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<document id="18D9595E7019194E91ECAA930BA73D5F" ID-DOI="10.11646/zootaxa.3637.3.8" ID-GBIF-Dataset="f1795ddd-2437-46c3-8d8f-2c3430565140" ID-ISSN="1175-5326" ID-Zenodo-Dep="221710" ID-ZooBank="19493296-421C-475E-AA1A-39129F2A4BBF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460591862340" checkinUser="plazi" docAuthor="Su, Yirui, Lu, Jinming &amp; Chen, Hongwei" docDate="2013" docId="038BF915FFF2FF93D3F0FA2120E3FDE7" docLanguage="en" docName="zt03637p373.pdf" docOrigin="Zootaxa 3637 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucophenga abbreviata" docType="treatment" docVersion="8" lastPageNumber="365" masterDocId="FFB2816DFFF1FF97D367FFDC2144FFF0" masterDocTitle="The genus Leucophenga (Diptera, Drosophilidae), part I: the abbreviata species group from the Oriental region with morphological and molecular evidence" masterLastPageNumber="373" masterPageNumber="361" pageNumber="364" updateTime="1698326086797" updateUser="plazi">
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<mods:title id="F19BB0D69A27A6B30383C3ADAC98AE02">The genus Leucophenga (Diptera, Drosophilidae), part I: the abbreviata species group from the Oriental region with morphological and molecular evidence</mods:title>
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<mods:namePart id="1102455397E608ECFDD2781B46DB7B95">Su, Yirui</mods:namePart>
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<mods:namePart id="9BF43DC78A11F5CE0C3CFC9BD0F9EECD">Lu, Jinming</mods:namePart>
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<mods:namePart id="5A7ACE1452E8298336C16C43846B13CD">Chen, Hongwei</mods:namePart>
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<mods:title id="A340DCEF25F6FB73A15EBA7EC2E788BF">Zootaxa</mods:title>
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<mods:date id="AD70D4D4B1FD5DC948BA5747F8B09ABD">2013</mods:date>
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<mods:number id="A968E723E6D1D4A1CA2BA948BE9D4473">3637</mods:number>
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<mods:identifier id="E772EF157FB9A773C47897485478374B" type="DOI">10.11646/zootaxa.3637.3.8</mods:identifier>
<mods:identifier id="BA38581B1E83276BA07E5BB84D9E86E6" type="GBIF-Dataset">f1795ddd-2437-46c3-8d8f-2c3430565140</mods:identifier>
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<treatment id="038BF915FFF2FF93D3F0FA2120E3FDE7" ID-DOI="http://doi.org/10.5281/zenodo.6157012" ID-GBIF-Taxon="119565826" ID-Zenodo-Dep="6157012" LSID="urn:lsid:plazi:treatment:038BF915FFF2FF93D3F0FA2120E3FDE7" httpUri="http://treatment.plazi.org/id/038BF915FFF2FF93D3F0FA2120E3FDE7" lastPageId="4" lastPageNumber="365" pageId="3" pageNumber="364">
<subSubSection id="C3381B88FFF2FF94D3F0FA212334F9E7" box="[151,624,1533,1559]" pageId="3" pageNumber="364" type="nomenclature">
<paragraph id="8B9D4803FFF2FF94D3F0FA212334F9E7" blockId="3.[151,624,1533,1559]" box="[151,624,1533,1559]" pageId="3" pageNumber="364">
<heading id="D0D5FF6FFFF2FF94D3F0FA212334F9E7" bold="true" box="[151,624,1533,1559]" fontSize="11" level="1" pageId="3" pageNumber="364" reason="1">
<emphasis id="B9569411FFF2FF94D3F0FA212334F9E7" bold="true" box="[151,624,1533,1559]" pageId="3" pageNumber="364">
<taxonomicName id="4C223380FFF2FF94D3F0FA212085F9E7" ID-CoL="6PT2Y" box="[151,449,1533,1559]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="abbreviata">
<emphasis id="B9569411FFF2FF94D3F0FA212085F9E7" bold="true" box="[151,449,1533,1559]" italics="true" pageId="3" pageNumber="364">Leucophenga abbreviata</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3381B88FFF2FF94D3F0F99F2507F9AC" box="[151,1091,1603,1633]" pageId="3" pageNumber="364" type="diagnosis">
<paragraph id="8B9D4803FFF2FF94D3F0F99F2507F9AC" blockId="3.[151,1436,1603,2031]" box="[151,1091,1603,1633]" pageId="3" pageNumber="364">
<emphasis id="B9569411FFF2FF94D3F0F99F2055F9AB" bold="true" box="[151,273,1603,1628]" pageId="3" pageNumber="364">Diagnosis.</emphasis>
M1 distally abbreviated, not reaching wing margin (
<figureCitation id="13195486FFF2FF94D03FF99822DEF9AC" box="[856,922,1604,1628]" captionStart="FIGURE 3" captionStartId="5.[151,249,1523,1546]" captionTargetBox="[151,1435,359,1492]" captionTargetId="figure@5.[151,1436,359,1492]" captionTargetPageId="5" captionText="FIGURE 3. Wing, mesonotum, scutellum and pleura of male: A, B, C. Leucophenga abbreviata (de Meijere, 1911); D, E, F. Leucophenga brevivena sp. nov.; G, H, I. Leucophenga sujuanae sp. nov.; J, K, L. Leucophenga zhenfangae sp. nov." httpUri="https://zenodo.org/record/221713/files/figure.png" pageId="3" pageNumber="364">Fig. 3</figureCitation>
A, 3D, 3G, 3J).
</paragraph>
</subSubSection>
<subSubSection id="C3381B88FFF2FF93D3A0F9B620E3FDE7" lastPageId="4" lastPageNumber="365" pageId="3" pageNumber="364" type="description">
<paragraph id="8B9D4803FFF2FF93D3A0F9B620B8FE3F" blockId="3.[151,1436,1603,2031]" lastBlockId="4.[151,1436,151,535]" lastPageId="4" lastPageNumber="365" pageId="3" pageNumber="364">
<emphasis id="B9569411FFF2FF94D3A0F9B6201CF972" bold="true" box="[199,344,1642,1667]" pageId="3" pageNumber="364">Description.</emphasis>
Male and female: Eyes red to brownish red. Ocellar triangle dark brown to black, with a pair of setae above ocellar setae. Postocellar seta usually small. Frons brown, narrow, nearly parallel, with a few minute setulae medially. All orbital setae large; proclinate and anterior reclinate orbital setae very close together, separated by distance less than 1/2 of that between anterior reclinate and posterior reclinate. Pedicel yellow; first flagellomere brownish; arista long plumose. Face mostly yellow; facial carina undeveloped. Clypeus yellow. Palpus brownish yellow, slender in both sexes. Vibrissa prominent; other orals small. Gena and postgena narrow. Mesonotum yellow to brown, not pollinose. Postpronotal lobe with 2–4 long setae and a few of shorter setae. Acrostichal setulae in ca. 12–14 irregular rows. Prescutellar setae large. Katepisternum yellowish, with small setae medially, and 2 large ones anteriorly and posteriorly, respectively. Subscutellum swollen. Basal medial-cubital crossvein absent. Wing costal vein between R2+3 and R4+5 distally with more than 5, 6 peg-like spinules on ventral surface; R2+3 sometimes curved to costa at tip. Halter mostly yellowish white. Legs mostly yellowish. Abdominal tergites variable in the color and pattern. Male terminalia: Epandrium usually with sparse pubescence and several setae around posterodorsal to ventral margins; apodeme usually developed (
<figureCitation id="13195486FFF5FF93D1CDFF602255FF24" box="[682,785,188,212]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="6.[151,249,1326,1349]" captionStartId-1="7.[151,249,1718,1741]" captionStartId-2="8.[151,249,1857,1880]" captionStartId-3="9.[151,249,1689,1712]" captionTargetBox-0="[366,1220,745,1293]" captionTargetBox-1="[360,1226,1107,1690]" captionTargetBox-2="[370,1214,1192,1812]" captionTargetBox-3="[358,1225,1105,1649]" captionTargetId-0="figure@6.[355,1232,735,1305]" captionTargetId-1="figure@7.[355,1231,1098,1696]" captionTargetId-2="figure@8.[366,1220,1175,1826]" captionTargetId-3="figure@9.[352,1234,1096,1666]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 5. Leucophenga abbreviata (de Meijere, 1911), male terminalia. A. Epandrium (epan), cercus (cer) and surstylus (sur) (lateral view); B. Hypandrium (hypd), paramere (pm) and aedeagal apodeme (aed a) (lateral view); C. Aedeagus (aed) and gonopods (gon) (lateral view). Scale lines = 0.1 mm." captionText-1="FIGURE 6. Leucophenga brevivena sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." captionText-2="FIGURE 7. Leucophenga sujuanae sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." captionText-3="FIGURE 8. Leucophenga zhenfangae sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." httpUri-0="https://zenodo.org/record/221715/files/figure.png" httpUri-1="https://zenodo.org/record/221716/files/figure.png" httpUri-2="https://zenodo.org/record/221717/files/figure.png" httpUri-3="https://zenodo.org/record/221718/files/figure.png" pageId="4" pageNumber="365">Figs 5–8</figureCitation>
A). Surstylus broad, flat, nearly entirely pubescent, with several setae on outer and inner surface (
<figureCitation id="13195486FFF5FF93D138FF032387FF07" box="[607,707,223,247]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="6.[151,249,1326,1349]" captionStartId-1="7.[151,249,1718,1741]" captionStartId-2="8.[151,249,1857,1880]" captionStartId-3="9.[151,249,1689,1712]" captionTargetBox-0="[366,1220,745,1293]" captionTargetBox-1="[360,1226,1107,1690]" captionTargetBox-2="[370,1214,1192,1812]" captionTargetBox-3="[358,1225,1105,1649]" captionTargetId-0="figure@6.[355,1232,735,1305]" captionTargetId-1="figure@7.[355,1231,1098,1696]" captionTargetId-2="figure@8.[366,1220,1175,1826]" captionTargetId-3="figure@9.[352,1234,1096,1666]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 5. Leucophenga abbreviata (de Meijere, 1911), male terminalia. A. Epandrium (epan), cercus (cer) and surstylus (sur) (lateral view); B. Hypandrium (hypd), paramere (pm) and aedeagal apodeme (aed a) (lateral view); C. Aedeagus (aed) and gonopods (gon) (lateral view). Scale lines = 0.1 mm." captionText-1="FIGURE 6. Leucophenga brevivena sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." captionText-2="FIGURE 7. Leucophenga sujuanae sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." captionText-3="FIGURE 8. Leucophenga zhenfangae sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." httpUri-0="https://zenodo.org/record/221715/files/figure.png" httpUri-1="https://zenodo.org/record/221716/files/figure.png" httpUri-2="https://zenodo.org/record/221717/files/figure.png" httpUri-3="https://zenodo.org/record/221718/files/figure.png" pageId="4" pageNumber="365">Figs 5–8</figureCitation>
A). Cercus separated from epandrium, with several setae, lacking pubescence (
<figureCitation id="13195486FFF5FF93D24CFED820D0FEEC" box="[299,404,260,284]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="6.[151,249,1326,1349]" captionStartId-1="7.[151,249,1718,1741]" captionStartId-2="8.[151,249,1857,1880]" captionStartId-3="9.[151,249,1689,1712]" captionTargetBox-0="[366,1220,745,1293]" captionTargetBox-1="[360,1226,1107,1690]" captionTargetBox-2="[370,1214,1192,1812]" captionTargetBox-3="[358,1225,1105,1649]" captionTargetId-0="figure@6.[355,1232,735,1305]" captionTargetId-1="figure@7.[355,1231,1098,1696]" captionTargetId-2="figure@8.[366,1220,1175,1826]" captionTargetId-3="figure@9.[352,1234,1096,1666]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 5. Leucophenga abbreviata (de Meijere, 1911), male terminalia. A. Epandrium (epan), cercus (cer) and surstylus (sur) (lateral view); B. Hypandrium (hypd), paramere (pm) and aedeagal apodeme (aed a) (lateral view); C. Aedeagus (aed) and gonopods (gon) (lateral view). Scale lines = 0.1 mm." captionText-1="FIGURE 6. Leucophenga brevivena sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." captionText-2="FIGURE 7. Leucophenga sujuanae sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." captionText-3="FIGURE 8. Leucophenga zhenfangae sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." httpUri-0="https://zenodo.org/record/221715/files/figure.png" httpUri-1="https://zenodo.org/record/221716/files/figure.png" httpUri-2="https://zenodo.org/record/221717/files/figure.png" httpUri-3="https://zenodo.org/record/221718/files/figure.png" pageId="4" pageNumber="365">Figs 5–8</figureCitation>
A). Hypandrium (gonopod in Bächli
<emphasis id="B9569411FFF5FF93D02DFED922C4FEEC" box="[842,896,260,284]" italics="true" pageId="4" pageNumber="365">et al</emphasis>
. 2004) anteriorly fused to aedeagal apodeme, laterally broad, usually with paramedian setae subbasally. Gonopods (dorsal arch in Bächli
<emphasis id="B9569411FFF5FF93D7FAFEF5258BFEB0" box="[1181,1231,296,320]" italics="true" pageId="4" pageNumber="365">et al</emphasis>
. 2004) fused with each other, forming slightly triangular plate, anterioventrally with curved, median rod. Paramere (outer paraphysis in Bächli
<emphasis id="B9569411FFF5FF93D262FEAD2073FE78" box="[261,311,368,392]" italics="true" pageId="4" pageNumber="365">et al</emphasis>
. 2004) contiguous to arm of aedeagal apodeme basally, lacking pubescence (
<figureCitation id="13195486FFF5FF93D7F8FEB32441FE77" box="[1183,1285,367,391]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="6.[151,249,1326,1349]" captionStartId-1="7.[151,249,1718,1741]" captionStartId-2="8.[151,249,1857,1880]" captionStartId-3="9.[151,249,1689,1712]" captionTargetBox-0="[366,1220,745,1293]" captionTargetBox-1="[360,1226,1107,1690]" captionTargetBox-2="[370,1214,1192,1812]" captionTargetBox-3="[358,1225,1105,1649]" captionTargetId-0="figure@6.[355,1232,735,1305]" captionTargetId-1="figure@7.[355,1231,1098,1696]" captionTargetId-2="figure@8.[366,1220,1175,1826]" captionTargetId-3="figure@9.[352,1234,1096,1666]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 5. Leucophenga abbreviata (de Meijere, 1911), male terminalia. A. Epandrium (epan), cercus (cer) and surstylus (sur) (lateral view); B. Hypandrium (hypd), paramere (pm) and aedeagal apodeme (aed a) (lateral view); C. Aedeagus (aed) and gonopods (gon) (lateral view). Scale lines = 0.1 mm." captionText-1="FIGURE 6. Leucophenga brevivena sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." captionText-2="FIGURE 7. Leucophenga sujuanae sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." captionText-3="FIGURE 8. Leucophenga zhenfangae sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." httpUri-0="https://zenodo.org/record/221715/files/figure.png" httpUri-1="https://zenodo.org/record/221716/files/figure.png" httpUri-2="https://zenodo.org/record/221717/files/figure.png" httpUri-3="https://zenodo.org/record/221718/files/figure.png" pageId="4" pageNumber="365">Figs 5–8</figureCitation>
B). Aedeagus glabrous (
<figureCitation id="13195486FFF5FF93D26DFE482036FE5C" box="[266,370,404,428]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="6.[151,249,1326,1349]" captionStartId-1="7.[151,249,1718,1741]" captionStartId-2="8.[151,249,1857,1880]" captionStartId-3="9.[151,249,1689,1712]" captionTargetBox-0="[366,1220,745,1293]" captionTargetBox-1="[360,1226,1107,1690]" captionTargetBox-2="[370,1214,1192,1812]" captionTargetBox-3="[358,1225,1105,1649]" captionTargetId-0="figure@6.[355,1232,735,1305]" captionTargetId-1="figure@7.[355,1231,1098,1696]" captionTargetId-2="figure@8.[366,1220,1175,1826]" captionTargetId-3="figure@9.[352,1234,1096,1666]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 5. Leucophenga abbreviata (de Meijere, 1911), male terminalia. A. Epandrium (epan), cercus (cer) and surstylus (sur) (lateral view); B. Hypandrium (hypd), paramere (pm) and aedeagal apodeme (aed a) (lateral view); C. Aedeagus (aed) and gonopods (gon) (lateral view). Scale lines = 0.1 mm." captionText-1="FIGURE 6. Leucophenga brevivena sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." captionText-2="FIGURE 7. Leucophenga sujuanae sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." captionText-3="FIGURE 8. Leucophenga zhenfangae sp. nov., male terminalia. A. Epandrium, cercus and surstylus; B. Hypandrium, paramere and aedeagal apodeme; C. Aedeagus and gonopods. Scale lines = 0.1 mm." httpUri-0="https://zenodo.org/record/221715/files/figure.png" httpUri-1="https://zenodo.org/record/221716/files/figure.png" httpUri-2="https://zenodo.org/record/221717/files/figure.png" httpUri-3="https://zenodo.org/record/221718/files/figure.png" pageId="4" pageNumber="365">Figs 5–8</figureCitation>
C); basal bridges contiguous to median rod of gonopod; apodeme with a pair of arms each contiguous to base of paramere.
</paragraph>
<paragraph id="8B9D4803FFF5FF93D3A0FE0120E3FDE7" blockId="4.[151,1436,151,535]" pageId="4" pageNumber="365">In the following descriptions of each species, only characters that depart from the upper universal characters are provided for brevity.</paragraph>
</subSubSection>
</treatment>
</document>
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