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<document id="ECC48CFFA6C4AE148C16A4502AFD00A3" ID-CLB-Dataset="27590" ID-DOI="10.11646/zootaxa.4551.4.3" ID-GBIF-Dataset="ddc0d912-7a93-428e-a987-8eb155da7d02" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623047" ID-ZooBank="886BB4EA-3A68-47F3-BBBC-1B8D9603D13C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554270427687" checkinUser="plazi" docAuthor="Buchsbaum, Ulf &amp; Grehan, John R." docDate="2019" docId="03B5224FC137FFEDFF41FF65FC93F91A" docLanguage="en" docName="zootaxa.4551.4.3.pdf" docOrigin="Zootaxa 4551 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Endoclita meifenga Buchsbaum &amp; Grehan 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="436" masterDocId="FF8C5A37C135FFE9FFD6FFB8FFE8FFA5" masterDocTitle="New species of Endoclita (Lepidoptera: Hepialidae) and revived species status of E. kosemponis from Taiwan" masterLastPageNumber="444" masterPageNumber="432" pageNumber="434" updateTime="1698716150949" updateUser="ExternalLinkService">
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<mods:title id="51A39B9A8E57DF908412C3A17B6189FD">New species of Endoclita (Lepidoptera: Hepialidae) and revived species status of E. kosemponis from Taiwan</mods:title>
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<mods:namePart id="7F5B1AEC5AEF1C7D91A2926747C032F8">Buchsbaum, Ulf</mods:namePart>
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<mods:namePart id="BD4D38602A4484F1393398783B7CAA90">Grehan, John R.</mods:namePart>
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<mods:title id="114FE934515B080642D31AB65EE2208B">Zootaxa</mods:title>
</mods:titleInfo>
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<mods:date id="851707B76AC27A24B782D3A8BBB4FDE2">2019</mods:date>
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<mods:number id="50985A1E70858153AFB4A2D330C28806">2019-02-04</mods:number>
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<mods:identifier id="EBBD78A83AD2EDC345377721C04BFC86" type="DOI">10.11646/zootaxa.4551.4.3</mods:identifier>
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<treatment id="03B5224FC137FFEDFF41FF65FC93F91A" ID-DOI="http://doi.org/10.5281/zenodo.5943871" ID-GBIF-Taxon="154534049" ID-Zenodo-Dep="5943871" LSID="urn:lsid:plazi:treatment:03B5224FC137FFEDFF41FF65FC93F91A" httpUri="http://treatment.plazi.org/id/03B5224FC137FFEDFF41FF65FC93F91A" lastPageId="4" lastPageNumber="436" pageId="2" pageNumber="434">
<subSubSection id="C306C0D2C137FFEBFF41FF65FD07FF52" box="[151,751,221,247]" pageId="2" pageNumber="434" type="nomenclature">
<paragraph id="8BA39359C137FFEBFF41FF65FD07FF52" blockId="2.[151,751,221,280]" box="[151,751,221,247]" pageId="2" pageNumber="434">
<heading id="D0EB2435C137FFEBFF41FF65FD07FF52" bold="true" box="[151,751,221,247]" fontSize="11" level="1" pageId="2" pageNumber="434" reason="1">
<emphasis id="B9684F4BC137FFEBFF41FF65FD07FF52" bold="true" box="[151,751,221,247]" pageId="2" pageNumber="434">
<taxonomicName id="4C1CE8DAC137FFEBFF41FF65FD4DFF52" authority="Buchsbaum &amp; Grehan" authorityName="Buchsbaum &amp; Grehan" authorityYear="2019" box="[151,677,221,247]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="meifenga" status="sp. nov.">
<emphasis id="B9684F4BC137FFEBFF41FF65FE6BFF52" bold="true" box="[151,387,221,247]" italics="true" pageId="2" pageNumber="434">Endoclita meifenga</emphasis>
Buchsbaum &amp; Grehan
</taxonomicName>
<taxonomicNameLabel id="A25BF230C137FFEBFD7BFF65FD07FF52" box="[685,751,221,247]" pageId="2" pageNumber="434" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C306C0D2C137FFEBFF41FF47FDDBFEBD" box="[151,563,255,280]" pageId="2" pageNumber="434" type="description">
<paragraph id="8BA39359C137FFEBFF41FF47FDDBFEBD" blockId="2.[151,751,221,280]" box="[151,563,255,280]" pageId="2" pageNumber="434">
<figureCitation id="13278FDCC137FFEBFF41FF47FF37FEBD" box="[151,223,255,280]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1944,1967]" captionTargetBox="[372,1214,1068,1901]" captionTargetId="figure@2.[372,1214,1068,1901]" captionTargetPageId="2" captionText="FIGURES 1–2. Voucher specimens: Endoclita meifenga sp. n., male HT (NMNS). 1a, dorsal; 1b, ventral; 1c, habitus at light sheet (reoriented from original vertical positon with head uppermost). 2, E. kosemponis types (ZMHB). 2a, dorsal male LT; 2b, ventral; 2c dorsal female. Photos by Ulf Buchsbaum." httpUri="https://zenodo.org/record/2623049/files/figure.png" pageId="2" pageNumber="434">Figs. 1</figureCitation>
a–c, 4a, 5a, 6a, 7a, 8a, 9a, 12a
</paragraph>
</subSubSection>
<subSubSection id="C306C0D2C137FFEBFF41FEFFFAFBFEC5" box="[151,1299,327,352]" pageId="2" pageNumber="434" type="etymology">
<paragraph id="8BA39359C137FFEBFF41FEFFFAFBFEC5" blockId="2.[151,1437,327,1000]" box="[151,1299,327,352]" pageId="2" pageNumber="434">
<emphasis id="B9684F4BC137FFEBFF41FEFFFEFEFEC5" bold="true" box="[151,278,327,352]" pageId="2" pageNumber="434">Etymology</emphasis>
. Named for the Meifeng
<collectingRegion id="49D85DBBC137FFEBFDE5FEFFFD73FEC5" box="[563,667,327,352]" country="United Kingdom" name="Highland" pageId="2" pageNumber="434">Highland</collectingRegion>
Experimental Station where the
<typeStatus id="54A72DFBC137FFEBFBDCFEFFFB84FEC5" box="[1034,1132,327,352]" pageId="2" pageNumber="434" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="C306C0D2C137FFEBFF11FED4FE6DFE69" pageId="2" pageNumber="434" type="materials_examined">
<paragraph id="8BA39359C137FFEBFF11FED4FE6DFE69" blockId="2.[151,1437,327,1000]" pageId="2" pageNumber="434">
<materialsCitation id="3B749904C137FFEBFF11FED4FE6DFE69" ID-GBIF-Occurrence="2234226090" collectingDate="2011-05-11" collectionCode="NMNS" collectorName="Chen &amp; U. Buchsbaum. Deposited" country="Taiwan" county="Central" elevation="2100" location="Meifeng" municipality="Puli" pageId="2" pageNumber="434" specimenCount="1" specimenCount-male="1" stateProvince="Nantou" typeStatus="holotype">
<emphasis id="B9684F4BC137FFEBFF11FED4FEDCFE20" bold="true" box="[199,308,364,389]" pageId="2" pageNumber="434">
<typeStatus id="54A72DFBC137FFEBFF11FED4FEDCFE20" box="[199,308,364,389]" pageId="2" pageNumber="434" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D1A58D0C137FFEBFEEAFED5FEB9FE21" box="[316,337,365,388]" pageId="2" pageNumber="434" type="male">♂</specimenCount>
with the following labels (separated by forward slashes):
<collectingCounty id="62C2EBD5C137FFEBFC34FED4FBDDFE20" box="[994,1077,364,389]" pageId="2" pageNumber="434">Central</collectingCounty>
<collectingCountry id="F30BD3C9C137FFEBFBEAFED4FB7AFE20" box="[1084,1170,364,389]" name="Taiwan" pageId="2" pageNumber="434">Taiwan</collectingCountry>
; near
<collectingMunicipality id="6BC70923C137FFEBFB00FED4FAEEFE20" box="[1238,1286,364,389]" pageId="2" pageNumber="434">Puli</collectingMunicipality>
,
<collectingRegion id="49D85DBBC137FFEBFAC4FED5FA8CFE21" box="[1298,1380,365,388]" country="Taiwan" name="Nantou" pageId="2" pageNumber="434">Nantou</collectingRegion>
Co.;
<location id="8EC3C582C137FFEBFF41FE37FF11FE0D" LSID="urn:lsid:plazi:treatment:03B5224FC137FFEDFF41FF65FC93F91A:8EC3C582C137FFEBFF41FE37FF11FE0D" box="[151,249,399,424]" country="Taiwan" county="Central" municipality="Puli" name="Meifeng" pageId="2" pageNumber="434" stateProvince="Nantou">Meifeng</location>
, ca.
<quantity id="4CE43EBCC137FFEBFEE7FE28FE62FE02" box="[305,394,400,424]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="434" unit="m" value="2100.0">
<elevation id="0031746AC137FFEBFEE7FE28FE62FE02" box="[305,394,400,424]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="434" unit="m" value="2100.0">2100 m</elevation>
</quantity>
NN; 24°05’19 N / 121°10’26 E; LF,
<date id="FFA2B599C137FFEBFC90FE28FC0BFE0D" box="[838,995,400,424]" pageId="2" pageNumber="434" value="2011-05-11">
<collectingDate id="EFE64C71C137FFEBFC90FE28FC0BFE0D" box="[838,995,400,424]" pageId="2" pageNumber="434" value="2011-05-11">11 May 2011</collectingDate>
</date>
; leg. Mei-Yu
<collectorName id="26E9F68FC137FFEBFB50FE37FB29FE0D" box="[1158,1217,399,424]" pageId="2" pageNumber="434">Chen</collectorName>
&amp;
<collectorName id="26E9F68FC137FFEBFB3DFE28FEE1FE68" pageId="2" pageNumber="434">U. Buchsbaum. Deposited</collectorName>
in
<collectionCode id="ED0D0B9CC137FFEBFEFAFE0CFE69FE69" box="[300,385,436,460]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="2" pageNumber="434">NMNS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C306C0D2C137FFEBFF11FE6FFB0AFC90" pageId="2" pageNumber="434" type="diagnosis">
<paragraph id="8BA39359C137FFEBFF11FE6FFB0AFC90" blockId="2.[151,1437,327,1000]" pageId="2" pageNumber="434">
<emphasis id="B9684F4BC137FFEBFF11FE6FFEA9FE55" bold="true" box="[199,321,471,496]" pageId="2" pageNumber="434">Diagnosis.</emphasis>
A medium sized
<taxonomicName id="4C1CE8DAC137FFEBFDDDFE6FFD90FE55" box="[523,632,471,496]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B9684F4BC137FFEBFDDDFE6FFD90FE55" box="[523,632,471,496]" italics="true" pageId="2" pageNumber="434">Endoclita</emphasis>
</taxonomicName>
species with a darker shaded triangular region over much of the discal area and a trapezoid ‘recess’ in the central anterior discal cell. These features are also present in
<taxonomicName id="4C1CE8DAC137FFEBFB23FE45FED5FD9D" authority="(Moore, 1879)" authorityName="Moore" authorityYear="1879" baseAuthorityName="Moore" baseAuthorityYear="1879" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="chalybeatus">
<emphasis id="B9684F4BC137FFEBFB23FE45FA74FDB0" box="[1269,1436,508,533]" italics="true" pageId="2" pageNumber="434">E. chalybeatus</emphasis>
(
<bibRefCitation id="EF8DEEA8C137FFEBFF48FD98FEDDFD9D" author="Moore, F." box="[158,309,544,568]" pageId="2" pageNumber="434" refId="ref6375" refString="Moore, F. (1879) Descriptions of new genera and species of Asiatic Lepidoptera Heterocera. Proceedings of the Zoological Society of London, 1879, 387 ‾ 417. https: // doi. org / 10.1111 / j. 1096 - 3642.1879. tb 02671. x" type="book" year="1879">Moore, 1879</bibRefCitation>
)
</taxonomicName>
(northeastern
<collectingCountry id="F30BD3C9C137FFEBFE32FDA7FDCAFD9D" box="[484,546,543,568]" name="India" pageId="2" pageNumber="434">India</collectingCountry>
),
<taxonomicName id="4C1CE8DAC137FFEBFDEEFD99FD48FD9D" baseAuthorityName="Poujade" baseAuthorityYear="1886" box="[568,672,543,568]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="B9684F4BC137FFEBFDEEFD99FD48FD9D" box="[568,672,543,568]" italics="true" pageId="2" pageNumber="434">E. davidi</emphasis>
</taxonomicName>
(
<collectingCountry id="F30BD3C9C137FFEBFD67FDA7FCEDFD9D" box="[689,773,543,568]" name="Taiwan" pageId="2" pageNumber="434">Taiwan</collectingCountry>
and western
<collectingCountry id="F30BD3C9C137FFEBFC74FDA7FC01FD9D" box="[930,1001,543,568]" name="China" pageId="2" pageNumber="434">China</collectingCountry>
),
<taxonomicName id="4C1CE8DAC137FFEBFC29FD99FAFDFD9D" authority="(Taiwan)" authorityName="Buchsbaum &amp; Grehan" authorityYear="2019" baseAuthorityName="Taiwan" baseAuthorityYear="1916" box="[1023,1301,543,568]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="kosemponis">
<emphasis id="B9684F4BC137FFEBFC29FD99FB4CFD9D" box="[1023,1188,543,568]" italics="true" pageId="2" pageNumber="434">E. kosemponis</emphasis>
(
<collectingRegion id="49D85DBBC137FFEBFB63FDA7FAFDFD9D" box="[1205,1301,543,568]" country="China" name="Taiwan)" pageId="2" pageNumber="434">Taiwan)</collectingRegion>
</taxonomicName>
,
<taxonomicName id="4C1CE8DAC137FFEBFAF5FD99FEA0FDF9" authority="(Tindale, 1942)" authorityName="Tindale" authorityYear="1942" baseAuthorityName="Tindale" baseAuthorityYear="1942" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="B9684F4BC137FFEBFAF5FD99FA74FD9D" box="[1315,1436,545,568]" italics="true" pageId="2" pageNumber="434">E. magnus</emphasis>
(
<bibRefCitation id="EF8DEEA8C137FFEBFF48FDFCFEA8FDF9" author="Tindale, N. B." box="[158,320,580,605]" pageId="2" pageNumber="434" pagination="151 - 168" refId="ref6756" refString="Tindale, N. B. (1942) Revision of the ghost moths (Lepidoptera Homoneura, family Hepialidae). Part V. Records of the South Australian Museum, 7, 151 - 168." type="journal article" year="1942">Tindale, 1942</bibRefCitation>
)
</taxonomicName>
(southwestern
<collectingCountry id="F30BD3C9C137FFEBFE2DFDFCFDD1FDF8" box="[507,569,580,605]" name="India" pageId="2" pageNumber="434">India</collectingCountry>
),
<taxonomicName id="4C1CE8DAC137FFEBFD86FDFDFC59FDF9" authority="(Moore, 1879)" authorityName="Moore" authorityYear="1879" baseAuthorityName="Moore" baseAuthorityYear="1879" box="[592,945,580,605]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="B9684F4BC137FFEBFD86FDFDFD17FDF8" box="[592,767,580,605]" italics="true" pageId="2" pageNumber="434">E. malabaricus</emphasis>
(
<bibRefCitation id="EF8DEEA8C137FFEBFCC4FDFCFC41FDF9" author="Moore, F." box="[786,937,580,604]" pageId="2" pageNumber="434" refId="ref6375" refString="Moore, F. (1879) Descriptions of new genera and species of Asiatic Lepidoptera Heterocera. Proceedings of the Zoological Society of London, 1879, 387 ‾ 417. https: // doi. org / 10.1111 / j. 1096 - 3642.1879. tb 02671. x" type="book" year="1879">Moore, 1879</bibRefCitation>
)
</taxonomicName>
(southwestern
<collectingCountry id="F30BD3C9C137FFEBFBB2FDFCFB4AFDF8" box="[1124,1186,580,605]" name="India" pageId="2" pageNumber="434">India</collectingCountry>
),
<taxonomicName id="4C1CE8DAC137FFEBFB6FFDFDFF30FD25" authority="(Walker, 1856)" authorityName="Walker" authorityYear="1856" baseAuthorityName="Walker" baseAuthorityYear="1856" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="signifer">
<emphasis id="B9684F4BC137FFEBFB6FFDFDFADAFDF9" box="[1209,1330,580,604]" italics="true" pageId="2" pageNumber="434">E. signifer</emphasis>
(
<bibRefCitation id="EF8DEEA8C137FFEBFA92FDFCFF38FD25" author="Walker, F." pageId="2" pageNumber="434" refId="ref6989" refString="Walker, F. (1856) List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Part VII. Lepidoptera Heterocera. Trustees of BMNH, London, 299 pp." type="book" year="1856">Walker, 1856</bibRefCitation>
)
</taxonomicName>
(northeastern
<collectingCountry id="F30BD3C9C137FFEBFEAAFDDFFE51FD25" box="[380,441,615,640]" name="India" pageId="2" pageNumber="434">India</collectingCountry>
), and
<taxonomicName id="4C1CE8DAC137FFEBFE29FDD1FD90FD25" baseAuthorityName="Moore" baseAuthorityYear="1877" box="[511,632,617,640]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B9684F4BC137FFEBFE29FDD1FD90FD25" box="[511,632,617,640]" italics="true" pageId="2" pageNumber="434">E. sinensis</emphasis>
</taxonomicName>
(
<collectingCountry id="F30BD3C9C137FFEBFD5EFDDFFD34FD25" box="[648,732,615,640]" name="Taiwan" pageId="2" pageNumber="434">Taiwan</collectingCountry>
and eastern
<collectingCountry id="F30BD3C9C137FFEBFCBAFDDFFC5BFD25" box="[876,947,615,640]" name="China" pageId="2" pageNumber="434">China</collectingCountry>
), but absent from the Taiwanese species
<taxonomicName id="4C1CE8DAC137FFEBFA52FDD1FF03FD00" authorityName="Buchsbaum &amp; Hsu" authorityYear="2018" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="atayala">
<emphasis id="B9684F4BC137FFEBFA52FDD1FF03FD00" italics="true" pageId="2" pageNumber="434">E. atayala</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1CE8DAC137FFEBFEF6FD35FE6DFD01" authorityName="Ueda" authorityYear="1987" box="[288,389,653,676]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="inouei">
<emphasis id="B9684F4BC137FFEBFEF6FD35FE6DFD01" box="[288,389,653,676]" italics="true" pageId="2" pageNumber="434">E. inouei</emphasis>
</taxonomicName>
. Within the triangular shaded discal cell species present in
<collectingCountry id="F30BD3C9C137FFEBFBCEFD34FB85FD00" box="[1048,1133,652,677]" name="Taiwan" pageId="2" pageNumber="434">Taiwan</collectingCountry>
, the FW of
<taxonomicName id="4C1CE8DAC137FFEBFB38FD35FA9CFD01" authority="Buchsbaum &amp; Grehan, 2019" authorityName="Buchsbaum &amp; Grehan" authorityYear="2019" box="[1262,1396,652,676]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="meifenga" status="sp. nov.">
<emphasis id="B9684F4BC137FFEBFB38FD35FA9CFD01" box="[1262,1396,652,676]" italics="true" pageId="2" pageNumber="434">E. meifenga</emphasis>
</taxonomicName>
<emphasis id="B9684F4BC137FFEBFAACFD34FF46FD6D" bold="true" pageId="2" pageNumber="434">
<taxonomicNameLabel id="A25BF230C137FFEBFAACFD34FF46FD6D" pageId="2" pageNumber="434" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C1CE8DAC137FFEBFE31FD09FD63FD6D" authorityName="Buchsbaum &amp; Grehan" authorityYear="2019" baseAuthorityName="Strand" baseAuthorityYear="1916" box="[487,651,687,712]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="kosemponis">
<emphasis id="B9684F4BC137FFEBFE31FD09FD63FD6D" box="[487,651,687,712]" italics="true" pageId="2" pageNumber="434">E. kosemponis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1CE8DAC137FFEBFD12FD09FCD6FD6D" baseAuthorityName="Moore" baseAuthorityYear="1877" box="[708,830,689,712]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B9684F4BC137FFEBFD12FD09FCD6FD6D" box="[708,830,689,712]" italics="true" pageId="2" pageNumber="434">E. sinensis</emphasis>
</taxonomicName>
by the presence of a prominent costal lobe. The FW pattern of
<taxonomicName id="4C1CE8DAC137FFEBFEDCFD6DFE78FD49" authority="Buchsbaum &amp; Grehan, 2019" authorityName="Buchsbaum &amp; Grehan" authorityYear="2019" box="[266,400,724,748]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="meifenga" status="sp. nov.">
<emphasis id="B9684F4BC137FFEBFEDCFD6DFE78FD49" box="[266,400,724,748]" italics="true" pageId="2" pageNumber="434">E. meifenga</emphasis>
</taxonomicName>
<emphasis id="B9684F4BC137FFEBFE40FD6CFE3EFD48" bold="true" box="[406,470,724,749]" pageId="2" pageNumber="434">
<taxonomicNameLabel id="A25BF230C137FFEBFE40FD6CFE3EFD48" box="[406,470,724,749]" pageId="2" pageNumber="434" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is very similar to
<emphasis id="B9684F4BC137FFEBFD76FD6DFCE4FD48" box="[672,780,724,749]" italics="true" pageId="2" pageNumber="434">
<taxonomicName id="4C1CE8DAC137FFEBFD76FD6DFCE0FD48" baseAuthorityName="Poujade" baseAuthorityYear="1886" box="[672,776,724,749]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="davidi">E. davidi</taxonomicName>
,
</emphasis>
but there is a pale submarginal band extending across Rs2 and Rs3 that is absent in
<emphasis id="B9684F4BC137FFEBFE1CFD41FDD4FCB5" box="[458,572,759,784]" italics="true" pageId="2" pageNumber="434">
<taxonomicName id="4C1CE8DAC137FFEBFE1CFD41FDD0FCB5" baseAuthorityName="Poujade" baseAuthorityYear="1886" box="[458,568,759,784]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="davidi">E. davidi</taxonomicName>
.
</emphasis>
The male genitalia of
<taxonomicName id="4C1CE8DAC137FFEBFC98FD41FC31FCB5" authority="Buchsbaum &amp; Grehan, 2019" authorityName="Buchsbaum &amp; Grehan" authorityYear="2019" box="[846,985,760,784]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="meifenga" status="sp. nov.">
<emphasis id="B9684F4BC137FFEBFC98FD41FC31FCB5" box="[846,985,760,784]" italics="true" pageId="2" pageNumber="434">E. meifenga</emphasis>
</taxonomicName>
<emphasis id="B9684F4BC137FFEBFC32FD4FFBC0FCB5" bold="true" box="[996,1064,759,784]" pageId="2" pageNumber="434">
<taxonomicNameLabel id="A25BF230C137FFEBFC32FD4FFBC0FCB5" box="[996,1064,759,784]" pageId="2" pageNumber="434" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C1CE8DAC137FFEBFBB1FD41FB38FCB5" baseAuthorityName="Poujade" baseAuthorityYear="1886" box="[1127,1232,759,784]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="B9684F4BC137FFEBFBB1FD41FB38FCB5" box="[1127,1232,759,784]" italics="true" pageId="2" pageNumber="434">E. davidi</emphasis>
</taxonomicName>
are also distinct, including a contrasting concave vs convex shaped posterior ventral margin of the pseudotegumen.
</paragraph>
</subSubSection>
<subSubSection id="C306C0D2C137FFEDFF11FC87FF50FA65" lastPageId="4" lastPageNumber="436" pageId="2" pageNumber="434" type="description">
<paragraph id="8BA39359C137FFEBFF11FC87FDAAFCD9" blockId="2.[151,1437,327,1000]" pageId="2" pageNumber="434">
<emphasis id="B9684F4BC137FFEBFF11FC87FEB1FCFD" bold="true" box="[199,345,831,856]" pageId="2" pageNumber="434">Description.</emphasis>
Male
<typeStatus id="54A72DFBC137FFEBFE49FC87FDE9FCFD" box="[415,513,831,856]" pageId="2" pageNumber="434" type="holotype">holotype</typeStatus>
(
<figureCitation id="13278FDCC137FFEBFDD9FC87FDA5FCFD" box="[527,589,831,856]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1944,1967]" captionTargetBox="[372,1214,1068,1901]" captionTargetId="figure@2.[372,1214,1068,1901]" captionTargetPageId="2" captionText="FIGURES 1–2. Voucher specimens: Endoclita meifenga sp. n., male HT (NMNS). 1a, dorsal; 1b, ventral; 1c, habitus at light sheet (reoriented from original vertical positon with head uppermost). 2, E. kosemponis types (ZMHB). 2a, dorsal male LT; 2b, ventral; 2c dorsal female. Photos by Ulf Buchsbaum." httpUri="https://zenodo.org/record/2623049/files/figure.png" pageId="2" pageNumber="434">Fig. 1</figureCitation>
a–c): Wingspan
<quantity id="4CE43EBCC137FFEBFCD1FCF8FCB0FCF2" box="[775,856,832,856]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="434" unit="mm" value="71.0">71 mm</quantity>
; FW length:
<quantity id="4CE43EBCC137FFEBFC31FCF8FBD0FCF2" box="[999,1080,832,856]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="434" unit="mm" value="33.0">33 mm</quantity>
, width:
<quantity id="4CE43EBCC137FFEBFB59FCF8FB37FCF2" box="[1167,1247,832,856]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="434" unit="mm" value="14.0">14 mm</quantity>
, ratio 2.4: 1; HW length:
<quantity id="4CE43EBCC137FFEBFF3AFCDCFED4FCD9" box="[236,316,868,892]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="434" unit="mm" value="29.0">29 mm</quantity>
, width: 12, ratio 2.4: 1.
</paragraph>
<paragraph id="8BA39359C137FFEBFF11FC3FFBD5FC4D" blockId="2.[151,1437,327,1000]" pageId="2" pageNumber="434">
<emphasis id="B9684F4BC137FFEBFF11FC3FFEE3FC05" box="[199,267,903,928]" italics="true" pageId="2" pageNumber="434">Head.</emphasis>
Eyes prominent. Antenna filiform, pale yellowish brown, scape cylindrical, pedicel rounded and slightly barrel shaped, flagellum with 22 segments and covered with numerous sensillae caeticae, lamellar scales absent, annuli slightly larger centrally, narrowing to apex. Interocular-antennal scales absent.
</paragraph>
<caption id="DF63C3D1C137FFEBFF41F820FD58F849" httpUri="https://zenodo.org/record/2623049/files/figure.png" pageId="2" pageNumber="434" startId="2.[151,264,1944,1967]" targetBox="[372,1214,1068,1901]" targetPageId="2">
<paragraph id="8BA39359C137FFEBFF41F820FD58F849" blockId="2.[151,1436,1944,2028]" pageId="2" pageNumber="434">
<emphasis id="B9684F4BC137FFEBFF41F820FED3F80B" bold="true" box="[151,315,1944,1967]" pageId="2" pageNumber="434">FIGURES 1–2.</emphasis>
Voucher specimens:
<taxonomicName id="4C1CE8DAC137FFEBFDC2F821FD33F80A" authority="Buchsbaum &amp; Grehan, 2019" authorityName="Buchsbaum &amp; Grehan" authorityYear="2019" box="[532,731,1945,1967]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="meifenga" status="sp. nov.">
<emphasis id="B9684F4BC137FFEBFDC2F821FD33F80A" box="[532,731,1945,1967]" italics="true" pageId="2" pageNumber="434">Endoclita meifenga</emphasis>
</taxonomicName>
<emphasis id="B9684F4BC137FFEBFD34F821FCF4F80B" bold="true" box="[738,796,1945,1966]" pageId="2" pageNumber="434">
<taxonomicNameLabel id="A25BF230C137FFEBFD34F821FCF4F80B" box="[738,796,1945,1966]" pageId="2" pageNumber="434" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male HT (NMNS). 1a, dorsal; 1b, ventral; 1c, habitus at light sheet (reoriented from original vertical positon with head uppermost). 2,
<taxonomicName id="4C1CE8DAC137FFEBFCAFF801FBE3F868" authorityName="Buchsbaum &amp; Grehan" authorityYear="2019" baseAuthorityName="Strand" baseAuthorityYear="1916" box="[889,1035,1975,1998]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="434" phylum="Arthropoda" rank="species" species="kosemponis">
<emphasis id="B9684F4BC137FFEBFCAFF801FBE3F868" box="[889,1035,1975,1998]" italics="true" pageId="2" pageNumber="434">E. kosemponis</emphasis>
</taxonomicName>
types (ZMHB). 2a, dorsal male LT; 2b, ventral; 2c dorsal female. Photos by Ulf Buchsbaum.
</paragraph>
</caption>
<caption id="DF63C3D1C136FFEAFF41FC67FCC3FBF4" httpUri="https://zenodo.org/record/2623051/files/figure.png" pageId="3" pageNumber="435" startId="3.[151,250,991,1013]" targetBox="[340,1247,193,969]" targetPageId="3">
<paragraph id="8BA39359C136FFEAFF41FC67FCC3FBF4" blockId="3.[151,1436,991,1105]" pageId="3" pageNumber="435">
<emphasis id="B9684F4BC136FFEAFF41FC67FEFBFC51" bold="true" box="[151,275,991,1013]" pageId="3" pageNumber="435">FIGURE 3.</emphasis>
Voucher specimens:
<emphasis id="B9684F4BC136FFEAFE3BFC67FDBFFC50" box="[493,599,991,1013]" italics="true" pageId="3" pageNumber="435">
<taxonomicName id="4C1CE8DAC136FFEAFE3BFC67FDBBFC50" box="[493,595,991,1013]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="435" phylum="Arthropoda" rank="genus">Endoclita</taxonomicName>
.
</emphasis>
3a, dorsal LT female
<taxonomicName id="4C1CE8DAC136FFEAFCE1FC58FC4CFC50" baseAuthorityName="Moore" baseAuthorityYear="1877" box="[823,932,992,1013]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="435" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B9684F4BC136FFEAFCE1FC58FC4CFC50" box="[823,932,992,1013]" italics="true" pageId="3" pageNumber="435">E. sinensis</emphasis>
</taxonomicName>
(incorrectly labelled as male—see Tindale 1958: 167); 3b, dorsal ST female
<taxonomicName id="4C1CE8DAC136FFEAFE66FC46FDF5FBB6" baseAuthorityName="Moore" baseAuthorityYear="1877" box="[432,541,1022,1043]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="435" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B9684F4BC136FFEAFE66FC46FDF5FBB6" box="[432,541,1022,1043]" italics="true" pageId="3" pageNumber="435">E. sinensis</emphasis>
</taxonomicName>
(NHMUK). Photos by David Lees, ©The Trustees of the Natural History Museum. 3c, dorsal male; 3d, dorsal female (NMNS). Photos by Ulf Buchsbaum; 3e, dorsal HT female
<taxonomicName id="4C1CE8DAC136FFEAFBEFFBA5FB4AFB97" authorityName="Buchsbaum &amp; Hsu" authorityYear="2018" box="[1081,1186,1052,1074]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="435" phylum="Arthropoda" rank="species" species="atayala">
<emphasis id="B9684F4BC136FFEAFBEFFBA5FB4AFB97" box="[1081,1186,1052,1074]" italics="true" pageId="3" pageNumber="435">E. atayala</emphasis>
</taxonomicName>
(from Buchsbaum et al. 2018); 3f, dorsal HT female
<taxonomicName id="4C1CE8DAC136FFEAFE6EFB84FDFBFBF4" authorityName="Ueda" authorityYear="1987" box="[440,531,1084,1105]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="435" phylum="Arthropoda" rank="species" species="inouei">
<emphasis id="B9684F4BC136FFEAFE6EFB84FDFBFBF4" box="[440,531,1084,1105]" italics="true" pageId="3" pageNumber="435">E. inouei</emphasis>
</taxonomicName>
. Photo by Kyoichiro Ueda.
</paragraph>
</caption>
<paragraph id="8BA39359C136FFEAFF11FBC0FE06F8BB" blockId="3.[151,1437,1144,2002]" pageId="3" pageNumber="435">
<emphasis id="B9684F4BC136FFEAFF11FBC0FEF0FB34" box="[199,280,1144,1169]" italics="true" pageId="3" pageNumber="435">Thorax</emphasis>
. Dorsally covered with pale to dark olive green scales, scutum III dark yellowish brown, anteriorly free of scales, lateral and ventral thorax olive green; FW costal margin with shallow costal lobe anterior to the discal white spot, margin almost straight from base to costal lobe, and distally a shallow convex curve to apex; outer margin a shallow curve merging into posterior margin; venation ‘hepialine’ (sensu
<bibRefCitation id="EF8DEEA8C136FFEAFB58FB5CFABFFB58" author="Dumbleton, L. J." box="[1166,1367,1252,1277]" pageId="3" pageNumber="435" pagination="981" refId="ref5947" refString="Dumbleton, L. J. (1966) Genitalia, classification and zoogeography of the New Zealand Hepialidae (Lepidoptera). New Zealand Journal of Science, 9, 920 ‾ 981." type="journal article" year="1966">Dumbleton 1966</bibRefCitation>
), Sc1 extending into costal lobe (
<figureCitation id="13278FDCC136FFEAFE08FAB0FDDDFA85" box="[478,565,1288,1313]" captionStart="FIGURES 4–5" captionStartId="5.[151,264,1609,1632]" captionTargetBox="[188,1398,733,1588]" captionTargetId="figure@5.[188,1398,733,1588]" captionTargetPageId="5" captionText="FIGURES 4–5. Thoracic. appendages: 4, wing venation. 4a, Endoclita meifenga sp. n (HT); 4b, E. kosemponis (LT). 5, Endoclita male legs. 5a E. meifenga sp. n; 5b, E. kosemponis LT. 5c, E. sinensis (NMNS)" httpUri="https://zenodo.org/record/2623053/files/figure.png" pageId="3" pageNumber="435">Fig. 4a</figureCitation>
). FW dorsal ground colour pale greyish brown covered with olive green markings edged in pale greenish white. Single largest olive green patch forming a triangular shape bounded anteriorly by R and posteriorly by CuA2 and extending distally just beyond the Cu-M-Rs cross veins; a greyish brown anterior trapezoid ‘recess’ patch in anterior discal cell anteriorly bounded by R (
<figureCitation id="13278FDCC136FFEAFB56FACCFB0AFA28" box="[1152,1250,1396,1421]" captionStart="FIGURE 12" captionStartId="9.[151,250,1873,1895]" captionTargetBox="[355,1236,601,1836]" captionTargetId="figure@9.[340,1247,591,1851]" captionTargetPageId="9" captionText="FIGURE 12. Male FW venation, discal shading (blue outline) and anterior recess (green outline): 12a, E. meifenga sp. n. HT); 12b, E. chalybeatus (India, Meghalaya, NHMUK); 12c, E. davidi (holotype, China, MNHN), 12d, E. kosemponis (LT); 12e, E. magnus (Shenbaganur, India, NHMUK); 12f, E. malabaricus (Ooty, India, NHMUK); 12g, E. signifer (LT, Meghalaya, India, NHMUK); 12h, E. sinensis (NMNS); ad—anterior discal cell, md—medial discal cell, pd—posterior discal cell, pdbpostdiscal band, premarginal band, smb—submarginal band, mb—marginal band." httpUri="https://zenodo.org/record/2623060/files/figure.png" pageId="3" pageNumber="435">Fig. 12a</figureCitation>
). A premarginal band of olive green extends obliquely from CuA to the costal margin near the apex, comprising elliptical shapes between the veins, and darkest between costal margin and M
<subScript id="1798911CC136FFEAFCEBFA73FCADFA7C" attach="left" box="[829,837,1483,1497]" fontSize="6" pageId="3" pageNumber="435">1</subScript>
. Adjacent submarginal band pale, extending to costal margin at apex with darker marginal band between apex and almost to Rs4 (
<figureCitation id="13278FDCC136FFEAFC20FA5BFBBFFA5E" box="[1014,1111,1507,1532]" captionStart="FIGURE 12" captionStartId="9.[151,250,1873,1895]" captionTargetBox="[355,1236,601,1836]" captionTargetId="figure@9.[340,1247,591,1851]" captionTargetPageId="9" captionText="FIGURE 12. Male FW venation, discal shading (blue outline) and anterior recess (green outline): 12a, E. meifenga sp. n. HT); 12b, E. chalybeatus (India, Meghalaya, NHMUK); 12c, E. davidi (holotype, China, MNHN), 12d, E. kosemponis (LT); 12e, E. magnus (Shenbaganur, India, NHMUK); 12f, E. malabaricus (Ooty, India, NHMUK); 12g, E. signifer (LT, Meghalaya, India, NHMUK); 12h, E. sinensis (NMNS); ad—anterior discal cell, md—medial discal cell, pd—posterior discal cell, pdbpostdiscal band, premarginal band, smb—submarginal band, mb—marginal band." httpUri="https://zenodo.org/record/2623060/files/figure.png" pageId="3" pageNumber="435">Fig. 12a</figureCitation>
). An ovoid white spot edged basally with dark olive green is located on the basal side of the M
<subScript id="1798911CC136FFEAFCB9F9AEFC9FF981" attach="left" box="[879,887,1558,1572]" fontSize="6" pageId="3" pageNumber="435">1</subScript>
–M
<subScript id="1798911CC136FFEAFC48F9AEFC4EF981" attach="left" box="[926,934,1558,1572]" fontSize="6" pageId="3" pageNumber="435">2</subScript>
cross vein. Costal region with 4–5 pale olive patches edged with pale greenish white and separated by pale grey. Ventral FW with costal pocket; pale grey to greyish green with dorsal ornamentation between costal margin and Sc; posteriorly oriented row of piliform scales along Sc; softer piliform scales across much of the basal and central wing surface. Dorsal HW grey with pale olive green scales over anterior apical region and marginal scales. Ventral HW pale grey to greyish green with dorsal ornamentation between costal margin and Sc. Legs pale greyish olive; leg length ratio pro: meso: meta 1:1.2:0.77; proleg lacking epiphysis; metatibia with yellowish to reddish brown androconia and large androconial gland along dorsal length of tibia (
<figureCitation id="13278FDCC136FFEAFE59F8BDFE09F8BB" box="[399,481,1797,1822]" captionStart="FIGURES 4–5" captionStartId="5.[151,264,1609,1632]" captionTargetBox="[188,1398,733,1588]" captionTargetId="figure@5.[188,1398,733,1588]" captionTargetPageId="5" captionText="FIGURES 4–5. Thoracic. appendages: 4, wing venation. 4a, Endoclita meifenga sp. n (HT); 4b, E. kosemponis (LT). 5, Endoclita male legs. 5a E. meifenga sp. n; 5b, E. kosemponis LT. 5c, E. sinensis (NMNS)" httpUri="https://zenodo.org/record/2623053/files/figure.png" pageId="3" pageNumber="435">Fig. 5a</figureCitation>
).
</paragraph>
<paragraph id="8BA39359C136FFEDFF11F891FE1FFEB9" blockId="3.[151,1437,1144,2002]" lastBlockId="4.[151,1436,151,824]" lastPageId="4" lastPageNumber="436" pageId="3" pageNumber="435">
<emphasis id="B9684F4BC136FFEAFF11F891FED9F8E7" box="[199,305,1833,1858]" italics="true" pageId="3" pageNumber="435">Abdomen</emphasis>
(
<figureCitation id="13278FDCC136FFEAFE92F891FE07F8E7" box="[324,495,1833,1858]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1745,1768]" captionTargetBox="[151,1436,747,1725]" captionTargetId="figure@6.[151,1436,747,1725]" captionTargetPageId="6" captionText="FIGURES 6–8. Abdomen of Endoclita species (male unless otherwise specified): 6, second tergite and sternite. 6a, E. meifenga sp. n. (HT); 6b, E. kosemponis (LT); 6c, E. kosemponis female (ST), missing anterior margin of sternum II; 6d, E. sinensis (NMNS). 7, tergosternal connection. 7a, E. meifenga sp. n. (HT); 7b, E. kosemponis (LT). 8, tergites and sternite VIII. 8a, E. meifenga sp. n. (LT); 8b, E. kosemponis (LT); 8c, E. sinensis (vm—detached saccular margin of the genitalia) (NMNS); 8d, E. kosemponis female (ST)." httpUri="https://zenodo.org/record/2623056/files/figure.png" pageId="3" pageNumber="435">Fig. 6a, 7a, 8a</figureCitation>
) Pale greyish olive; tergites and sternites moderately sclerotized; tergum II with lateral ridge extending anterio-medially to lateral tuberculate plate, anterior ridge extending from tergosternal connection to dorsal median (
<figureCitation id="13278FDCC136FFEAFE30F8C9FDD0F82F" box="[486,568,1905,1930]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1745,1768]" captionTargetBox="[151,1436,747,1725]" captionTargetId="figure@6.[151,1436,747,1725]" captionTargetPageId="6" captionText="FIGURES 6–8. Abdomen of Endoclita species (male unless otherwise specified): 6, second tergite and sternite. 6a, E. meifenga sp. n. (HT); 6b, E. kosemponis (LT); 6c, E. kosemponis female (ST), missing anterior margin of sternum II; 6d, E. sinensis (NMNS). 7, tergosternal connection. 7a, E. meifenga sp. n. (HT); 7b, E. kosemponis (LT). 8, tergites and sternite VIII. 8a, E. meifenga sp. n. (LT); 8b, E. kosemponis (LT); 8c, E. sinensis (vm—detached saccular margin of the genitalia) (NMNS); 8d, E. kosemponis female (ST)." httpUri="https://zenodo.org/record/2623056/files/figure.png" pageId="3" pageNumber="435">Fig. 6a</figureCitation>
); tergosternal connection with broad, triangular central region, lacking tergal knob, lateral and dorsal brace almost at right angles, dorsal brace not fused to anterior ridge of TII (
<figureCitation id="13278FDCC136FFEAFAECF82DFA66F80B" box="[1338,1422,1941,1966]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1745,1768]" captionTargetBox="[151,1436,747,1725]" captionTargetId="figure@6.[151,1436,747,1725]" captionTargetPageId="6" captionText="FIGURES 6–8. Abdomen of Endoclita species (male unless otherwise specified): 6, second tergite and sternite. 6a, E. meifenga sp. n. (HT); 6b, E. kosemponis (LT); 6c, E. kosemponis female (ST), missing anterior margin of sternum II; 6d, E. sinensis (NMNS). 7, tergosternal connection. 7a, E. meifenga sp. n. (HT); 7b, E. kosemponis (LT). 8, tergites and sternite VIII. 8a, E. meifenga sp. n. (LT); 8b, E. kosemponis (LT); 8c, E. sinensis (vm—detached saccular margin of the genitalia) (NMNS); 8d, E. kosemponis female (ST)." httpUri="https://zenodo.org/record/2623056/files/figure.png" pageId="3" pageNumber="435">Fig. 7a</figureCitation>
); Segment II pleurum with lateral pouch, sternum II elongate and laterally concave, anterio-lateral arms subrectangular and obliquely angled, anterior margin forming broad ‘V’ shape (
<figureCitation id="13278FDCC131FFEDFBD8FF2FFBB7FF15" box="[1038,1119,151,176]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1745,1768]" captionTargetBox="[151,1436,747,1725]" captionTargetId="figure@6.[151,1436,747,1725]" captionTargetPageId="6" captionText="FIGURES 6–8. Abdomen of Endoclita species (male unless otherwise specified): 6, second tergite and sternite. 6a, E. meifenga sp. n. (HT); 6b, E. kosemponis (LT); 6c, E. kosemponis female (ST), missing anterior margin of sternum II; 6d, E. sinensis (NMNS). 7, tergosternal connection. 7a, E. meifenga sp. n. (HT); 7b, E. kosemponis (LT). 8, tergites and sternite VIII. 8a, E. meifenga sp. n. (LT); 8b, E. kosemponis (LT); 8c, E. sinensis (vm—detached saccular margin of the genitalia) (NMNS); 8d, E. kosemponis female (ST)." httpUri="https://zenodo.org/record/2623056/files/figure.png" pageId="4" pageNumber="436">Fig. 6a</figureCitation>
, one lateral arm broken off); tergum VIII an isosceles trapezoid with lateral edges curving anteriorly and narrowing to posterior margin; sternum VIII sub-rectangular, slightly wider posteriorly, anterior and lateral margins straight, posterior margin with medial rectangular concavity (
<figureCitation id="13278FDCC131FFEDFE4EFEBCFE02FEB9" box="[408,490,260,285]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1745,1768]" captionTargetBox="[151,1436,747,1725]" captionTargetId="figure@6.[151,1436,747,1725]" captionTargetPageId="6" captionText="FIGURES 6–8. Abdomen of Endoclita species (male unless otherwise specified): 6, second tergite and sternite. 6a, E. meifenga sp. n. (HT); 6b, E. kosemponis (LT); 6c, E. kosemponis female (ST), missing anterior margin of sternum II; 6d, E. sinensis (NMNS). 7, tergosternal connection. 7a, E. meifenga sp. n. (HT); 7b, E. kosemponis (LT). 8, tergites and sternite VIII. 8a, E. meifenga sp. n. (LT); 8b, E. kosemponis (LT); 8c, E. sinensis (vm—detached saccular margin of the genitalia) (NMNS); 8d, E. kosemponis female (ST)." httpUri="https://zenodo.org/record/2623056/files/figure.png" pageId="4" pageNumber="436">Fig. 8a</figureCitation>
).
</paragraph>
<paragraph id="8BA39359C131FFEDFF11FE9FFD55FD99" blockId="4.[151,1436,151,824]" pageId="4" pageNumber="436">
<emphasis id="B9684F4BC131FFEDFF11FE9FFE87FEE5" box="[199,367,295,320]" italics="true" pageId="4" pageNumber="436">Male genitalia</emphasis>
(
<figureCitation id="13278FDCC131FFEDFE57FE9FFE3EFEE5" box="[385,470,295,320]" captionStart="FIGURES 9–11" captionStartId="7.[151,264,1480,1503]" captionTargetBox="[217,1369,337,1459]" captionTargetId="figure@7.[217,1369,337,1459]" captionTargetPageId="7" captionText="FIGURES 9–11. Genitalia: 9a, E. meifenga sp. n (HT); 9b, E. kosemponis (LT): dpp—dorsal posterior projection, pvfposterio-ventral flange, mpp—medial posterior projection, mt—medial tooth, avn—anterio-ventral nexus, va—valva; 9c, E. sinensis (vincular margin with medial spine detached with abdomen—ref. Fig. 8c); 9d, E. davidi (HT). 10, external female genitlaia: 10a, E. kosemponis; 10b, E. sinensis. 11, internal female genitalia: 11a, E. kosemponis; 11b, E. sinensis." httpUri="https://zenodo.org/record/2623058/files/figure.png" pageId="4" pageNumber="436">Fig. 9a</figureCitation>
). Tegumen (intermediate plate) elongate. Saccus (vinculum) triangular, lateral edge almost straight, posterior margin with medial apical tooth and lateral, curved apodemal ridges. Tergal lobes not observed. Pseudotegumen well developed, margin adjacent to anogenital field with a ragged or broken edge with a triangular dorso-posterior projection, a blunt medial posterior projection angled to the median, and a strongly sclerotized ventral W-shaped bridge meeting across the median as an anterio-ventral nexus, and dorso-posterior margin fused across the median forming a narrow tubular junction. Fultura superior membranous, fultura and inferior sub-square with concave lateral edges. Valva small relative to pseudotegumen, narrow, digitiform, apex rounded. Phallus membranous, without cornutus.
</paragraph>
<paragraph id="8BA39359C131FFEDFF11FDFFFE1DFD0D" blockId="4.[151,1436,151,824]" pageId="4" pageNumber="436">
<emphasis id="B9684F4BC131FFEDFF11FDFFFE62FDC5" bold="true" box="[199,394,583,608]" pageId="4" pageNumber="436">DNA sequences:</emphasis>
The following sequences were obtained from the COI gene and were processed according to the BOLD System (boldsystems.org/). In the absence of comparable species in BOLD, only the available sequences are presented below.
</paragraph>
<paragraph id="8BA39359C131FFEDFF11FD0CFE78FD68" blockId="4.[151,1436,151,824]" box="[199,400,692,717]" pageId="4" pageNumber="436">Female unknown.</paragraph>
<paragraph id="8BA39359C131FFEDFF11FD6FFD48FCB0" blockId="4.[151,1436,151,824]" pageId="4" pageNumber="436">
<emphasis id="B9684F4BC131FFEDFF11FD6FFCACFD55" bold="true" box="[199,836,727,752]" pageId="4" pageNumber="436">
<taxonomicName id="4C1CE8DAC131FFEDFF11FD6FFD00FD55" authority="Buchsbaum &amp; Grehan" authorityName="Buchsbaum &amp; Grehan" authorityYear="2019" box="[199,744,727,752]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="436" phylum="Arthropoda" rank="species" species="meifenga" status="sp. nov.">
<emphasis id="B9684F4BC131FFEDFF11FD6FFE5BFD55" bold="true" box="[199,435,727,752]" italics="true" pageId="4" pageNumber="436">Endoclita meifenga</emphasis>
Buchsbaum &amp; Grehan
</taxonomicName>
<taxonomicNameLabel id="A25BF230C131FFEDFD2CFD6FFCACFD55" box="[762,836,727,752]" pageId="4" pageNumber="436" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Sample ID: BC ZSM Lep 59526, Process ID: GWOTK290-12, BIN URI: BOLD: ABX1145
</paragraph>
<paragraph id="8BA39359C131FFEDFF11FCA7FDF0FC9D" blockId="4.[151,1436,151,824]" box="[199,536,799,824]" pageId="4" pageNumber="436">Sample Nucleotide Sequence:</paragraph>
<paragraph id="8BA39359C131FFEDFF41FCD0FDA4FB61" blockId="4.[151,1436,872,1220]" box="[151,1436,872,1220]" pageId="4" pageNumber="436">AACTTTATATTTTATTTTTGGTATTTGAGCAGGAATAATTGGTACTTCATTAAGATTATTAATTCGAACAG AATTAGGAAACCCAGGATCTTTAATTGGAGATGATCAAATTTATAATGTAATTGTAACAGCACATGCTTT TATTATAATTTTTTTTATAGTTATACCAATTATAATTGGAGGATTTGGAAATTGATTAGTTCCATTAATATT AGGAGCACCAGATATAGCTTTCCCACGATTAAATAATATAAGATTCTGGCTATTACCCCCATCATTAATA TTATTAATTTCTAGAAGAATCGTTGAAAATGGAGCAGGAACAGGCTGAACTGTTTACCCCCCACTATCT GCAAATATTGCACATGCTGGAAGATCTGTTGATTTAGCAATTTTTTCTCTACATTTAGCAGGAATTTCTT CTATTTTAGGGGCAGTAAATTTTATTACAACTGTAATTAATATACGATCAGAAGGAATATCTTTTGATCG AATACCTTTATTTGTTTGAAGAGTAGCAATTACTGCTTTATTACTTTTATTATCATTACCTGTACTAGCAG GAGCTATTACTATATTATTAACAGATCGAAATTTAAATACTTCATTTTTTGATCCTGCGGGAGGAGGAGA TCCAATTTTATATCAACATTTATTT</paragraph>
<paragraph id="8BA39359C131FFEDFF41FB4CFE7CFAA9" blockId="4.[151,404,1268,1293]" box="[151,404,1268,1293]" pageId="4" pageNumber="436">Amino Acid Sequence</paragraph>
<paragraph id="8BA39359C131FFEDFF41FA84FF50FA65" blockId="4.[151,1434,1340,1472]" box="[151,1434,1340,1472]" pageId="4" pageNumber="436">TLYFIFGIWAGMIGTSLSLLIRTELGNPGSLIGDDQIYNVIVTAHAFIMIFFMVMPIMIGGFGNWLVPLMLGA PDMAFPRLNNMSFWLLPPSLMLLISSSIVENGAGTGWTVYPPLSANIAHAGSSVDLAIFSLHLAGISSILGA VNFITTVINMRSEGMSFDRMPLFVWSVAITALLLLLSLPVLAGAITMLLTDRNLNTSFFDPAGGGDPILYQH LF</paragraph>
</subSubSection>
<subSubSection id="C306C0D2C131FFEDFF11FA57FC93F91A" pageId="4" pageNumber="436" type="distribution">
<paragraph id="8BA39359C131FFEDFF11FA57FC93F91A" blockId="4.[151,1436,1519,1727]" pageId="4" pageNumber="436">
<emphasis id="B9684F4BC131FFEDFF11FA57FE88F9AD" bold="true" box="[199,352,1519,1544]" pageId="4" pageNumber="436">Distribution.</emphasis>
Known only from the
<typeStatus id="54A72DFBC131FFEDFD89FA49FD67F9AC" box="[607,655,1521,1545]" pageId="4" pageNumber="436">type</typeStatus>
locality in northern
<collectingCountry id="F30BD3C9C131FFEDFCA5FA57FC2DF9AD" box="[883,965,1519,1544]" name="Taiwan" pageId="4" pageNumber="436">Taiwan</collectingCountry>
(
<figureCitation id="13278FDCC131FFEDFC02FA57FBCBF9AD" box="[980,1059,1519,1544]" captionStart="FIGURE 13" captionStartId="10.[151,250,1278,1300]" captionTargetBox="[151,1436,591,1256]" captionTargetId="figure@10.[151,1436,591,1256]" captionTargetPageId="10" captionText="FIGURE 13. Distribution of E. meifenga sp. n. 13a, forest habitat of the type locality at the Meifeng Experimental Farm (Photo Ulf Buchsbaum); 13b, type locality in Taiwan." httpUri="https://zenodo.org/record/2623062/files/figure.png" pageId="4" pageNumber="436">Fig. 13</figureCitation>
a–b). The Highland Experimental Farm (http://mf.shengchi.com.tw/enweb/) is an open area of agricultural crops and dry meadows at an elevation of about
<quantity id="4CE43EBCC131FFEDFF08F980FED3F9F5" box="[222,315,1592,1616]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="436" unit="m" value="2100.0">2,100 m</quantity>
and surrounded by natural mountain forest. In the days preceding collecting, the weather was dry and warm followed by misty and cloudy conditions with light rain and an evening temperature of about 15°C on the day of collection (May, 11
<superScript id="7C693E11C131FFEDFE2DF9C7FDEFF928" attach="none" box="[507,519,1663,1677]" fontSize="6" pageId="4" pageNumber="436">th</superScript>
) which took place just a few days before a typhoon hit
<collectingCountry id="F30BD3C9C131FFEDFB73F93AFB1FF93E" box="[1189,1271,1666,1691]" name="Taiwan" pageId="4" pageNumber="436">Taiwan</collectingCountry>
(the principal typhoon season usually lasting from about June through October).
</paragraph>
</subSubSection>
</treatment>
</document>
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