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<mods:title id="46B0F9896D3EA2C11F8804E5A92D2DA6">Phylogeny and Taxonomic Revision of the Endemic Malagasy Genus Ptychochromis (Teleostei: Cichlidae), with the Description of Five New Species and a Diagnosis for Katria, New Genus</mods:title>
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<subSubSection id="C327657CF070FF85D5AFFADE014CFB59" box="[729,1120,1213,1237]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8B8236F7F070FF85D5AFFADE014CFB59" blockId="13.[729,1120,1213,1237]" box="[729,1120,1213,1237]" pageId="13" pageNumber="14">
<heading id="D0CA819BF070FF85D5AFFADE014CFB59" box="[729,1120,1213,1237]" centered="true" fontSize="36" level="2" pageId="13" pageNumber="14" reason="8">
<taxonomicName id="4C3D4D74F070FF85D5AFFADE014CFB59" ID-CoL="756X" ID-ENA="96999" authority="Steindachner, 1880" authorityName="Steindachner" authorityYear="1880" box="[729,1120,1213,1237]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B949EAE5F070FF85D5AFFADE0651FB59" bold="true" box="[729,893,1213,1237]" italics="true" pageId="13" pageNumber="14">Ptychochromis</emphasis>
<bibRefCitation id="EFAC4B06F070FF85D4F1FADE014CFB59" author="Steindachner, F." box="[903,1120,1213,1237]" pageId="13" pageNumber="14" pagination="238 - 266" refId="ref30605" refString="Steindachner, F. 1880. Ichthyologische Beitrage (IX). I. Uber eine Sammlung von Flussfischen von Tohizona auf Madagascar. II. Uber zwei neue * Agonus * - Arten aus Californien. III. Uber einige Fischarten aus dem nordlichen Japan, gesammelt von Professor Dybowski. Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Klasse der Kaiserlichen Akademie der Wissenschaften 82: 238 - 266." type="journal article" year="1880">Steindachner, 1880</bibRefCitation>
</taxonomicName>
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<paragraph id="8B8236F7F070FF85D5C6FA8B0188FA65" blockId="13.[662,1188,1256,1514]" pageId="13" pageNumber="14">
DIAGNOSIS: A ptychochromin genus diagnosed by the presence of a free second epibranchial toothplate (fig. 7A,
<emphasis id="B949EAE5F070FF85D343FB4107F8FAD4" italics="true" pageId="13" pageNumber="14">Character 10: 1</emphasis>
), the third infraorbital (IO3) excluded from the orbit by IO2 (fig. 8F,
<emphasis id="B949EAE5F070FF85D343FB3E07EAFA1E" italics="true" pageId="13" pageNumber="14">Character 11:1</emphasis>
), the presence of a well-developed palatine groove (fig. 6FG,
<emphasis id="B949EAE5F070FF85D40EFBF4010EFA23" box="[888,1058,1431,1455]" italics="true" pageId="13" pageNumber="14">Character 12:1</emphasis>
), and ventral displacement of the palatine-palatine ligament insertion (fig. 6FG,
<emphasis id="B949EAE5F070FF85D49BFBB101B8FA66" box="[1005,1172,1490,1514]" italics="true" pageId="13" pageNumber="14">Character 13:1</emphasis>
).
</paragraph>
<paragraph id="8B8236F7F070FF85D5A5F8740149F9A3" blockId="13.[723,1125,1559,1583]" box="[723,1125,1559,1583]" pageId="13" pageNumber="14">
<taxonomicName id="4C3D4D74F070FF85D5A5F87406FEF9A3" authority="STIASSNY &amp; SPARKS, 2006" authorityName="STIASSNY &amp; SPARKS" authorityYear="2006" box="[723,978,1559,1583]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="onilahy" status="sp. nov.">
<emphasis id="B949EAE5F070FF85D5A5F87406FEF9A3" bold="true" box="[723,978,1559,1583]" italics="true" pageId="13" pageNumber="14">Ptychochromis onilahy</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27A579EF070FF85D494F8740149F9A3" box="[994,1125,1559,1583]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
</paragraph>
<paragraph id="8B8236F7F070FF85D5A9F8570176F9C0" blockId="13.[735,1114,1588,1612]" box="[735,1114,1588,1612]" pageId="13" pageNumber="14">
<heading id="D0CA819BF070FF85D5A9F8570176F9C0" box="[735,1114,1588,1612]" centered="true" fontSize="36" level="3" pageId="13" pageNumber="14" reason="7">
<figureCitation id="13062A72F070FF85D5A9F85706AEF9C0" box="[735,898,1588,1612]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="17.[112,150,1705,1727]" captionStartId-1="18.[151,189,1004,1026]" captionTargetBox-0="[118,1156,911,1669]" captionTargetBox-1="[154,1200,206,970]" captionTargetId-0="figure-251@17.[118,1157,911,1670]" captionTargetId-1="figure-307@18.[154,1201,206,970]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Fig. 11. Ptychochromis onilahy, new species, holotype, MNHN 1962-0201, 84.3 mm SL, southwestern Madagascar, Province of Tulear, Onilahy River. Scale bar 5 1 cm. Drawing by Ian Hart." captionText-1="Fig. 12. Ptychochromis onilahy, new species, paratype, MNHN 2006-0780: (A) palatine, (B) LPJ, (C) first gill arch, and (D) infraorbital series." figureDoi-0="http://doi.org/10.5281/zenodo.5385357" figureDoi-1="http://doi.org/10.5281/zenodo.5385361" httpUri-0="https://zenodo.org/record/5385357/files/figure.png" httpUri-1="https://zenodo.org/record/5385361/files/figure.png" pageId="13" pageNumber="14">Figures 1112</figureCitation>
, Plate 2A, Table 4
</heading>
</paragraph>
<paragraph id="8B8236F7F070FF85D5E0F8170617F928" blockId="13.[662,1189,1652,1754]" pageId="13" pageNumber="14">
<taxonomicName id="4C3D4D74F070FF85D5E0F817069AF905" baseAuthorityName="Bleeker" baseAuthorityYear="1868" box="[662,950,1652,1673]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="oligacanthus">
<emphasis id="B949EAE5F070FF85D5E0F817069AF905" box="[662,950,1652,1673]" italics="true" pageId="13" pageNumber="14">Ptychochromis oligacanthus</emphasis>
</taxonomicName>
, race du Sud-Ouest:
<bibRefCitation id="EFAC4B06F070FF85D5C6F8ED061BF928" author="Kiener, A." box="[688,823,1678,1700]" pageId="13" pageNumber="14" pagination="1 - 244" refId="ref29702" refString="Kiener, A. 1963. Poissons, peche et pisciculture a Madagascar. Publication Centre Technique Forestier Tropical 24: 1 - 244." type="journal article" year="1963">Kiener, 1963</bibRefCitation>
.
</paragraph>
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<paragraph id="8B8236F7F070FF85D5E0F8CA01B7F956" blockId="13.[662,1189,1652,1754]" pageId="13" pageNumber="14">
<taxonomicName id="4C3D4D74F070FF85D5E0F8CA0690F932" baseAuthorityName="Bleeker" baseAuthorityYear="1868" box="[662,956,1705,1726]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="oligacanthus">
<emphasis id="B949EAE5F070FF85D5E0F8CA0690F932" box="[662,956,1705,1726]" italics="true" pageId="13" pageNumber="14">Ptychochromis oligacanthus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3D4D74F070FF85D4AFF8CA0188F933" box="[985,1188,1705,1727]" pageId="13" pageNumber="14" rank="subSpecies" subSpecies="geographique">race géographique</taxonomicName>
Sud-Ouest: Kiener and Mauge´, 1966, fig. 26.
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