273 lines
34 KiB
XML
273 lines
34 KiB
XML
<document ID-DOI="10.3897/zookeys.12.167" ID-GBIF-Dataset="3026c846-a607-4826-b2a9-8f7a531baec5" ID-GBIF-Taxon="163971734" ID-ISSN="1313–2970" ID-Zenodo-Dep="576460" ID-ZooBank="B32EF8D5-C3C6-4C8C-8845-284125448425" LSID="urn:lsid:zoobank.org:act:AEBAEF74-7FF7-4E99-8108-D0D85ACE06C3" approvalRequired="25" approvalRequired_for_textStreams="25" checkinTime="1587995767728" checkinUser="plazi" docAuthor="Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & VandenSpiegel, Didier" docDate="2009" docId="936787D7FFA7FFA38ABB47C4FE39FC63" docLanguage="en" docName="ZK_article_1994.pdf" docOrigin="ZooKeys 12 (12)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Eutrichodesmus curticornis Golovatch, Geoffroy, Mauries & VandenSpiegel 2009, sp. n." docType="treatment" docUuid="AEBAEF74-7FF7-4E99-8108-D0D85ACE06C3" docUuidSource="ZooBank" docVersion="4" lastPageId="23" lastPageNumber="24" masterDocId="6F5EFFAFFFB3FFB48A374557FFC6FF89" masterDocTitle="Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species" masterLastPageNumber="46" masterPageNumber="1" pageId="20" pageNumber="21" updateTime="1643565021146" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei</mods:namePart>
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<mods:affiliation>Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Ecologie & Gestion de la Biodiversité, UMR 7179 du CNRS, Equipe EVOLTRAIT, Brunoy, France & D 362 Dc 0 - & CC</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mauriès, Jean-Paul</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Muséum national d’Histoire naturelle, Département Systématique et Evolution, Section Arthropodes, Paris, France & - & - 42 F & D 01 - 28 Ad- 43 F</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>VandenSpiegel, Didier</mods:namePart>
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<mods:affiliation>Musée Royal de l’Afrique centrale, Tervuren, Belgium & A & EE & F- 04802 E 68 Cb</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-06-18</mods:number>
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||
</mods:detail>
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<mods:detail type="volume">
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<mods:number>12</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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||
<mods:number>12</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>46</mods:end>
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</mods:extent>
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</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.3897/zookeys.12.167</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">3026c846-a607-4826-b2a9-8f7a531baec5</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576460</mods:identifier>
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||
<mods:identifier type="ZooBank">B32EF8D5-C3C6-4C8C-8845-284125448425</mods:identifier>
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||
</mods:mods>
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<treatment ID-GBIF-Taxon="163971734" LSID="urn:lsid:zoobank.org:act:AEBAEF74-7FF7-4E99-8108-D0D85ACE06C3" httpUri="http://treatment.plazi.org/id/936787D7FFA7FFA38ABB47C4FE39FC63" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
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<subSubSection box="[140,1108,658,686]" pageId="20" pageNumber="21" type="nomenclature">
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||
<paragraph blockId="20.[140,1108,658,756]" box="[140,1108,658,686]" pageId="20" pageNumber="21">
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||
<heading box="[140,1108,658,686]" level="2" pageId="20" pageNumber="21">
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||
<emphasis bold="true" box="[140,1108,658,686]" pageId="20" pageNumber="21">
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||
<taxonomicName authority="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityName="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityYear="2009" box="[140,1032,658,686]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="curticornis" status="sp. n.">
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<emphasis bold="true" box="[140,463,659,686]" italics="true" pageId="20" pageNumber="21">
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<emphasis box="[140,328,659,686]" inLineHeading="true" pageId="20" pageNumber="21" reason="1">Eutrichodesmus</emphasis>
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curticornis
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</emphasis>
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Golovatch, Geoffroy, Mauriès & VandenSpiegel
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1043,1108,659,685]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="20" pageNumber="21" type="description">
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||
<paragraph blockId="20.[140,1108,658,756]" box="[140,982,694,721]" pageId="20" pageNumber="21">
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||
<uri box="[140,982,694,721]" pageId="20" pageNumber="21">
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||
urn:lsid:zoobank.org:act:
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<uuid box="[418,982,694,721]" pageId="20" pageNumber="21">
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||
AEBAEF74-7FF7-4E99-8108-D0D85
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<collectingRegion box="[861,922,694,721]" country="Indonesia" name="Aceh" pageId="20" pageNumber="21">ACE</collectingRegion>
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06C3
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</uuid>
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</uri>
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</paragraph>
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<paragraph blockId="20.[140,1108,658,756]" box="[140,260,730,756]" pageId="20" pageNumber="21">
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Figs
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<figureCitation box="[192,260,730,756]" captionStart-0="Figure 16" captionStart-1="Figure 17" captionStart-2="Figure 18" captionStartId-0="20.[140,212,1622,1644]" captionStartId-1="21.[140,212,1590,1612]" captionStartId-2="22.[140,211,1558,1580]" captionTargetBox-0="[140,1108,928,1602]" captionTargetBox-1="[140,1108,513,1570]" captionTargetBox-2="[160,1087,165,1538]" captionTargetId-0="figure@20.[137,1111,928,1602]" captionTargetId-1="figure@21.[140,1108,510,1573]" captionTargetId-2="figure@22.[160,1087,165,1539]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="Figure 16. Eutrichodesmus curticornis sp. n., ♀ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." captionText-1="Figure 17. Eutrichodesmus curticornis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, dorsal view; C, midbody segment, dorsal view; D, posterior part of body, caudal view. Scale bars: A, 1.0 mm; B-D, 0.1 mm." captionText-2="Figure 18. Eutrichodesmus curticornis sp. n., ♀ paratype (A-F) and ♁ holotype (G-H); A, anterior part of body, ventral view; B, midbody segments, ventral view; C, posterior part of body, ventral view; D, cross-section of a midbody segment, caudal view; E, texture of tegument; F, legs, caudal view; G, H, right gonopod, lateral and mesal views, respectively. – Scale bars: A-D, G, 0.1 mm; E, 0.01 mm; F, H, 0.02 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3813124" figureDoi-1="http://doi.org/10.5281/zenodo.3813126" figureDoi-2="http://doi.org/10.5281/zenodo.3813128" httpUri-0="https://zenodo.org/record/3813124/files/figure.png" httpUri-1="https://zenodo.org/record/3813126/files/figure.png" httpUri-2="https://zenodo.org/record/3813128/files/figure.png" pageId="20" pageNumber="21">16-18</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="20" pageNumber="21" type="materials_examined">
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||
<paragraph blockId="20.[140,1107,800,897]" pageId="20" pageNumber="21">
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||
<emphasis bold="true" box="[140,314,800,826]" pageId="20" pageNumber="21">
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<emphasis box="[140,198,800,826]" inLineHeading="true" pageId="20" pageNumber="21" reason="1">Type</emphasis>
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material:
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2610648301" collectingDate="2003-01-19" collectorName="L. Deharveng & A. Bedos" country="Vietnam" county="Anh Son" location="Hang Lung Bo Cave" municipality="Hoi Son" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="2" stateProvince="Nghe An" typeStatus="holotype">
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<collectingCountry box="[323,424,800,826]" name="Vietnam" pageId="20" pageNumber="21">Vietnam</collectingCountry>
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,
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<collectingRegion box="[437,613,800,827]" country="Vietnam" name="Nghe An" pageId="20" pageNumber="21">Nghê An Prov.</collectingRegion>
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,
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<collectingCounty box="[626,731,800,826]" pageId="20" pageNumber="21">Anh Son</collectingCounty>
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:
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<collectingMunicipality box="[745,846,800,826]" pageId="20" pageNumber="21">Hoi Son</collectingMunicipality>
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,
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||
<location LSID="urn:lsid:plazi:treatment:936787D7FFA7FFA38ABB47C4FE39FC63:1E11601AFFA7FFA0896B4676FB89FCB3" box="[860,1103,800,827]" country="Vietnam" county="Anh Son" municipality="Hoi Son" name="Hang Lung Bo Cave" pageId="20" pageNumber="21" stateProvince="Nghe An">Hang Lung Bo Cave</location>
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, hand collected,
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<collectingDate box="[321,437,836,862]" pageId="20" pageNumber="21" value="2003-01-19">
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<date box="[321,437,836,862]" pageId="20" pageNumber="21" value="2003-01-19">19.I.2003</date>
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</collectingDate>
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, leg.
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<collectorName box="[498,655,835,862]" pageId="20" pageNumber="21">L. Deharveng</collectorName>
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and
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<collectorName box="[714,817,835,862]" pageId="20" pageNumber="21">A. Bedos</collectorName>
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(VIET-901),
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<typeStatus box="[977,1076,835,861]" pageId="20" pageNumber="21">holotype</typeStatus>
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♁ (MNHN JC 321),
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<typeStatus box="[359,469,871,897]" pageId="20" pageNumber="21">paratypes</typeStatus>
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:
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<specimenCount box="[480,517,871,897]" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
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(MNHN JC 321),
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<specimenCount box="[750,787,871,897]" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
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(SEM)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3813124" ID-Zenodo-Dep="3813124" httpUri="https://zenodo.org/record/3813124/files/figure.png" pageId="20" pageNumber="21" startId="20.[140,212,1622,1644]" targetBox="[140,1108,928,1602]" targetPageId="20">
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||
<paragraph blockId="20.[140,1107,1622,1676]" pageId="20" pageNumber="21">
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||
<emphasis bold="true" box="[140,250,1622,1644]" pageId="20" pageNumber="21">
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||
<figureCitation box="[140,243,1622,1644]" captionStart-0="Figure 17" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStartId-0="21.[140,212,1590,1612]" captionStartId-1="4.[140,213,1558,1580]" captionStartId-2="5.[140,212,1442,1464]" captionStartId-3="7.[140,212,1622,1644]" captionStartId-4="8.[140,212,1498,1520]" captionStartId-5="9.[140,212,1558,1580]" captionTargetBox-0="[140,1108,513,1570]" captionTargetBox-1="[140,1108,502,1538]" captionTargetBox-2="[140,1108,165,1422]" captionTargetBox-3="[140,1108,623,1602]" captionTargetBox-4="[140,1108,165,1478]" captionTargetBox-5="[140,1108,478,1538]" captionTargetId-0="figure@21.[140,1108,510,1573]" captionTargetId-1="figure@4.[140,1108,502,1539]" captionTargetId-2="figure@5.[140,1108,165,1423]" captionTargetId-3="figure@7.[140,1108,622,1602]" captionTargetId-4="figure@8.[140,1108,165,1478]" captionTargetId-5="figure@9.[140,1108,478,1538]" captionTargetPageId-0="21" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="Figure 17. Eutrichodesmus curticornis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, dorsal view; C, midbody segment, dorsal view; D, posterior part of body, caudal view. Scale bars: A, 1.0 mm; B-D, 0.1 mm." captionText-1="Figure 2. Eutrichodesmus distinctus sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, lateral and dorsal views, respectively; C, F, midbody segments, lateral and dorsal views, respectively; D, G, posterior part of body, lateral and dorsal views, respectively. – Scale bars: A, 0.5 mm; B-G, 0.2 mm." captionText-2="Figure 3. Eutrichodesmus distinctus sp. n., ♁ paratypes; A, midbody segments, ventral view; B, telson, ventral view; C, anterior part of body, ventral view; D, texture of tegument, dorsal view; E, both gonopods in situ, ventral view; F-H, gonopod, ventral, mesal and lateral views, respectively. – Scale bars: A, 0.5 mm; B, E, G, 0.1 mm; C, H, 0.2 mm; D, 0.02 mm; F, 0.05 mm." captionText-3="Figure 4. Eutrichodesmus regularis sp. n., ♁ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." captionText-4="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." captionText-5="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3813126" figureDoi-1="http://doi.org/10.5281/zenodo.3769973" figureDoi-2="http://doi.org/10.5281/zenodo.3769975" figureDoi-3="http://doi.org/10.5281/zenodo.3769977" figureDoi-4="http://doi.org/10.5281/zenodo.3769979" figureDoi-5="http://doi.org/10.5281/zenodo.3769981" httpUri-0="https://zenodo.org/record/3813126/files/figure.png" httpUri-1="https://zenodo.org/record/3769973/files/figure.png" httpUri-2="https://zenodo.org/record/3769975/files/figure.png" httpUri-3="https://zenodo.org/record/3769977/files/figure.png" httpUri-4="https://zenodo.org/record/3769979/files/figure.png" httpUri-5="https://zenodo.org/record/3769981/files/figure.png" pageId="20" pageNumber="21">Figure 16</figureCitation>
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.
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</emphasis>
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<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[255,491,1622,1644]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="curticornis" status="sp. n.">
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<emphasis box="[255,491,1622,1644]" italics="true" pageId="20" pageNumber="21">Eutrichodesmus curticornis</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel box="[496,547,1623,1644]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
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,
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||
<specimenCount box="[556,568,1623,1644]" pageId="20" pageNumber="21" type="female">♀</specimenCount>
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<typeStatus box="[579,661,1623,1644]" pageId="20" pageNumber="21">paratype</typeStatus>
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; habitus, lateral view. (Photographed not to scale by L. Deharveng).
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</paragraph>
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</caption>
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||
<subSubSection box="[185,764,167,193]" pageId="21" pageNumber="22" type="etymology">
|
||
<paragraph blockId="21.[140,1108,167,475]" box="[185,764,167,193]" pageId="21" pageNumber="22">
|
||
<emphasis bold="true" box="[185,263,167,193]" pageId="21" pageNumber="22">
|
||
<emphasis box="[185,258,167,193]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Name</emphasis>
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||
:
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||
</emphasis>
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To emphasize the unusually short antennae.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="diagnosis">
|
||
<paragraph blockId="21.[140,1108,167,475]" pageId="21" pageNumber="22">
|
||
<emphasis bold="true" box="[185,312,201,227]" pageId="21" pageNumber="22">
|
||
<emphasis box="[185,307,201,227]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Diagnosis</emphasis>
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||
:
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||
</emphasis>
|
||
Differs from congeners by the perfect volvation, coupled with the very short antennae, legs and paraterga, the contiguous paramedian tubercles above the antennal sockets, the regular mixostictic pattern of metatergal tuberculation and a few minor details of gonopod structure (in particular, the shape of the telopodite and distofemoral process).
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="description">
|
||
<paragraph blockId="21.[140,1108,167,475]" pageId="21" pageNumber="22">
|
||
<emphasis bold="true" box="[185,335,377,403]" pageId="21" pageNumber="22">
|
||
<emphasis box="[185,330,377,403]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Description</emphasis>
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||
:
|
||
</emphasis>
|
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Length of adults of both sexes ca
|
||
<quantity box="[719,858,378,405]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.3" metricValueMin="4.7" pageId="21" pageNumber="22" unit="mm" value="5.0" valueMax="5.3" valueMin="4.7">4.7-5.3 mm</quantity>
|
||
, width
|
||
<quantity box="[944,1103,378,405]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="21" pageNumber="22" unit="mm" value="0.675" valueMax="0.7" valueMin="0.65">0.65–0.7 mm</quantity>
|
||
, body broadest at segment 3 or 4.
|
||
<typeStatus box="[525,632,413,439]" pageId="21" pageNumber="22">Holotype</typeStatus>
|
||
ca
|
||
<quantity box="[672,763,413,440]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="21" pageNumber="22" unit="mm" value="4.7">4.7 mm</quantity>
|
||
long and
|
||
<quantity box="[880,986,413,440]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="21" pageNumber="22" unit="mm" value="0.65">0.65 mm</quantity>
|
||
wide. Coloration uniformly pallid, shown pinkish because of a photographic artifact (Fig. 16).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3813126" ID-Zenodo-Dep="3813126" httpUri="https://zenodo.org/record/3813126/files/figure.png" pageId="21" pageNumber="22" startId="21.[140,212,1590,1612]" targetBox="[140,1108,513,1570]" targetPageId="21">
|
||
<paragraph blockId="21.[140,1109,1590,1676]" pageId="21" pageNumber="22">
|
||
<emphasis bold="true" box="[140,253,1590,1612]" pageId="21" pageNumber="22">
|
||
<figureCitation box="[140,248,1590,1612]" captionStart-0="Figure 18" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStartId-0="22.[140,211,1558,1580]" captionStartId-1="4.[140,213,1558,1580]" captionStartId-2="5.[140,212,1442,1464]" captionStartId-3="7.[140,212,1622,1644]" captionStartId-4="8.[140,212,1498,1520]" captionStartId-5="9.[140,212,1558,1580]" captionStartId-6="10.[140,212,1622,1644]" captionTargetBox-0="[160,1087,165,1538]" captionTargetBox-1="[140,1108,502,1538]" captionTargetBox-2="[140,1108,165,1422]" captionTargetBox-3="[140,1108,623,1602]" captionTargetBox-4="[140,1108,165,1478]" captionTargetBox-5="[140,1108,478,1538]" captionTargetBox-6="[140,1108,308,1602]" captionTargetId-0="figure@22.[160,1087,165,1539]" captionTargetId-1="figure@4.[140,1108,502,1539]" captionTargetId-2="figure@5.[140,1108,165,1423]" captionTargetId-3="figure@7.[140,1108,622,1602]" captionTargetId-4="figure@8.[140,1108,165,1478]" captionTargetId-5="figure@9.[140,1108,478,1538]" captionTargetId-6="figure@10.[140,1108,308,1603]" captionTargetPageId-0="22" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionText-0="Figure 18. Eutrichodesmus curticornis sp. n., ♀ paratype (A-F) and ♁ holotype (G-H); A, anterior part of body, ventral view; B, midbody segments, ventral view; C, posterior part of body, ventral view; D, cross-section of a midbody segment, caudal view; E, texture of tegument; F, legs, caudal view; G, H, right gonopod, lateral and mesal views, respectively. – Scale bars: A-D, G, 0.1 mm; E, 0.01 mm; F, H, 0.02 mm." captionText-1="Figure 2. Eutrichodesmus distinctus sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, lateral and dorsal views, respectively; C, F, midbody segments, lateral and dorsal views, respectively; D, G, posterior part of body, lateral and dorsal views, respectively. – Scale bars: A, 0.5 mm; B-G, 0.2 mm." captionText-2="Figure 3. Eutrichodesmus distinctus sp. n., ♁ paratypes; A, midbody segments, ventral view; B, telson, ventral view; C, anterior part of body, ventral view; D, texture of tegument, dorsal view; E, both gonopods in situ, ventral view; F-H, gonopod, ventral, mesal and lateral views, respectively. – Scale bars: A, 0.5 mm; B, E, G, 0.1 mm; C, H, 0.2 mm; D, 0.02 mm; F, 0.05 mm." captionText-3="Figure 4. Eutrichodesmus regularis sp. n., ♁ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." captionText-4="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." captionText-5="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." captionText-6="Figure 7. Eutrichodesmus regularis sp. n., ♁ paratypes; A-F, gonopods, frontoventral, submesal, mesal, mesal, mesal and lateral views, respectively. – Scale bars: A-C, 0.1 mm; D, 0.05 mm; E, F, 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3813128" figureDoi-1="http://doi.org/10.5281/zenodo.3769973" figureDoi-2="http://doi.org/10.5281/zenodo.3769975" figureDoi-3="http://doi.org/10.5281/zenodo.3769977" figureDoi-4="http://doi.org/10.5281/zenodo.3769979" figureDoi-5="http://doi.org/10.5281/zenodo.3769981" figureDoi-6="http://doi.org/10.5281/zenodo.3769983" httpUri-0="https://zenodo.org/record/3813128/files/figure.png" httpUri-1="https://zenodo.org/record/3769973/files/figure.png" httpUri-2="https://zenodo.org/record/3769975/files/figure.png" httpUri-3="https://zenodo.org/record/3769977/files/figure.png" httpUri-4="https://zenodo.org/record/3769979/files/figure.png" httpUri-5="https://zenodo.org/record/3769981/files/figure.png" httpUri-6="https://zenodo.org/record/3769983/files/figure.png" pageId="21" pageNumber="22">Figure 17</figureCitation>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[262,501,1590,1612]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="curticornis" status="sp. n.">
|
||
<emphasis box="[262,501,1590,1612]" italics="true" pageId="21" pageNumber="22">Eutrichodesmus curticornis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[509,563,1591,1612]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
|
||
,
|
||
<specimenCount box="[576,588,1591,1612]" pageId="21" pageNumber="22" type="female">♀</specimenCount>
|
||
<typeStatus box="[602,684,1591,1612]" pageId="21" pageNumber="22">paratype</typeStatus>
|
||
;
|
||
<emphasis bold="true" box="[695,714,1590,1612]" pageId="21" pageNumber="22">A</emphasis>
|
||
, habitus, lateral view;
|
||
<emphasis bold="true" box="[931,948,1590,1612]" pageId="21" pageNumber="22">B</emphasis>
|
||
, anterior part of body, dorsal view;
|
||
<emphasis bold="true" box="[314,333,1622,1644]" pageId="21" pageNumber="22">C</emphasis>
|
||
, midbody segment, dorsal view;
|
||
<emphasis bold="true" box="[643,662,1622,1644]" pageId="21" pageNumber="22">D</emphasis>
|
||
, posterior part of body, caudal view. Scale bars: A, 1.0 mm; B-D,
|
||
<quantity box="[305,382,1655,1676]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="mm" value="0.1">0.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3813128" ID-Zenodo-Dep="3813128" httpUri="https://zenodo.org/record/3813128/files/figure.png" pageId="22" pageNumber="23" startId="22.[140,211,1558,1580]" targetBox="[160,1087,165,1538]" targetPageId="22">
|
||
<paragraph blockId="22.[140,1108,1558,1676]" pageId="22" pageNumber="23">
|
||
<emphasis bold="true" box="[140,251,1558,1580]" pageId="22" pageNumber="23">
|
||
<figureCitation box="[140,245,1558,1580]" captionStart-0="Figure 19" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStartId-0="24.[140,212,1073,1095]" captionStartId-1="4.[140,213,1558,1580]" captionStartId-2="5.[140,212,1442,1464]" captionStartId-3="7.[140,212,1622,1644]" captionStartId-4="8.[140,212,1498,1520]" captionStartId-5="9.[140,212,1558,1580]" captionStartId-6="10.[140,212,1622,1644]" captionStartId-7="11.[140,212,1622,1644]" captionTargetBox-0="[140,1108,165,1053]" captionTargetBox-1="[140,1108,502,1538]" captionTargetBox-2="[140,1108,165,1422]" captionTargetBox-3="[140,1108,623,1602]" captionTargetBox-4="[140,1108,165,1478]" captionTargetBox-5="[140,1108,478,1538]" captionTargetBox-6="[140,1108,308,1602]" captionTargetBox-7="[140,1108,865,1602]" captionTargetId-0="figure@24.[140,1108,165,1053]" captionTargetId-1="figure@4.[140,1108,502,1539]" captionTargetId-2="figure@5.[140,1108,165,1423]" captionTargetId-3="figure@7.[140,1108,622,1602]" captionTargetId-4="figure@8.[140,1108,165,1478]" captionTargetId-5="figure@9.[140,1108,478,1538]" captionTargetId-6="figure@10.[140,1108,308,1603]" captionTargetId-7="figure@11.[140,1108,865,1605]" captionTargetPageId-0="24" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionText-0="Figure 19. Eutrichodesmus asteroides sp. n., ♁ paratype from Hang Cha Noi Cave; habitus, lateral view. (Photographed by L. Deharveng)." captionText-1="Figure 2. Eutrichodesmus distinctus sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, lateral and dorsal views, respectively; C, F, midbody segments, lateral and dorsal views, respectively; D, G, posterior part of body, lateral and dorsal views, respectively. – Scale bars: A, 0.5 mm; B-G, 0.2 mm." captionText-2="Figure 3. Eutrichodesmus distinctus sp. n., ♁ paratypes; A, midbody segments, ventral view; B, telson, ventral view; C, anterior part of body, ventral view; D, texture of tegument, dorsal view; E, both gonopods in situ, ventral view; F-H, gonopod, ventral, mesal and lateral views, respectively. – Scale bars: A, 0.5 mm; B, E, G, 0.1 mm; C, H, 0.2 mm; D, 0.02 mm; F, 0.05 mm." captionText-3="Figure 4. Eutrichodesmus regularis sp. n., ♁ paratype; habitus, lateral view. (Photographed not to scale by L. Deharveng)." captionText-4="Figure 5. Eutrichodesmus regularis sp. n., ♁ paratype; A, habitus, lateral view; B, E, anterior part of body, frontodorsal and ventral views, respectively; C, midbody segments, dorsal view; D, F, posterior part of body, dorsal and ventral views, respectively. – Scale bars: A, C, 0.5 mm; B, D-F, 0.2 mm." captionText-5="Figure 6. Eutrichodesmus regularis sp. n., ♁ paratype; A, texture of tegument, dorsal view; B, metatergal tubercle, dorsal view; C, metatergal seta, dorsal view; D, cross-section of a midbody segment, caudal view; E, midbody legs, caudal view; F, segments 7 and 8, ventral view; G, left gonopod in situ, caudoventral view. – Scale bars: A, B, 0.05 mm; C, 0.01 mm; D, F, 0.2 mm; E, G, 0.1 mm." captionText-6="Figure 7. Eutrichodesmus regularis sp. n., ♁ paratypes; A-F, gonopods, frontoventral, submesal, mesal, mesal, mesal and lateral views, respectively. – Scale bars: A-C, 0.1 mm; D, 0.05 mm; E, F, 0.2 mm." captionText-7="Figure 8. Eutrichodesmus aster sp. n., ♁ paratype from Tham Han Cave; habitus, lateral view. (Photographed not to scale by L. Deharveng)." figureDoi-0="http://doi.org/10.5281/zenodo.3813130" figureDoi-1="http://doi.org/10.5281/zenodo.3769973" figureDoi-2="http://doi.org/10.5281/zenodo.3769975" figureDoi-3="http://doi.org/10.5281/zenodo.3769977" figureDoi-4="http://doi.org/10.5281/zenodo.3769979" figureDoi-5="http://doi.org/10.5281/zenodo.3769981" figureDoi-6="http://doi.org/10.5281/zenodo.3769983" figureDoi-7="http://doi.org/10.5281/zenodo.3769985" httpUri-0="https://zenodo.org/record/3813130/files/figure.png" httpUri-1="https://zenodo.org/record/3769973/files/figure.png" httpUri-2="https://zenodo.org/record/3769975/files/figure.png" httpUri-3="https://zenodo.org/record/3769977/files/figure.png" httpUri-4="https://zenodo.org/record/3769979/files/figure.png" httpUri-5="https://zenodo.org/record/3769981/files/figure.png" httpUri-6="https://zenodo.org/record/3769983/files/figure.png" httpUri-7="https://zenodo.org/record/3769985/files/figure.png" pageId="22" pageNumber="23">Figure 18</figureCitation>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[259,492,1558,1580]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="curticornis" status="sp. n.">
|
||
<emphasis box="[259,492,1558,1580]" italics="true" pageId="22" pageNumber="23">Eutrichodesmus curticornis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[500,552,1559,1580]" pageId="22" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
|
||
,
|
||
<specimenCount box="[564,576,1559,1580]" pageId="22" pageNumber="23" type="female">♀</specimenCount>
|
||
<typeStatus box="[590,668,1559,1580]" pageId="22" pageNumber="23">paratype</typeStatus>
|
||
(A-F) and ♁
|
||
<typeStatus box="[799,879,1558,1580]" pageId="22" pageNumber="23">holotype</typeStatus>
|
||
(G-H);
|
||
<emphasis bold="true" box="[959,978,1558,1580]" pageId="22" pageNumber="23">A</emphasis>
|
||
, anterior part of body, ventral view;
|
||
<emphasis bold="true" box="[353,370,1590,1612]" pageId="22" pageNumber="23">B</emphasis>
|
||
, midbody segments, ventral view;
|
||
<emphasis bold="true" box="[700,718,1590,1612]" pageId="22" pageNumber="23">C</emphasis>
|
||
, posterior part of body, ventral view;
|
||
<emphasis bold="true" box="[1082,1101,1590,1612]" pageId="22" pageNumber="23">D</emphasis>
|
||
, cross-section of a midbody segment, caudal view;
|
||
<emphasis bold="true" box="[593,608,1622,1644]" pageId="22" pageNumber="23">E</emphasis>
|
||
, texture of tegument;
|
||
<emphasis bold="true" box="[813,827,1622,1644]" pageId="22" pageNumber="23">F</emphasis>
|
||
, legs, caudal view;
|
||
<emphasis bold="true" box="[1002,1052,1622,1644]" pageId="22" pageNumber="23">G, H</emphasis>
|
||
, right gonopod, lateral and mesal views, respectively. – Scale bars: A-D, G,
|
||
<quantity box="[761,836,1655,1676]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="mm" value="0.1">0.1 mm</quantity>
|
||
; E,
|
||
<quantity box="[870,957,1655,1676]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="mm" value="0.01">0.01 mm</quantity>
|
||
; F, H,
|
||
<quantity box="[1017,1104,1655,1676]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="23" unit="mm" value="0.02">0.02 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="23.[140,1108,167,1003]" pageId="23" pageNumber="24">
|
||
Adults with 20 segments (Figs 16, 17A), pattern of conglobation typical of “doratodesmids”. Head (Fig. 18A) with a contiguous pair of paramedian tubercles above antennal sockets; antennae very short and clavate, but tegument (Fig. 18E) and many other characters (Figs 18B, C, F) much as in
|
||
<taxonomicName authority="Golovatch & Geoffroy & Mauriès & VandenSpiegel, 2009" authorityName="Golovatch & Geoffroy & Mauriès & VandenSpiegel" authorityYear="2009" box="[662,797,272,298]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="distinctus" status="sp. n.">
|
||
<emphasis box="[662,797,272,298]" italics="true" pageId="23" pageNumber="24">E. distinctus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[806,870,273,299]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
|
||
; collum rather regularly tuberculate (Fig. 17B). Metaterga behind collum with three transverse, regular, mixostictic rows of rounded, flat, undifferentiated tubercles (Figs 17A, C), each crowned with a broken-off seta (Fig. 18E). Paraterga subvertical, rather wide, slightly surpassing level of venter, with only a slight impression at base and nearly continuing general outline of convex dorsum (Fig. 18D); paraterga 2 (Figs 17A, 18A) evidently lobulate only anterolaterally but not laterally (Fig. 17A). Following paraterga broadly rounded, very slightly bilobate laterally and unilobate caudolaterally (Figs 17A, 18B). Limbus rather irregularly crenulate, almost hidden by nearby abundant microvilli (Fig. 18E). Pore formula normal, ozopores poorly visible, located dorsally near base of ventrocaudal lobulation (Fig. 17A). Epiproct short, with only slightly differentiated tubercles (Fig. 17D).
|
||
</paragraph>
|
||
<paragraph blockId="23.[140,1108,167,1003]" box="[185,983,695,721]" pageId="23" pageNumber="24">Legs very short and stout, barely reaching edge of paraterga (Fig. 18F).</paragraph>
|
||
<paragraph blockId="23.[140,1108,167,1003]" pageId="23" pageNumber="24">Gonopods (Figs 18G, H) relatively complex. Coxae abundantly micropapillate and setose, with a usual, apicolateral lobe. Telopodite elongate, slender, only very slightly arcuate, with a marked, denticulate distofemoral process (dp) at about midway and an evident distolateral tooth; seminal groove terminating without hairpad at base of the tooth.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="23" pageNumber="24" type="discussion">
|
||
<paragraph blockId="23.[140,1108,167,1003]" pageId="23" pageNumber="24">
|
||
<emphasis bold="true" box="[185,298,906,932]" pageId="23" pageNumber="24">
|
||
<emphasis box="[185,293,906,932]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">Remarks</emphasis>
|
||
:
|
||
</emphasis>
|
||
This pallid species shows short antennae, legs and paraterga, all these traits apparently being related to the small body size. It is also a typical “doratodesmid”, possibly yet one more troglobite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |