treatments-xml/data/93/67/87/936787D7FFAFFF948ABB442EFEDEFC2D.xml

<document ID-DOI="10.3897/zookeys.12.167" ID-GBIF-Dataset="3026c846-a607-4826-b2a9-8f7a531baec5" ID-GBIF-Taxon="163971733" ID-ISSN="1313–2970" ID-Zenodo-Dep="576460" ID-ZooBank="B32EF8D5-C3C6-4C8C-8845-284125448425" LSID="urn:lsid:zoobank.org:act:E7EB907C-AA5B-4612-B292-403142A55D5B" approvalRequired="25" approvalRequired_for_textStreams="25" checkinTime="1587995767728" checkinUser="plazi" docAuthor="Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; VandenSpiegel, Didier" docDate="2009" docId="936787D7FFAFFF948ABB442EFEDEFC2D" docLanguage="en" docName="ZK_article_1994.pdf" docOrigin="ZooKeys 12 (12)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Eutrichodesmus griseus Golovatch, Geoffroy, Mauries &amp; VandenSpiegel 2009, sp. n." docType="treatment" docUuid="E7EB907C-AA5B-4612-B292-403142A55D5B" docUuidSource="ZooBank" docVersion="3" lastPageId="32" lastPageNumber="33" masterDocId="6F5EFFAFFFB3FFB48A374557FFC6FF89" masterDocTitle="Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species" masterLastPageNumber="46" masterPageNumber="1" pageId="28" pageNumber="29" updateTime="1643565021146" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei</mods:namePart>
<mods:affiliation>Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Ecologie &amp; Gestion de la Biodiversité, UMR 7179 du CNRS, Equipe EVOLTRAIT, Brunoy, France &amp; D 362 Dc 0 - &amp; CC</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mauriès, Jean-Paul</mods:namePart>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Systématique et Evolution, Section Arthropodes, Paris, France &amp; - &amp; - 42 F &amp; D 01 - 28 Ad- 43 F</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>VandenSpiegel, Didier</mods:namePart>
<mods:affiliation>Musée Royal de l’Afrique centrale, Tervuren, Belgium &amp; A &amp; EE &amp; F- 04802 E 68 Cb</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-06-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>12</mods:number>
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<mods:number>12</mods:number>
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<mods:identifier type="DOI">10.3897/zookeys.12.167</mods:identifier>
<mods:identifier type="GBIF-Dataset">3026c846-a607-4826-b2a9-8f7a531baec5</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576460</mods:identifier>
<mods:identifier type="ZooBank">B32EF8D5-C3C6-4C8C-8845-284125448425</mods:identifier>
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<treatment ID-GBIF-Taxon="163971733" LSID="urn:lsid:zoobank.org:act:E7EB907C-AA5B-4612-B292-403142A55D5B" httpUri="http://treatment.plazi.org/id/936787D7FFAFFF948ABB442EFEDEFC2D" lastPageId="32" lastPageNumber="33" pageId="28" pageNumber="29">
<subSubSection box="[140,1065,377,404]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph blockId="28.[140,1065,377,474]" box="[140,1065,377,404]" pageId="28" pageNumber="29">
<heading box="[140,1065,377,404]" level="2" pageId="28" pageNumber="29">
<emphasis bold="true" box="[140,1065,377,404]" pageId="28" pageNumber="29">
<taxonomicName authority="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityName="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityYear="2009" box="[140,988,377,404]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="griseus" status="sp. n.">
<emphasis bold="true" box="[140,414,377,404]" italics="true" pageId="28" pageNumber="29">
<emphasis box="[140,328,377,404]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">Eutrichodesmus</emphasis>
griseus
</emphasis>
Golovatch, Geoffroy, Mauriès &amp; VandenSpiegel
</taxonomicName>
, 
<taxonomicNameLabel box="[1000,1065,378,404]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="description">
<paragraph blockId="28.[140,1065,377,474]" box="[140,970,413,440]" pageId="28" pageNumber="29">
<uri box="[140,970,413,440]" pageId="28" pageNumber="29">
urn:lsid:zoobank.org:act: 
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</uri>
</paragraph>
<paragraph blockId="28.[140,1065,377,474]" box="[140,261,448,474]" pageId="28" pageNumber="29">
<figureCitation box="[140,261,448,474]" captionStart-0="Figure 23" captionStart-1="Figure 24" captionStart-2="Figure 25" captionStart-3="Figure 26" captionStartId-0="28.[140,212,1622,1644]" captionStartId-1="29.[140,212,1394,1416]" captionStartId-2="30.[140,212,1507,1529]" captionStartId-3="31.[140,212,1590,1612]" captionTargetBox-0="[140,1108,938,1602]" captionTargetBox-1="[139,1107,165,1375]" captionTargetBox-2="[140,1108,165,1487]" captionTargetBox-3="[140,1108,512,1561]" captionTargetId-0="figure@28.[137,1111,938,1602]" captionTargetId-1="figure@29.[139,1108,165,1375]" captionTargetId-2="figure@30.[140,1109,165,1488]" captionTargetId-3="figure@31.[140,1108,511,1570]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionTargetPageId-2="30" captionTargetPageId-3="31" captionText-0="Figure 23. Eutrichodesmus griseus sp. n., ♀ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." captionText-1="Figure 24. Eutrichodesmus griseus sp. n., ♀ paratype; A, habitus, lateral view; B, F, anterior part of body, lateral and frontal views, respectively; C, G, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view; E, epiproct, dorsolateral view. – Scale bars: A, 0.5 mm; B-G, 0.1 mm." captionText-2="Figure 25. Eutrichodesmus griseus sp. n., ♀ paratype; A, D, posterior part of body, dorsal and ventral views, respectively; B, anterior part of body, ventral view; C, midbody segments, ventral view; E, crosssection of a midbody segment, caudal view; F, texture of tegument, dorsal view. – Scale bars: A-E, 0.1 mm; F, 0.01 mm." captionText-3="Figure 26. Eutrichodesmus griseus sp. n., paratypes ♀ (A, B) and ♁ (C, D, E); A, limbus, dorsal view; B, metatergal seta, sublateral view; C-E, left gonopod, mesal, lateral and subfrontal views, respectively. – Scale bars: A, 0.01 mm; B, 0.005 mm; C-E, 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3769993" figureDoi-1="http://doi.org/10.5281/zenodo.3769995" figureDoi-2="http://doi.org/10.5281/zenodo.3769997" figureDoi-3="http://doi.org/10.5281/zenodo.3769999" httpUri-0="https://zenodo.org/record/3769993/files/figure.png" httpUri-1="https://zenodo.org/record/3769995/files/figure.png" httpUri-2="https://zenodo.org/record/3769997/files/figure.png" httpUri-3="https://zenodo.org/record/3769999/files/figure.png" pageId="28" pageNumber="29">Figs 23-26</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="materials_examined">
<paragraph blockId="28.[140,1108,518,862]" pageId="28" pageNumber="29">
<emphasis bold="true" box="[140,317,518,545]" pageId="28" pageNumber="29">
<emphasis box="[140,198,519,545]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">Type</emphasis>
material:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2610648303" collectingDate="2003-01-27" collectorName="L. Deharveng &amp; A. Bedos" country="Vietnam" county="Kien Luong" location="Nui Hon Chong" municipality="Hon Chong" pageId="28" pageNumber="29" specimenCount="2" specimenCount-female="2" stateProvince="Kien Giang" typeStatus="holotype">
<collectingCountry box="[329,430,519,545]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
, 
<collectingRegion box="[447,656,519,545]" country="Vietnam" name="Kien Giang" pageId="28" pageNumber="29">Kien Giang Prov.</collectingRegion>
, 
<collectingCounty box="[673,818,519,545]" pageId="28" pageNumber="29">Kien Luong</collectingCounty>
: 
<collectingMunicipality box="[833,980,519,545]" pageId="28" pageNumber="29">Hon Chong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:936787D7FFAFFF948ABB442EFEDEFC2D:1E11601AFFAFFFA889D24750FF1BFDCD" country="Vietnam" county="Kien Luong" municipality="Hon Chong" name="Nui Hon Chong" pageId="28" pageNumber="29" stateProvince="Kien Giang">Nui Hon Chong</location>
, outside 
<location LSID="urn:lsid:plazi:treatment:936787D7FFAFFF948ABB442EFEDEFC2D:1E11601AFFAFFFA88B77477DFEBEFDCD" box="[320,376,554,580]" country="Vietnam" county="Kien Luong" municipality="Hon Chong" name="Cave" pageId="28" pageNumber="29" stateProvince="Kien Giang">Cave</location>
2 near 
<location LSID="urn:lsid:plazi:treatment:936787D7FFAFFF948ABB442EFEDEFC2D:1E11601AFFAFFFA88BFC477DFDB6FDCD" box="[459,624,554,580]" country="Vietnam" county="Kien Luong" municipality="Hon Chong" name="Hang Hai Cot" pageId="28" pageNumber="29" stateProvince="Kien Giang">Hang Hai Côt</location>
, grasses, soil, 
<location LSID="urn:lsid:plazi:treatment:936787D7FFAFFF948ABB442EFEDEFC2D:1E11601AFFAFFFA8893F477DFC91FDCD" box="[776,855,554,580]" country="Vietnam" county="Kien Luong" municipality="Hon Chong" name="Berlese" pageId="28" pageNumber="29" stateProvince="Kien Giang">Berlese</location>
extraction, 
<date box="[988,1103,554,580]" pageId="28" pageNumber="29" value="2003-01-27">
<collectingDate box="[988,1103,554,580]" pageId="28" pageNumber="29" value="2003-01-27">27.I.2003</collectingDate>
</date>
, leg. 
<collectorName box="[190,348,589,616]" pageId="28" pageNumber="29">L. Deharveng</collectorName>
and 
<collectorName box="[411,515,589,615]" pageId="28" pageNumber="29">A. Bedos</collectorName>
(VIET-923), 
<typeStatus box="[679,778,589,615]" pageId="28" pageNumber="29">holotype</typeStatus>
♁ (MNHN JC 323), 
<typeStatus pageId="28" pageNumber="29">paratypes</typeStatus>
: 1 ♁, 
<specimenCount box="[271,308,624,651]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
(MNHN JC 323), 
<specimenCount box="[541,578,624,651]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
(SEM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="etymology">
<paragraph blockId="28.[140,1108,518,862]" pageId="28" pageNumber="29">
<emphasis bold="true" box="[185,263,659,685]" pageId="28" pageNumber="29">
<emphasis box="[185,258,659,685]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">Name</emphasis>
:
</emphasis>
To emphasize the greyish coloration of the vertex, collum, metaterga and most of the telson.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="diagnosis">
<paragraph blockId="28.[140,1108,518,862]" pageId="28" pageNumber="29">
<emphasis bold="true" box="[185,312,729,755]" pageId="28" pageNumber="29">
<emphasis box="[185,307,729,755]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">Diagnosis</emphasis>
:
</emphasis>
Differs from congeners by the peculiar, grey to blackish coloration, coupled with the relatively short and high paraterga (thus the body apparently not capable of complete volvation) and a few minor details of gonopod structure (in particular, the shape of the telopodite and distofemoral process).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769993" ID-Zenodo-Dep="3769993" httpUri="https://zenodo.org/record/3769993/files/figure.png" pageId="28" pageNumber="29" startId="28.[140,212,1622,1644]" targetBox="[140,1108,938,1602]" targetPageId="28">
<paragraph blockId="28.[140,1107,1622,1676]" pageId="28" pageNumber="29">
<emphasis bold="true" box="[140,251,1622,1644]" pageId="28" pageNumber="29">Figure 23.</emphasis>
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[256,456,1622,1644]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="griseus" status="sp. n.">
<emphasis box="[256,456,1622,1644]" italics="true" pageId="28" pageNumber="29">
Eutrichodesmus 
<taxonomicName box="[399,456,1623,1644]" pageId="28" pageNumber="29" rank="species" species="griseus" status="sp. n.">griseus</taxonomicName>
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[462,513,1623,1644]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
, ♀ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769995" ID-Zenodo-Dep="3769995" httpUri="https://zenodo.org/record/3769995/files/figure.png" pageId="29" pageNumber="30" startId="29.[140,212,1394,1416]" targetBox="[139,1107,165,1375]" targetPageId="29">
<paragraph blockId="29.[140,1108,1394,1481]" pageId="29" pageNumber="30">
<emphasis bold="true" box="[140,250,1394,1416]" pageId="29" pageNumber="30">Figure 24.</emphasis>
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[255,455,1395,1416]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="griseus" status="sp. n.">
<emphasis box="[255,455,1395,1416]" italics="true" pageId="29" pageNumber="30">Eutrichodesmus griseus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[460,511,1395,1416]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
, ♀ paratype; 
<emphasis bold="true" box="[634,653,1394,1416]" pageId="29" pageNumber="30">A</emphasis>
, habitus, lateral view; 
<emphasis bold="true" box="[858,875,1394,1416]" pageId="29" pageNumber="30">B</emphasis>
, 
<emphasis bold="true" box="[887,901,1394,1416]" pageId="29" pageNumber="30">F</emphasis>
, anterior part of body, lateral and frontal views, respectively; 
<emphasis bold="true" box="[491,541,1426,1448]" pageId="29" pageNumber="30">C, G</emphasis>
, midbody segments, lateral and dorsal views, respectively; 
<emphasis bold="true" box="[1082,1101,1426,1448]" pageId="29" pageNumber="30">D</emphasis>
, posterior part of body, lateral view; 
<emphasis bold="true" box="[466,481,1458,1480]" pageId="29" pageNumber="30">E</emphasis>
, epiproct, dorsolateral view. – Scale bars: A, 0.5 mm; B-G, 0.1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769997" ID-Zenodo-Dep="3769997" httpUri="https://zenodo.org/record/3769997/files/figure.png" pageId="30" pageNumber="31" startId="30.[140,212,1507,1529]" targetBox="[140,1108,165,1487]" targetPageId="30">
<paragraph blockId="30.[140,1107,1507,1625]" pageId="30" pageNumber="31">
<emphasis bold="true" box="[140,253,1507,1529]" pageId="30" pageNumber="31">Figure 25.</emphasis>
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[260,463,1507,1529]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="griseus" status="sp. n.">
<emphasis box="[260,463,1507,1529]" italics="true" pageId="30" pageNumber="31">
<taxonomicName box="[260,397,1507,1528]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">Eutrichodesmus</taxonomicName>
griseus
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[470,523,1508,1529]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
, ♀ paratype; 
<emphasis bold="true" box="[654,673,1507,1529]" pageId="30" pageNumber="31">A</emphasis>
, 
<emphasis bold="true" box="[687,706,1507,1529]" pageId="30" pageNumber="31">D</emphasis>
, posterior part of body, dorsal and ventral views, respectively; 
<emphasis bold="true" box="[321,338,1539,1561]" pageId="30" pageNumber="31">B</emphasis>
, anterior part of body, ventral view; 
<emphasis bold="true" box="[683,702,1539,1561]" pageId="30" pageNumber="31">C</emphasis>
, midbody segments, ventral view; 
<emphasis bold="true" box="[1027,1042,1539,1561]" pageId="30" pageNumber="31">E</emphasis>
, crosssection of a midbody segment, caudal view; 
<emphasis bold="true" box="[559,573,1571,1593]" pageId="30" pageNumber="31">F</emphasis>
, texture of tegument, dorsal view. – Scale bars: A-E, 0.1 mm; F, 0.01 mm.
</paragraph>
</caption>
<subSubSection lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="description">
<paragraph blockId="31.[140,1108,166,475]" pageId="31" pageNumber="32">
<emphasis bold="true" box="[185,335,166,192]" pageId="31" pageNumber="32">
<emphasis box="[185,330,166,192]" inLineHeading="true" pageId="31" pageNumber="32" reason="1">Description</emphasis>
:
</emphasis>
Length of adults of both sexes ca 6.0- 
<quantity box="[765,858,166,193]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="31" pageNumber="32" unit="mm" value="6.5">6.5 mm</quantity>
, width 
<quantity box="[944,1103,166,193]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="31" pageNumber="32" unit="mm" value="0.725" valueMax="0.75" valueMin="0.7">0.7–0.75 mm</quantity>
, body broadest at segment 3 or 4. 
<typeStatus box="[512,619,202,228]" pageId="31" pageNumber="32">Holotype</typeStatus>
ca 
<quantity box="[656,745,202,229]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="32" unit="mm" value="4.5">4.5 mm</quantity>
long and 
<quantity box="[857,946,202,229]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="31" pageNumber="32" unit="mm" value="2.5">2.5 mm</quantity>
wide. Coloration of vertex, collum, following metaterga, and of dorsal and lateral parts of telson greyish to dark grey. Clypeolabral part of head, venter and legs contrastingly pallid (
<figureCitation box="[148,232,307,334]" captionStart="Figure 23" captionStartId="28.[140,212,1622,1644]" captionTargetBox="[140,1108,938,1602]" captionTargetId="figure@28.[137,1111,938,1602]" captionTargetPageId="28" captionText="Figure 23. Eutrichodesmus griseus sp. n., ♀ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." figureDoi="http://doi.org/10.5281/zenodo.3769993" httpUri="https://zenodo.org/record/3769993/files/figure.png" pageId="31" pageNumber="32">Fig. 23</figureCitation>
). 
<typeStatus box="[253,360,307,333]" pageId="31" pageNumber="32">Holotype</typeStatus>
light grey.
</paragraph>
<paragraph blockId="31.[140,1108,166,475]" lastBlockId="32.[140,1108,167,932]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
Adults with 20 segments (
<figureCitation box="[484,572,343,369]" captionStart="Figure 23" captionStartId="28.[140,212,1622,1644]" captionTargetBox="[140,1108,938,1602]" captionTargetId="figure@28.[137,1111,938,1602]" captionTargetPageId="28" captionText="Figure 23. Eutrichodesmus griseus sp. n., ♀ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." figureDoi="http://doi.org/10.5281/zenodo.3769993" httpUri="https://zenodo.org/record/3769993/files/figure.png" pageId="31" pageNumber="32">Figs 23</figureCitation>
, 
<figureCitation box="[585,637,343,369]" captionStart="Figure 24" captionStartId="29.[140,212,1394,1416]" captionTargetBox="[139,1107,165,1375]" captionTargetId="figure@29.[139,1108,165,1375]" captionTargetPageId="29" captionText="Figure 24. Eutrichodesmus griseus sp. n., ♀ paratype; A, habitus, lateral view; B, F, anterior part of body, lateral and frontal views, respectively; C, G, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view; E, epiproct, dorsolateral view. – Scale bars: A, 0.5 mm; B-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769995" httpUri="https://zenodo.org/record/3769995/files/figure.png" pageId="31" pageNumber="32">24A</figureCitation>
), volvation apparently incomplete due to relatively short paraterga. Head (
<figureCitation box="[508,608,378,405]" captionStart="Figure 25" captionStartId="30.[140,212,1507,1529]" captionTargetBox="[140,1108,165,1487]" captionTargetId="figure@30.[140,1109,165,1488]" captionTargetPageId="30" captionText="Figure 25. Eutrichodesmus griseus sp. n., ♀ paratype; A, D, posterior part of body, dorsal and ventral views, respectively; B, anterior part of body, ventral view; C, midbody segments, ventral view; E, crosssection of a midbody segment, caudal view; F, texture of tegument, dorsal view. – Scale bars: A-E, 0.1 mm; F, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3769997" httpUri="https://zenodo.org/record/3769997/files/figure.png" pageId="31" pageNumber="32">Fig. 25B</figureCitation>
) with a poorly separated pair of paramedian tubercles above antennal sockets; antennae relatively short and clavate; tegument (
<figureCitation captionStart="Figure 25" captionStartId="30.[140,212,1507,1529]" captionTargetBox="[140,1108,165,1487]" captionTargetId="figure@30.[140,1109,165,1488]" captionTargetPageId="30" captionText="Figure 25. Eutrichodesmus griseus sp. n., ♀ paratype; A, D, posterior part of body, dorsal and ventral views, respectively; B, anterior part of body, ventral view; C, midbody segments, ventral view; E, crosssection of a midbody segment, caudal view; F, texture of tegument, dorsal view. – Scale bars: A-E, 0.1 mm; F, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3769997" httpUri="https://zenodo.org/record/3769997/files/figure.png" pageId="31" pageNumber="32">Fig. 25F</figureCitation>
) and many other characters (
<figureCitation box="[522,606,448,475]" captionStart="Figure 25" captionStartId="30.[140,212,1507,1529]" captionTargetBox="[140,1108,165,1487]" captionTargetId="figure@30.[140,1109,165,1488]" captionTargetPageId="30" captionText="Figure 25. Eutrichodesmus griseus sp. n., ♀ paratype; A, D, posterior part of body, dorsal and ventral views, respectively; B, anterior part of body, ventral view; C, midbody segments, ventral view; E, crosssection of a midbody segment, caudal view; F, texture of tegument, dorsal view. – Scale bars: A-E, 0.1 mm; F, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3769997" httpUri="https://zenodo.org/record/3769997/files/figure.png" pageId="31" pageNumber="32">Figs 25</figureCitation>
C-E) much as in 
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[807,941,448,474]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="distinctus" status="sp. n.">
<emphasis box="[807,941,448,474]" italics="true" pageId="31" pageNumber="32">E. distinctus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[950,1015,449,475]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
; collum rather regularly tuberculate, anterior row of tubercles especially dense and directed dorsofrontally, thus slightly resembling a pyrgodesmid condition (
<figureCitation box="[913,1094,202,228]" captionStart="Figure 24" captionStartId="29.[140,212,1394,1416]" captionTargetBox="[139,1107,165,1375]" captionTargetId="figure@29.[139,1108,165,1375]" captionTargetPageId="29" captionText="Figure 24. Eutrichodesmus griseus sp. n., ♀ paratype; A, habitus, lateral view; B, F, anterior part of body, lateral and frontal views, respectively; C, G, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view; E, epiproct, dorsolateral view. – Scale bars: A, 0.5 mm; B-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769995" httpUri="https://zenodo.org/record/3769995/files/figure.png" pageId="32" pageNumber="33">Figs 24A, B, F</figureCitation>
). Metaterga behind collum with three transverse, regular, nearly isostictic rows of round- ed, flat, undifferentiated tubercles (
<figureCitation box="[548,635,272,298]" captionStart="Figure 24" captionStartId="29.[140,212,1394,1416]" captionTargetBox="[139,1107,165,1375]" captionTargetId="figure@29.[139,1108,165,1375]" captionTargetPageId="29" captionText="Figure 24. Eutrichodesmus griseus sp. n., ♀ paratype; A, habitus, lateral view; B, F, anterior part of body, lateral and frontal views, respectively; C, G, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view; E, epiproct, dorsolateral view. – Scale bars: A, 0.5 mm; B-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769995" httpUri="https://zenodo.org/record/3769995/files/figure.png" pageId="32" pageNumber="33">Figs 24</figureCitation>
A-G), each crowned with a short, 2-segmented, conspicuously denticulate and slightly claviform seta (
<figureCitation box="[874,981,307,334]" captionStart="Figure 26" captionStartId="31.[140,212,1590,1612]" captionTargetBox="[140,1108,512,1561]" captionTargetId="figure@31.[140,1108,511,1570]" captionTargetPageId="31" captionText="Figure 26. Eutrichodesmus griseus sp. n., paratypes ♀ (A, B) and ♁ (C, D, E); A, limbus, dorsal view; B, metatergal seta, sublateral view; C-E, left gonopod, mesal, lateral and subfrontal views, respectively. – Scale bars: A, 0.01 mm; B, 0.005 mm; C-E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769999" httpUri="https://zenodo.org/record/3769999/files/figure.png" pageId="32" pageNumber="33">Fig. 26B</figureCitation>
). Paraterga directed ventrolaterad, short but rather broad, barely reaching level of venter, nearly continuing general outline of convex dorsum (
<figureCitation box="[685,772,378,404]" captionStart="Figure 24" captionStartId="29.[140,212,1394,1416]" captionTargetBox="[139,1107,165,1375]" captionTargetId="figure@29.[139,1108,165,1375]" captionTargetPageId="29" captionText="Figure 24. Eutrichodesmus griseus sp. n., ♀ paratype; A, habitus, lateral view; B, F, anterior part of body, lateral and frontal views, respectively; C, G, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view; E, epiproct, dorsolateral view. – Scale bars: A, 0.5 mm; B-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769995" httpUri="https://zenodo.org/record/3769995/files/figure.png" pageId="32" pageNumber="33">Figs 24</figureCitation>
A-D, 25E); paraterga 2 (
<figureCitation captionStart="Figure 24" captionStartId="29.[140,212,1394,1416]" captionTargetBox="[139,1107,165,1375]" captionTargetId="figure@29.[139,1108,165,1375]" captionTargetPageId="29" captionText="Figure 24. Eutrichodesmus griseus sp. n., ♀ paratype; A, habitus, lateral view; B, F, anterior part of body, lateral and frontal views, respectively; C, G, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view; E, epiproct, dorsolateral view. – Scale bars: A, 0.5 mm; B-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769995" httpUri="https://zenodo.org/record/3769995/files/figure.png" pageId="32" pageNumber="33">Fig. 24B</figureCitation>
) evidently lobulate only laterally. Following paraterga broadly rounded, evidently 3- to 5-lobate laterally, at least bilobate also caudolaterally (
<figureCitation box="[844,931,448,474]" captionStart="Figure 24" captionStartId="29.[140,212,1394,1416]" captionTargetBox="[139,1107,165,1375]" captionTargetId="figure@29.[139,1108,165,1375]" captionTargetPageId="29" captionText="Figure 24. Eutrichodesmus griseus sp. n., ♀ paratype; A, habitus, lateral view; B, F, anterior part of body, lateral and frontal views, respectively; C, G, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view; E, epiproct, dorsolateral view. – Scale bars: A, 0.5 mm; B-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769995" httpUri="https://zenodo.org/record/3769995/files/figure.png" pageId="32" pageNumber="33">Figs 24</figureCitation>
A-D, G). Limbus distinctly spinulate (
<figureCitation box="[415,518,483,510]" captionStart="Figure 26" captionStartId="31.[140,212,1590,1612]" captionTargetBox="[140,1108,512,1561]" captionTargetId="figure@31.[140,1108,511,1570]" captionTargetPageId="31" captionText="Figure 26. Eutrichodesmus griseus sp. n., paratypes ♀ (A, B) and ♁ (C, D, E); A, limbus, dorsal view; B, metatergal seta, sublateral view; C-E, left gonopod, mesal, lateral and subfrontal views, respectively. – Scale bars: A, 0.01 mm; B, 0.005 mm; C-E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769999" httpUri="https://zenodo.org/record/3769999/files/figure.png" pageId="32" pageNumber="33">Fig. 26A</figureCitation>
), almost hidden by nearby abundant microvilli (
<figureCitation captionStart="Figure 25" captionStartId="30.[140,212,1507,1529]" captionTargetBox="[140,1108,165,1487]" captionTargetId="figure@30.[140,1109,165,1488]" captionTargetPageId="30" captionText="Figure 25. Eutrichodesmus griseus sp. n., ♀ paratype; A, D, posterior part of body, dorsal and ventral views, respectively; B, anterior part of body, ventral view; C, midbody segments, ventral view; E, crosssection of a midbody segment, caudal view; F, texture of tegument, dorsal view. – Scale bars: A-E, 0.1 mm; F, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3769997" httpUri="https://zenodo.org/record/3769997/files/figure.png" pageId="32" pageNumber="33">Fig. 25F</figureCitation>
). Pore formula normal, ozopores located on top of porosteles (always caudalmost lateral lobulation) (
<figureCitation box="[355,438,554,580]" captionStart="Figure 24" captionStartId="29.[140,212,1394,1416]" captionTargetBox="[139,1107,165,1375]" captionTargetId="figure@29.[139,1108,165,1375]" captionTargetPageId="29" captionText="Figure 24. Eutrichodesmus griseus sp. n., ♀ paratype; A, habitus, lateral view; B, F, anterior part of body, lateral and frontal views, respectively; C, G, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view; E, epiproct, dorsolateral view. – Scale bars: A, 0.5 mm; B-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769995" httpUri="https://zenodo.org/record/3769995/files/figure.png" pageId="32" pageNumber="33">Figs 24</figureCitation>
A-D, G). Epiproct rather heavily tuberculate (
<figureCitation box="[954,1095,554,580]" captionStart="Figure 24" captionStartId="29.[140,212,1394,1416]" captionTargetBox="[139,1107,165,1375]" captionTargetId="figure@29.[139,1108,165,1375]" captionTargetPageId="29" captionText="Figure 24. Eutrichodesmus griseus sp. n., ♀ paratype; A, habitus, lateral view; B, F, anterior part of body, lateral and frontal views, respectively; C, G, midbody segments, lateral and dorsal views, respectively; D, posterior part of body, lateral view; E, epiproct, dorsolateral view. – Scale bars: A, 0.5 mm; B-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769995" httpUri="https://zenodo.org/record/3769995/files/figure.png" pageId="32" pageNumber="33">Figs 24D, E</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769999" ID-Zenodo-Dep="3769999" httpUri="https://zenodo.org/record/3769999/files/figure.png" pageId="31" pageNumber="32" startId="31.[140,212,1590,1612]" targetBox="[140,1108,512,1561]" targetPageId="31">
<paragraph blockId="31.[140,1108,1590,1676]" pageId="31" pageNumber="32">
<emphasis bold="true" box="[140,252,1590,1612]" pageId="31" pageNumber="32">Figure 26.</emphasis>
<taxonomicName authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[260,461,1590,1612]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="griseus" status="sp. n.">
<emphasis box="[260,461,1590,1612]" italics="true" pageId="31" pageNumber="32">Eutrichodesmus griseus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[468,521,1591,1612]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
, paratypes ♀ (A, B) and ♁ (C, D, E); 
<emphasis bold="true" box="[889,908,1590,1612]" pageId="31" pageNumber="32">A</emphasis>
, limbus, dorsal view; 
<emphasis bold="true" box="[140,157,1622,1644]" pageId="31" pageNumber="32">B</emphasis>
, metatergal seta, sublateral view; 
<emphasis bold="true" box="[466,508,1622,1644]" pageId="31" pageNumber="32">C-E</emphasis>
, left gonopod, mesal, lateral and subfrontal views, respectively. – Scale bars: A, 0.01 mm; B, 0.005 mm; C-E, 0.2 mm.
</paragraph>
</caption>
<paragraph blockId="32.[140,1108,167,932]" box="[185,1070,589,616]" pageId="32" pageNumber="33">
Legs relatively short and stout, barely reaching edge of paraterga (
<figureCitation box="[922,1007,589,616]" captionStart="Figure 25" captionStartId="30.[140,212,1507,1529]" captionTargetBox="[140,1108,165,1487]" captionTargetId="figure@30.[140,1109,165,1488]" captionTargetPageId="30" captionText="Figure 25. Eutrichodesmus griseus sp. n., ♀ paratype; A, D, posterior part of body, dorsal and ventral views, respectively; B, anterior part of body, ventral view; C, midbody segments, ventral view; E, crosssection of a midbody segment, caudal view; F, texture of tegument, dorsal view. – Scale bars: A-E, 0.1 mm; F, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3769997" httpUri="https://zenodo.org/record/3769997/files/figure.png" pageId="32" pageNumber="33">Figs 25</figureCitation>
C-E).
</paragraph>
<paragraph blockId="32.[140,1108,167,932]" pageId="32" pageNumber="33">
Gonopods (
<figureCitation box="[322,408,624,651]" captionStart="Figure 26" captionStartId="31.[140,212,1590,1612]" captionTargetBox="[140,1108,512,1561]" captionTargetId="figure@31.[140,1108,511,1570]" captionTargetPageId="31" captionText="Figure 26. Eutrichodesmus griseus sp. n., paratypes ♀ (A, B) and ♁ (C, D, E); A, limbus, dorsal view; B, metatergal seta, sublateral view; C-E, left gonopod, mesal, lateral and subfrontal views, respectively. – Scale bars: A, 0.01 mm; B, 0.005 mm; C-E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3769999" httpUri="https://zenodo.org/record/3769999/files/figure.png" pageId="32" pageNumber="33">Figs 26</figureCitation>
C-E) relatively simple. Coxae abundantly micropapillate but only with two macrosetae near base of a small apicolateral lobe. Telopodite elongate, rather slender, evidently arcuate, with a rudimentary, subtriangular, distofemoral process (dp) at about midway and an evident lateral, digitiform solenomere bearing a large hairpad apically and subapically, and a simple dentiform acropodite mesally.
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="33" type="discussion">
<paragraph blockId="32.[140,1108,167,932]" pageId="32" pageNumber="33">
<emphasis bold="true" box="[185,298,800,826]" pageId="32" pageNumber="33">
<emphasis box="[185,293,800,826]" inLineHeading="true" pageId="32" pageNumber="33" reason="1">Remarks</emphasis>
:
</emphasis>
Because this pigmented species obviously shows incomplete volvation, it is not a typical “doratodesmid”, definitely epigean. Moreover, because of the elevated anterior row of tubercles on the collum, superficially it slightly resembles certain 
<taxonomicName authorityName="Silvestri" authorityYear="1896" class="Diplopoda" family="Pyrgodesmidae" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Pyrgodesmidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>