year="1909">Moser, 1909</bibRefCitation>
</taxonomicName>
,also from
<collectingCountry id="F30B96C5A67F0660FDDBF862FD48E3D4" box="[544,682,1995,2021]" name="Hong Kong" pageId="74" pageNumber="75">Hong Kong</collectingCountry>
.Soon after,
<bibRefCitation id="EF8DABA4A67F0660FCCAF863FBE6E3D5" author="Dalla Torre K. W. von" box="[817,1028,1994,2021]" pageId="74" pageNumber="75" pagination="135 - 290" refId="ref43620" refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany." type="book chapter" year="1912">Dalla Torre (1912)</bibRefCitation>
realized the primary homonymy and proposed the name
<taxonomicName id="4C1CADD6A67E0661FE29FEA0FD2CE512" authorityName="Dalla Torre" authorityYear="1912" box="[466,718,265,291]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="scutulata">
<emphasis id="B9680A47A67E0661FE29FEA0FD2CE512" box="[466,718,265,291]" italics="true" pageId="75" pageNumber="76">Holotrichia scutulata</emphasis>
</taxonomicName>
for
<taxonomicName id="4C1CADD6A67E0661FD05FEA3FC94E512" authorityName="Moser" authorityYear="1909" box="[766,886,266,291]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67E0661FD05FEA3FC94E512" box="[766,886,266,291]" italics="true" pageId="75" pageNumber="76">H. scutata</emphasis>
</taxonomicName>
of
<bibRefCitation id="EF8DABA4A67E0661FC66FEA3FBA3E512" author="Moser J." box="[925,1089,265,292]" pageId="75" pageNumber="76" pagination="468 - 488" refId="ref45295" refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488." type="journal article" year="1909">Moser (1909)</bibRefCitation>
. We have studied
<specimenCount id="9D1A1DDCA67E0661FAE8FEA3FA95E515" box="[1299,1399,266,292]" pageId="75" pageNumber="76" type="generic">one type</specimenCount>
specimen of
<taxonomicName id="4C1CADD6A67E0661FEB4FE84FD86E577" authority="Reitter, 1902" authorityName="Reitter" authorityYear="1902" box="[335,612,300,326]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67E0661FEB4FE84FE24E577" box="[335,454,301,326]" italics="true" pageId="75" pageNumber="76">H. scutata</emphasis>
<bibRefCitation id="EF8DABA4A67E0661FE36FE85FD86E577" author="Reitter E." box="[461,612,300,326]" pageId="75" pageNumber="76" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
</taxonomicName>
(MFNB), which shows external features that characterize the genus
<taxonomicName id="4C1CADD6A67E0661FF47FEE6FEC6E558" authorityName="Pseudosinensis" authorityYear="2021" baseAuthorityName="Matsumoto" baseAuthorityYear="2011" box="[188,292,335,361]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A67E0661FF47FEE6FEC6E558" box="[188,292,335,361]" italics="true" pageId="75" pageNumber="76">Miridiba</emphasis>
</taxonomicName>
. Hence, we transfer
<taxonomicName id="4C1CADD6A67E0661FDE2FEF9FCD3E55B" authority="Reitter, 1902" authorityName="Reitter" authorityYear="1902" box="[537,817,335,362]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67E0661FDE2FEF9FD70E558" box="[537,658,336,361]" italics="true" pageId="75" pageNumber="76">H. scutata</emphasis>
<bibRefCitation id="EF8DABA4A67E0661FD62FEE6FCD3E55B" author="Reitter E." box="[665,817,335,362]" pageId="75" pageNumber="76" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
</taxonomicName>
to
<taxonomicName id="4C1CADD6A67E0661FCACFEE6FC5DE558" authorityName="Pseudosinensis" authorityYear="2021" baseAuthorityName="Matsumoto" baseAuthorityYear="2011" box="[855,959,335,361]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A67E0661FCACFEE6FC5DE558" box="[855,959,335,361]" italics="true" pageId="75" pageNumber="76">Miridiba</emphasis>
</taxonomicName>
and establish the genital morphotype VIII “
<emphasis id="B9680A47A67E0661FEFDFEDAFEBCE5BC" box="[262,350,371,397]" italics="true" pageId="75" pageNumber="76">Scutata</emphasis>
”. Besides, we have studied a type specimen of
<taxonomicName id="4C1CADD6A67E0661FC64FEDBFA95E5BC" authority="Dalla Torre, 1912" authorityName="Dalla Torre" authorityYear="1912" box="[927,1399,370,397]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="scutulata">
<emphasis id="B9680A47A67E0661FC64FEDBFB7FE5BD" box="[927,1181,370,396]" italics="true" pageId="75" pageNumber="76">Holotrichia scutulata</emphasis>
<bibRefCitation id="EF8DABA4A67E0661FB5FFEDBFA95E5BC" author="Dalla Torre K. W. von" box="[1188,1399,370,397]" pageId="75" pageNumber="76" pagination="135 - 290" refId="ref43620" refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany." type="book chapter" year="1912">Dalla Torre, 1912</bibRefCitation>
</taxonomicName>
(MFNB), verifying the similarities of external morphology and male genitalia (parameres shape, sclerotized structures and sensillae in the endophallus) between both taxa, which confirm the synonymy of
<taxonomicName id="4C1CADD6A6790666FF26FEA3FE8CE512" authorityName="Dalla Torre" authorityYear="1912" box="[221,366,265,291]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="scutulata">
<emphasis id="B9680A47A6790666FF26FEA3FE8CE512" box="[221,366,265,291]" italics="true" pageId="76" pageNumber="77">H. scutulata</emphasis>
</taxonomicName>
with
<taxonomicName id="4C1CADD6A6790666FE49FEA3FDC8E512" authorityName="Reitter" authorityYear="1902" box="[434,554,266,291]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A6790666FE49FEA3FDC8E512" box="[434,554,266,291]" italics="true" pageId="76" pageNumber="77">H. scutata</emphasis>
</taxonomicName>
. On the other hand, Keith (2010) described
<taxonomicName id="4C1CADD6A6790666FBCEFEA0FAE5E512" authorityName="Keith" authorityYear="2010" box="[1077,1287,265,291]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="B9680A47A6790666FBCEFEA0FAE5E512" box="[1077,1287,265,291]" italics="true" pageId="76" pageNumber="77">Miridiba lassallei</emphasis>
</taxonomicName>
based on a male specimen. According to the original description and illustrations (Keith 2010: figs 9–11), the characteristic shape of aedeagus, its asymmetry, the fusion and twist of dorsal branches, and fusion of ventral ones, distinguish it from other known
<taxonomicName id="4C1CADD6A6790666FD32FEDBFC77E5BD" box="[713,917,370,396]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9680A47A6790666FD32FEDBFCD3E5BD" box="[713,817,370,396]" italics="true" pageId="76" pageNumber="77">Miridiba</emphasis>
species.
</taxonomicName>
Nevertheless, after studying the
<typeStatus id="54A768F7A6790666FAEBFEDBFA95E5BD" box="[1296,1399,370,396]" pageId="76" pageNumber="77" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1CADD6A6790666FF25FE3EFE27E581" authority="(DKC)" baseAuthorityName="DKC" box="[222,453,406,432]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="B9680A47A6790666FF25FE3EFE8BE581" box="[222,361,406,432]" italics="true" pageId="76" pageNumber="77">M. lassallei</emphasis>
(DKC)
</taxonomicName>
, we have noted that the features of the external morphology and male genitalia that characterize
<taxonomicName id="4C1CADD6A6790666FE79FE13FD4CE5E2" authority="(Reitter, 1902)" baseAuthorityName="Reitter" baseAuthorityYear="1902" box="[386,686,441,467]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A6790666FE79FE13FE1FE5E2" box="[386,509,442,467]" italics="true" pageId="76" pageNumber="77">M. scutata</emphasis>
(
<bibRefCitation id="EF8DABA4A6790666FDF7FE10FD44E5E2" author="Reitter E." box="[524,678,441,467]" pageId="76" pageNumber="77" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
)
</taxonomicName>
are also present in
<taxonomicName id="4C1CADD6A6790666FC77FE13FBF6E5E2" box="[908,1044,441,467]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="B9680A47A6790666FC77FE13FBF6E5E2" box="[908,1044,441,467]" italics="true" pageId="76" pageNumber="77">M. lassallei</emphasis>
</taxonomicName>
. Therefore, we established the synonymy of
<taxonomicName id="4C1CADD6A6790666FEA5FE74FE05E5C7" box="[350,487,476,502]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="B9680A47A6790666FEA5FE74FE05E5C7" box="[350,487,476,502]" italics="true" pageId="76" pageNumber="77">M. lassallei</emphasis>
</taxonomicName>
with
<taxonomicName id="4C1CADD6A6790666FDD1FE74FD44E5C7" baseAuthorityName="Reitter" baseAuthorityYear="1902" box="[554,678,477,502]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A6790666FDD1FE74FD44E5C7" box="[554,678,477,502]" italics="true" pageId="76" pageNumber="77">M. scutata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF6386DDA67E0661FF46F8F6FE12E3D2" ID-DOI="http://doi.org/10.5281/zenodo.4770413" ID-Zenodo-Dep="4770413" httpUri="https://zenodo.org/record/4770413/files/figure.png" pageId="75" pageNumber="76" startId="75.[189,241,1887,1913]" targetBox="[277,1301,581,1832]" targetPageId="75">
<paragraph id="8BA3D655A67E0661FF46F8F6FE12E3D2" blockId="75.[189,1399,1887,2020]" pageId="75" pageNumber="76">
<emphasis id="B9680A47A67E0661FF46F8F6FE91E348" bold="true" box="[189,371,1887,1913]" pageId="75" pageNumber="76">Figs 94–97. 94</emphasis>
.
<taxonomicName id="4C1CADD6A67E0661FE7EF8F6FCE5E348" authority="(Reitter, 1902)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Reitter" baseAuthorityYear="1902" box="[389,775,1887,1913]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A67E0661FE7EF8F6FDAEE348" box="[389,588,1887,1913]" italics="true" pageId="75" pageNumber="76">Miridiba scutata</emphasis>
(
<bibRefCitation id="EF8DABA4A67E0661FDA6F8F6FD1CE348" author="Reitter E." box="[605,766,1887,1913]" pageId="75" pageNumber="76" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
)
</taxonomicName>
, lectotype, ♂ (MFNB). Habitus in drosal view.
<emphasis id="B9680A47A67E0661FAB1F8F6FF38E3AD" bold="true" pageId="75" pageNumber="76">95– 97</emphasis>
.
<taxonomicName id="4C1CADD6A67E0661FF10F82AFDBFE3AC" authority="Keith, 2010" authorityName="Keith" authorityYear="2010" box="[235,605,1922,1949]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="schoolmeesteri">
<emphasis id="B9680A47A67E0661FF10F82AFE26E3AD" box="[235,452,1922,1948]" italics="true" pageId="75" pageNumber="76">M. schoolmeesteri</emphasis>
Keith, 2010
</taxonomicName>
, holotype, ♂ (DKC).
<emphasis id="B9680A47A67E0661FC98F82BFC62E3AD" bold="true" box="[867,896,1922,1948]" pageId="75" pageNumber="76">95</emphasis>
. Habitus in dorsal view.
<emphasis id="B9680A47A67E0661FB4BF82BFB29E3AD" bold="true" box="[1200,1227,1922,1948]" pageId="75" pageNumber="76">96</emphasis>
. Parameres in dorsal (A) and lateral (B) views.
<emphasis id="B9680A47A67E0661FDAEF80FFD90E3F1" bold="true" box="[597,626,1958,1984]" pageId="75" pageNumber="76">97</emphasis>
. Endophallus and temones in lateral view. Scale bars: 94–95 = 5.0 mm; 96–97 = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA6790666FF46FDB4FCF3E656" box="[189,785,541,615]" pageId="76" pageNumber="77" type="distribution">
<paragraph id="8BA3D655A6790666FF46FDB4FEB4E606" blockId="76.[189,342,541,567]" box="[189,342,541,567]" pageId="76" pageNumber="77">
<heading id="D0EB6139A6790666FF46FDB4FEB4E606" bold="true" box="[189,342,541,567]" fontSize="11" level="3" pageId="76" pageNumber="77" reason="3">
<emphasis id="B9680A47A6790666FF46FDB4FEB4E606" bold="true" box="[189,342,541,567]" pageId="76" pageNumber="77">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6790666FF46FDE5FCF3E656" blockId="76.[189,785,588,615]" box="[189,785,588,615]" pageId="76" pageNumber="77">
<collectingCountry id="F30B96C5A6790666FF46FDE5FEE1E657" box="[189,259,588,614]" name="China" pageId="76" pageNumber="77">China</collectingCountry>
(
<collectingRegion id="49D818B7A6790666FEE8FDE5FE9EE657" box="[275,380,588,614]" country="China" name="Guangxi" pageId="76" pageNumber="77">Guangxi</collectingRegion>
,
<collectingCountry id="F30B96C5A6790666FE73FDE5FDEBE656" box="[392,521,588,615]" name="Hong Kong" pageId="76" pageNumber="77">Hongkong</collectingCountry>
),
<collectingCountry id="F30B96C5A6790666FDE7FDE5FD9DE657" box="[540,639,588,614]" name="Vietnam" pageId="76" pageNumber="77">Vietnam</collectingCountry>
(Nghia Lo).
</paragraph>
</subSubSection>
</treatment>
</document>
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