368 lines
36 KiB
XML
368 lines
36 KiB
XML
<document ID-DOI="10.3897/zookeys.12.167" ID-GBIF-Dataset="3026c846-a607-4826-b2a9-8f7a531baec5" ID-GBIF-Taxon="163971730" ID-ISSN="1313–2970" ID-Zenodo-Dep="576460" ID-ZooBank="B32EF8D5-C3C6-4C8C-8845-284125448425" LSID="urn:lsid:zoobank.org:act:B107B0B1-7423-466C-8E47-BE50F0EA0D5D" approvalRequired="25" approvalRequired_for_textStreams="25" checkinTime="1587995767728" checkinUser="plazi" docAuthor="Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & VandenSpiegel, Didier" docDate="2009" docId="936787D7FFB8FFBB8ABB4727FE5DFBDD" docLanguage="en" docName="ZK_article_1994.pdf" docOrigin="ZooKeys 12 (12)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Eutrichodesmus aster Golovatch, Geoffroy, Mauries & VandenSpiegel 2009, sp. n." docType="treatment" docUuid="B107B0B1-7423-466C-8E47-BE50F0EA0D5D" docUuidSource="ZooBank" docVersion="4" lastPageId="15" lastPageNumber="16" masterDocId="6F5EFFAFFFB3FFB48A374557FFC6FF89" masterDocTitle="Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species" masterLastPageNumber="46" masterPageNumber="1" pageId="11" pageNumber="12" updateTime="1643565021146" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei</mods:namePart>
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<mods:affiliation>Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Ecologie & Gestion de la Biodiversité, UMR 7179 du CNRS, Equipe EVOLTRAIT, Brunoy, France & D 362 Dc 0 - & CC</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mauriès, Jean-Paul</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Systématique et Evolution, Section Arthropodes, Paris, France & - & - 42 F & D 01 - 28 Ad- 43 F</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>VandenSpiegel, Didier</mods:namePart>
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<mods:affiliation>Musée Royal de l’Afrique centrale, Tervuren, Belgium & A & EE & F- 04802 E 68 Cb</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-06-18</mods:number>
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||
</mods:detail>
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||
<mods:detail type="volume">
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<mods:number>12</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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||
<mods:number>12</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>46</mods:end>
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</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.3897/zookeys.12.167</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">3026c846-a607-4826-b2a9-8f7a531baec5</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576460</mods:identifier>
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||
<mods:identifier type="ZooBank">B32EF8D5-C3C6-4C8C-8845-284125448425</mods:identifier>
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||
</mods:mods>
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<treatment ID-GBIF-Taxon="163971730" LSID="urn:lsid:zoobank.org:act:B107B0B1-7423-466C-8E47-BE50F0EA0D5D" httpUri="http://treatment.plazi.org/id/936787D7FFB8FFBB8ABB4727FE5DFBDD" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12">
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||
<subSubSection box="[140,1043,623,651]" pageId="11" pageNumber="12" type="nomenclature">
|
||
<paragraph blockId="11.[140,1043,623,721]" box="[140,1043,623,651]" pageId="11" pageNumber="12">
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||
<heading box="[140,1043,623,651]" level="2" pageId="11" pageNumber="12">
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||
<emphasis bold="true" box="[140,1043,623,651]" pageId="11" pageNumber="12">
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||
<taxonomicName authority="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityName="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityYear="2009" box="[140,966,623,651]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aster" status="sp. n.">
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<emphasis bold="true" box="[140,391,624,651]" italics="true" pageId="11" pageNumber="12">
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<emphasis box="[140,328,624,651]" inLineHeading="true" pageId="11" pageNumber="12" reason="1">Eutrichodesmus</emphasis>
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aster
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</emphasis>
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Golovatch, Geoffroy, Mauriès & VandenSpiegel
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</taxonomicName>
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,
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<taxonomicNameLabel box="[978,1043,624,650]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="11" pageNumber="12" type="description">
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||
<paragraph blockId="11.[140,1043,623,721]" box="[140,974,659,686]" pageId="11" pageNumber="12">
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||
<uri box="[140,974,659,686]" pageId="11" pageNumber="12">
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||
urn:lsid:zoobank.org:act:
|
||
<uuid box="[424,974,659,686]" pageId="11" pageNumber="12">B107B0B1-7423-466C-8E47-BE50F0EA0D5D</uuid>
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||
</uri>
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||
</paragraph>
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||
<paragraph blockId="11.[140,1043,623,721]" box="[140,245,695,721]" pageId="11" pageNumber="12">
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<figureCitation box="[140,245,695,721]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStart-2="Figure 11" captionStartId-0="11.[140,212,1622,1644]" captionStartId-1="12.[140,212,1558,1580]" captionStartId-2="14.[140,212,1622,1644]" captionTargetBox-0="[140,1108,865,1602]" captionTargetBox-1="[177,1070,165,1538]" captionTargetBox-2="[194,1057,699,1596]" captionTargetId-0="figure@11.[140,1108,865,1605]" captionTargetId-1="figure@12.[177,1070,165,1539]" captionTargetId-2="figure@14.[186,1062,681,1603]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="14" captionText-0="Figure 8. Eutrichodesmus aster sp. n., ♁ paratype from Tham Han Cave; habitus, lateral view. (Photographed not to scale by L. Deharveng)." captionText-1="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." captionText-2="Figure 11. Eutrichodesmus aster sp. n., ♁ paratype from Tham Han Cave; A, B, left gonopod, mesal and lateral views, respectively. – Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3769985" figureDoi-1="http://doi.org/10.5281/zenodo.3769987" figureDoi-2="http://doi.org/10.5281/zenodo.3813113" httpUri-0="https://zenodo.org/record/3769985/files/figure.png" httpUri-1="https://zenodo.org/record/3769987/files/figure.png" httpUri-2="https://zenodo.org/record/3813113/files/figure.png" pageId="11" pageNumber="12">Figs 8-11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="materials_examined">
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<paragraph blockId="11.[140,1107,765,827]" pageId="11" pageNumber="12">
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<materialsCitation ID-GBIF-Occurrence="2610648310" collectingDate="2003-12-20" collectorName="L. Deharveng & Deharveng, A & Phuong" country="Vietnam" county="Nghia Lo" location="Tham Han Cave" municipality="Xa Som" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Yen Bai" typeStatus="holotype">
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<emphasis bold="true" box="[140,312,765,791]" pageId="11" pageNumber="12">
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<typeStatus box="[140,198,765,791]" pageId="11" pageNumber="12">Type</typeStatus>
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material:
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</emphasis>
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<collectingCountry box="[319,420,765,791]" name="Vietnam" pageId="11" pageNumber="12">Vietnam</collectingCountry>
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,
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<collectingRegion box="[432,586,765,792]" country="Vietnam" name="Yen Bai" pageId="11" pageNumber="12">Yen Bai Prov.</collectingRegion>
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,
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<collectingCounty box="[597,710,765,792]" pageId="11" pageNumber="12">Nghia Lo</collectingCounty>
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:
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<collectingMunicipality box="[721,811,765,792]" pageId="11" pageNumber="12">Xa Som</collectingMunicipality>
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a,
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<location LSID="urn:lsid:plazi:treatment:936787D7FFB8FFBB8ABB4727FE5DFBDD:1E11601AFFB8FFBF897947AAFBC8FC9E" box="[846,1038,765,792]" country="Vietnam" county="Nghia Lo" municipality="Xa Som" name="Tham Han Cave" pageId="11" pageNumber="12" stateProvince="Yen Bai">Tham Han Cave</location>
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, hand collected,
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<date box="[254,398,800,826]" pageId="11" pageNumber="12" value="2003-12-20">
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<collectingDate box="[254,398,800,826]" pageId="11" pageNumber="12" value="2003-12-20">20.XII.2003</collectingDate>
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</date>
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, leg.
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<collectorName box="[457,615,800,827]" pageId="11" pageNumber="12">
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L.
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<collectorName box="[488,649,800,826]" pageId="11" pageNumber="12">Deharveng, A</collectorName>
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</collectorName>
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. Bedos and
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<collectorName box="[784,873,800,826]" pageId="11" pageNumber="12">Phuong</collectorName>
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(Vn0312-46), holo-
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</materialsCitation>
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3769985" ID-Zenodo-Dep="3769985" httpUri="https://zenodo.org/record/3769985/files/figure.png" pageId="11" pageNumber="12" startId="11.[140,212,1622,1644]" targetBox="[140,1108,865,1602]" targetPageId="11">
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<paragraph blockId="11.[140,1107,1622,1676]" pageId="11" pageNumber="12">
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<emphasis bold="true" box="[140,240,1622,1644]" pageId="11" pageNumber="12">Figure 8.</emphasis>
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<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[247,431,1622,1644]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aster" status="sp. n.">
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<emphasis box="[247,431,1622,1644]" italics="true" pageId="11" pageNumber="12">Eutrichodesmus aster</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[439,492,1623,1644]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
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, ♁ paratype from Tham Han Cave; habitus, lateral view. (Photographed not to scale by L. Deharveng).
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3769987" ID-Zenodo-Dep="3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="12" pageNumber="13" startId="12.[140,212,1558,1580]" targetBox="[177,1070,165,1538]" targetPageId="12">
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||
<paragraph blockId="12.[140,1108,1558,1676]" pageId="12" pageNumber="13">
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||
<emphasis bold="true" box="[140,238,1558,1580]" pageId="12" pageNumber="13">Figure 9.</emphasis>
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||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[244,428,1558,1580]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="aster" status="sp. n.">
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<emphasis box="[244,428,1558,1580]" italics="true" pageId="12" pageNumber="13">Eutrichodesmus aster</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[434,486,1559,1580]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
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, subadult paratype from Tham Han Cave;
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<emphasis bold="true" box="[886,905,1558,1580]" pageId="12" pageNumber="13">A</emphasis>
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, habitus, lateral view;
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<emphasis bold="true" box="[140,157,1590,1612]" pageId="12" pageNumber="13">B</emphasis>
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, anterior part of body, lateral view;
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<emphasis bold="true" box="[483,502,1590,1612]" pageId="12" pageNumber="13">C</emphasis>
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, midbody segments, lateral view;
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<emphasis bold="true" box="[811,830,1590,1612]" pageId="12" pageNumber="13">D</emphasis>
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, posterior part of body, lateral view;
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<emphasis bold="true" box="[194,209,1622,1644]" pageId="12" pageNumber="13">E</emphasis>
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, metatergal seta, sublateral view;
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<emphasis bold="true" box="[518,532,1622,1644]" pageId="12" pageNumber="13">F</emphasis>
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, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm.
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3813111" ID-Zenodo-Dep="3813111" httpUri="https://zenodo.org/record/3813111/files/figure.png" pageId="13" pageNumber="14" startId="13.[140,212,1558,1580]" targetBox="[177,1070,165,1538]" targetPageId="13">
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<paragraph blockId="13.[140,1108,1558,1676]" pageId="13" pageNumber="14">
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||
<emphasis bold="true" box="[140,253,1558,1580]" pageId="13" pageNumber="14">
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<figureCitation box="[140,246,1558,1580]" captionStart="Figure 12" captionStartId="16.[140,212,1064,1086]" captionTargetBox="[140,1108,165,1044]" captionTargetId="figure@16.[140,1108,165,1045]" captionTargetPageId="16" captionText="Figure 12. Eutrichodesmus filisetiger sp. n., ♁ paratype from Hang Doi Cave; habitus, lateral view. (Photographed not to scale by L. Deharveng)." figureDoi="http://doi.org/10.5281/zenodo.3813115" httpUri="https://zenodo.org/record/3813115/files/figure.png" pageId="13" pageNumber="14">Figure 10</figureCitation>
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.
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</emphasis>
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<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[261,446,1558,1580]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aster" status="sp. n.">
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<emphasis box="[261,446,1558,1580]" italics="true" pageId="13" pageNumber="14">Eutrichodesmus aster</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[455,508,1559,1580]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
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||
, subadult
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<typeStatus box="[608,687,1559,1580]" pageId="13" pageNumber="14">paratype</typeStatus>
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||
from Tham Han Cave;
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||
<emphasis bold="true" box="[919,966,1558,1580]" pageId="13" pageNumber="14">A, E</emphasis>
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||
, midbody segments, dorsal and ventral views, respectively;
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||
<emphasis bold="true" box="[568,615,1590,1612]" pageId="13" pageNumber="14">B, C</emphasis>
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||
, posterior part of body, dorsal and caudal views, respectively;
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||
<emphasis bold="true" box="[240,259,1622,1644]" pageId="13" pageNumber="14">D</emphasis>
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||
, anterior part of body, ventral view;
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||
<emphasis bold="true" box="[597,611,1622,1644]" pageId="13" pageNumber="14">F</emphasis>
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||
, cross-section of a midbody segment, caudal view;
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||
<emphasis bold="true" box="[1082,1102,1622,1644]" pageId="13" pageNumber="14">G</emphasis>
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||
, texture of tegument, dorsal view. – Scale bars: A, B, F,
|
||
<quantity box="[643,719,1654,1676]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="mm" value="0.5">0.5 mm</quantity>
|
||
; C-E,
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||
<quantity box="[779,855,1655,1676]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="mm" value="0.2">0.2 mm</quantity>
|
||
; G,
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||
<quantity box="[895,984,1654,1676]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="mm" value="0.05">0.05 mm</quantity>
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.
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||
</paragraph>
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</caption>
|
||
<paragraph blockId="14.[140,1108,167,616]" pageId="14" pageNumber="15">
|
||
type # (MNHN JC 319),
|
||
<typeStatus box="[442,552,167,193]" pageId="14" pageNumber="15">paratypes</typeStatus>
|
||
: 1 ♁,
|
||
<specimenCount box="[625,663,167,193]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount box="[686,786,167,193]" pageId="14" pageNumber="15" type="generic">2 subad.</specimenCount>
|
||
,
|
||
<specimenCount box="[798,863,167,193]" pageId="14" pageNumber="15" type="generic">1 juv.</specimenCount>
|
||
(MNHN JC 319), 1 ♁,
|
||
<specimenCount box="[177,273,202,228]" pageId="14" pageNumber="15" type="generic">1 subad.</specimenCount>
|
||
(ZMUM),
|
||
<specimenCount box="[409,505,202,228]" pageId="14" pageNumber="15" type="generic">1 subad.</specimenCount>
|
||
(SEM).
|
||
<materialsCitation ID-GBIF-Occurrence="2610648309" collectingDate="2003-12-19" collectorName="L. Deharveng & Deharveng, A & Phuong" country="Vietnam" location="Tham Le Cave" municipality="Nghia Lo" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Yen Bai" typeStatus="paratype">
|
||
<collectingCountry box="[604,705,202,228]" name="Vietnam" pageId="14" pageNumber="15">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion box="[717,870,202,228]" country="Vietnam" name="Yen Bai" pageId="14" pageNumber="15">Yen Bai Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality box="[882,994,202,228]" pageId="14" pageNumber="15">Nghia Lo</collectingMunicipality>
|
||
:
|
||
<location LSID="urn:lsid:plazi:treatment:936787D7FFB8FFBB8ABB4727FE5DFBDD:1E11601AFFBDFFBA89D9459DFF01FE8E" country="Vietnam" municipality="Nghia Lo" name="Tham Le Cave" pageId="14" pageNumber="15" stateProvince="Yen Bai">Tham Lê Cave</location>
|
||
, hand collected,
|
||
<collectingDate box="[384,527,237,263]" pageId="14" pageNumber="15" value="2003-12-19">
|
||
<date box="[384,527,237,263]" pageId="14" pageNumber="15" value="2003-12-19">19.XII.2003</date>
|
||
</collectingDate>
|
||
, leg.
|
||
<collectorName box="[583,739,237,264]" pageId="14" pageNumber="15">
|
||
L.
|
||
<collectorName box="[612,770,237,263]" pageId="14" pageNumber="15">Deharveng, A</collectorName>
|
||
</collectorName>
|
||
. Bedos and
|
||
<collectorName box="[900,989,237,263]" pageId="14" pageNumber="15">Phuong</collectorName>
|
||
(Vn0312- 38),
|
||
<typeStatus box="[193,303,273,299]" pageId="14" pageNumber="15">paratypes</typeStatus>
|
||
: 2 ♁ (MNHN JC 319)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation ID-GBIF-Occurrence="2610648308" collectingDate="2003-12-20" collectorName="L. Deharveng & Deharveng, A & Phuong" country="Vietnam" location="Hang Dan Khao Cave" municipality="Nghia Lo" pageId="14" pageNumber="15" specimenCount="3" specimenCount-female="2" stateProvince="Yen Bai" typeStatus="paratype">
|
||
<collectingCountry box="[586,687,272,298]" name="Vietnam" pageId="14" pageNumber="15">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion box="[698,851,272,299]" country="Vietnam" name="Yen Bai" pageId="14" pageNumber="15">Yen Bai Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality box="[863,975,272,299]" pageId="14" pageNumber="15">Nghia Lo</collectingMunicipality>
|
||
:
|
||
<location LSID="urn:lsid:plazi:treatment:936787D7FFB8FFBB8ABB4727FE5DFBDD:1E11601AFFBDFFBA89ED4446FEC8FEC7" country="Vietnam" municipality="Nghia Lo" name="Hang Dan Khao Cave" pageId="14" pageNumber="15" stateProvince="Yen Bai">Hang Dan Khao Cave</location>
|
||
, hand collected,
|
||
<date box="[469,613,308,334]" pageId="14" pageNumber="15" value="2003-12-20">
|
||
<collectingDate box="[469,613,308,334]" pageId="14" pageNumber="15" value="2003-12-20">20.XII.2003</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[677,838,307,334]" pageId="14" pageNumber="15">
|
||
L.
|
||
<collectorName box="[711,875,307,334]" pageId="14" pageNumber="15">Deharveng, A</collectorName>
|
||
</collectorName>
|
||
. Bedos and
|
||
<collectorName box="[1019,1108,307,333]" pageId="14" pageNumber="15">Phuong</collectorName>
|
||
(Vn0312-48), 1 ♁ (fragm.),
|
||
<specimenCount box="[465,502,343,369]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
|
||
(MNHN JC 319),
|
||
<specimenCount box="[735,772,343,369]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount box="[794,857,343,369]" pageId="14" pageNumber="15" type="generic">1 juv.</specimenCount>
|
||
(ZMUM)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[185,1021,378,404]" pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph blockId="14.[140,1108,167,616]" box="[185,1021,378,404]" pageId="14" pageNumber="15">
|
||
<emphasis bold="true" box="[185,263,378,404]" pageId="14" pageNumber="15">
|
||
<emphasis box="[185,258,378,404]" inLineHeading="true" pageId="14" pageNumber="15" reason="1">Name</emphasis>
|
||
:
|
||
</emphasis>
|
||
To refl ect the star-shaped and discoid body of the volvated animal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="15" type="diagnosis">
|
||
<paragraph blockId="14.[140,1108,167,616]" pageId="14" pageNumber="15">
|
||
<emphasis bold="true" box="[185,312,413,439]" pageId="14" pageNumber="15">
|
||
<emphasis box="[185,307,413,439]" inLineHeading="true" pageId="14" pageNumber="15" reason="1">Diagnosis</emphasis>
|
||
:
|
||
</emphasis>
|
||
Differs from all congeners, except
|
||
<taxonomicName baseAuthorityName="Hoffman" baseAuthorityYear="1977" box="[724,862,413,439]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="macclurei">
|
||
<emphasis box="[724,862,413,439]" italics="true" pageId="14" pageNumber="15">E. macclurei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="Hoffman" baseAuthorityYear="1977" box="[925,1062,413,439]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="reclinatus">
|
||
<emphasis box="[925,1062,413,439]" italics="true" pageId="14" pageNumber="15">E. reclinatus</emphasis>
|
||
</taxonomicName>
|
||
, by the extremely high mid-dorsal crests on metaterga 5-19. In addition, it can be distinguished from
|
||
<taxonomicName baseAuthorityName="Hoffman" baseAuthorityYear="1977" box="[298,433,483,509]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="macclurei">
|
||
<emphasis box="[298,433,483,509]" italics="true" pageId="14" pageNumber="15">E. macclurei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="Hoffman" baseAuthorityYear="1977" box="[491,626,483,509]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="reclinatus">
|
||
<emphasis box="[491,626,483,509]" italics="true" pageId="14" pageNumber="15">E. reclinatus</emphasis>
|
||
</taxonomicName>
|
||
in the presence of a mid-dorsal projection on metatergum 4, the slightly more strongly declivous and quadrilobate paraterga. The new species differs from all other species of the genus in minor details of gonopod structure (in particular, the shape of the telopodite and distofemoral process).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3813113" ID-Zenodo-Dep="3813113" httpUri="https://zenodo.org/record/3813113/files/figure.png" pageId="14" pageNumber="15" startId="14.[140,212,1622,1644]" targetBox="[194,1057,699,1596]" targetPageId="14">
|
||
<paragraph blockId="14.[140,1108,1622,1676]" pageId="14" pageNumber="15">
|
||
<emphasis bold="true" box="[140,250,1622,1644]" pageId="14" pageNumber="15">Figure 11.</emphasis>
|
||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[256,439,1622,1644]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aster" status="sp. n.">
|
||
<emphasis box="[256,439,1622,1644]" italics="true" pageId="14" pageNumber="15">Eutrichodesmus aster</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[445,496,1623,1644]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
|
||
, ♁
|
||
<typeStatus box="[529,608,1623,1644]" pageId="14" pageNumber="15">paratype</typeStatus>
|
||
from Tham Han Cave;
|
||
<emphasis bold="true" box="[829,875,1622,1644]" pageId="14" pageNumber="15">A, B</emphasis>
|
||
, left gonopod, mesal and lateral views, respectively. – Scale bar:
|
||
<quantity box="[492,569,1655,1676]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="mm" value="0.2">0.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="15" pageNumber="16" type="description">
|
||
<paragraph blockId="15.[140,1108,166,1108]" pageId="15" pageNumber="16">
|
||
<emphasis bold="true" box="[185,335,166,192]" pageId="15" pageNumber="16">
|
||
<emphasis box="[185,330,166,192]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Description</emphasis>
|
||
:
|
||
</emphasis>
|
||
Length of adults of both sexes ca
|
||
<quantity box="[742,870,167,193]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="15" pageNumber="16" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12-14 mm</quantity>
|
||
, width
|
||
<quantity box="[961,1103,167,193]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="15" pageNumber="16" unit="mm" value="2.2" valueMax="2.3" valueMin="2.1">2.1-2.3 mm</quantity>
|
||
, body broadest at segment 3 or 4.
|
||
<typeStatus box="[515,622,202,228]" pageId="15" pageNumber="16">Holotype</typeStatus>
|
||
ca
|
||
<quantity box="[660,743,202,228]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="16" unit="mm" value="12.0">12 mm</quantity>
|
||
long and
|
||
<quantity box="[856,945,202,228]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="16" unit="mm" value="2.2">2.2 mm</quantity>
|
||
wide. Coloration uniformly pallid, shown pinkish because of a photographic artifact (
|
||
<figureCitation box="[961,1028,237,263]" captionStart="Figure 8" captionStartId="11.[140,212,1622,1644]" captionTargetBox="[140,1108,865,1602]" captionTargetId="figure@11.[140,1108,865,1605]" captionTargetPageId="11" captionText="Figure 8. Eutrichodesmus aster sp. n., ♁ paratype from Tham Han Cave; habitus, lateral view. (Photographed not to scale by L. Deharveng)." figureDoi="http://doi.org/10.5281/zenodo.3769985" httpUri="https://zenodo.org/record/3769985/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="15.[140,1108,166,1108]" pageId="15" pageNumber="16">
|
||
Adults with 20 segments (
|
||
<figureCitation box="[487,560,272,298]" captionStart="Figure 8" captionStartId="11.[140,212,1622,1644]" captionTargetBox="[140,1108,865,1602]" captionTargetId="figure@11.[140,1108,865,1605]" captionTargetPageId="11" captionText="Figure 8. Eutrichodesmus aster sp. n., ♁ paratype from Tham Han Cave; habitus, lateral view. (Photographed not to scale by L. Deharveng)." figureDoi="http://doi.org/10.5281/zenodo.3769985" httpUri="https://zenodo.org/record/3769985/files/figure.png" pageId="15" pageNumber="16">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation box="[573,611,272,298]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="15" pageNumber="16">9A</figureCitation>
|
||
); body discoid, strongly flattened laterally; pattern of conglobation typical of “doratodesmids”. Head, antennae (Fig. 10D), tegument (Fig. 10G), sterna (Fig. 10E), gonopod aperture and many other characters much as in
|
||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[332,472,378,404]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="distinctus" status="sp. n.">
|
||
<emphasis box="[332,472,378,404]" italics="true" pageId="15" pageNumber="16">E. distinctus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[481,548,378,404]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
|
||
; collum rather irregularly tuberculate and only very slightly flattened medially at anterior edge (
|
||
<figureCitation box="[703,792,413,439]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="15" pageNumber="16">Fig. 9F</figureCitation>
|
||
). Metaterga behind collum with three transverse, rather irregular and mixostictic rows of rounded, flat, only slightly differentiated tubercles (
|
||
<figureCitation box="[513,582,483,510]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="15" pageNumber="16">Figs 9</figureCitation>
|
||
A-D), each crowned with a short, 2-segment- ed, flattened seta (
|
||
<figureCitation box="[361,454,519,545]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="15" pageNumber="16">Fig. 9E</figureCitation>
|
||
). Metaterga 4-19 each with a very high, slightly bifid, mid-dorsal process (
|
||
<figureCitation box="[377,453,554,580]" captionStart="Figure 8" captionStartId="11.[140,212,1622,1644]" captionTargetBox="[140,1108,865,1602]" captionTargetId="figure@11.[140,1108,865,1605]" captionTargetPageId="11" captionText="Figure 8. Eutrichodesmus aster sp. n., ♁ paratype from Tham Han Cave; habitus, lateral view. (Photographed not to scale by L. Deharveng)." figureDoi="http://doi.org/10.5281/zenodo.3769985" httpUri="https://zenodo.org/record/3769985/files/figure.png" pageId="15" pageNumber="16">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation box="[468,482,554,580]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="15" pageNumber="16">9</figureCitation>
|
||
A-D, 10A-C). Paraterga directed ventrolaterad up to subvertical, rather broad, surpassing level of venter (Fig. 10F); paraterga 2 strongly enlarged, very indistinctly lobulate anterolaterally (
|
||
<figureCitation box="[754,847,624,650]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="15" pageNumber="16">Fig. 9B</figureCitation>
|
||
). Following paraterga broadly rounded, evidently 4-lobate laterally and bilobate caudolaterally (
|
||
<figureCitation box="[1007,1077,659,686]" captionStart="Figure 9" captionStartId="12.[140,212,1558,1580]" captionTargetBox="[177,1070,165,1538]" captionTargetId="figure@12.[177,1070,165,1539]" captionTargetPageId="12" captionText="Figure 9. Eutrichodesmus aster sp. n., subadult paratype from Tham Han Cave; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, metatergal seta, sublateral view; F, anterior part of body, frontodorsal view. – Scale bars: A-D, F, 0.5 mm; E, 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3769987" httpUri="https://zenodo.org/record/3769987/files/figure.png" pageId="15" pageNumber="16">Figs 9</figureCitation>
|
||
A- D, 10E). Limbus strongly denticulate, almost hidden by nearby abundant microvilli (Fig. 10G). Pore formula normal, ozopores poorly visible, located dorsally near base of ventrocaudal lobulation (Figs 10A, B). Epiproct short, with differentiated tubercles (Fig. 10C).
|
||
</paragraph>
|
||
<paragraph blockId="15.[140,1108,166,1108]" box="[185,983,835,862]" pageId="15" pageNumber="16">Legs relatively long, slightly surpassing edge of paraterga (Figs 10E, F).</paragraph>
|
||
<paragraph blockId="15.[140,1108,166,1108]" pageId="15" pageNumber="16">
|
||
Gonopods (
|
||
<figureCitation box="[318,398,871,897]" captionStart="Figure 11" captionStartId="14.[140,212,1622,1644]" captionTargetBox="[194,1057,699,1596]" captionTargetId="figure@14.[186,1062,681,1603]" captionTargetPageId="14" captionText="Figure 11. Eutrichodesmus aster sp. n., ♁ paratype from Tham Han Cave; A, B, left gonopod, mesal and lateral views, respectively. – Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3813113" httpUri="https://zenodo.org/record/3813113/files/figure.png" pageId="15" pageNumber="16">Fig. 11</figureCitation>
|
||
) relatively simple. Coxae abundantly micropapillate and setose, with one normal, apicolateral lobe. Telopodite elongate, slender, slightly arcuate, with a short, bare, distofemoral process (dp) at about proximal one-third; seminal groove and acropodite (= solenomere) long, hairpad subapical.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="15" pageNumber="16" type="discussion">
|
||
<paragraph blockId="15.[140,1108,166,1108]" pageId="15" pageNumber="16">
|
||
<emphasis bold="true" box="[185,298,1011,1037]" pageId="15" pageNumber="16">
|
||
<emphasis box="[185,293,1011,1037]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Remarks</emphasis>
|
||
:
|
||
</emphasis>
|
||
This pallid species shows peculiarly high mid-dorsal projections, coupled with short, distally flattened tergal setae. It is also a typical “doratodesmid”, possibly another troglobite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |