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<document id="F115DBCF44BD6201AB935E5BB8CADEC0" ID-CLB-Dataset="126748" ID-DOI="10.1111/zoj.12330" ID-GBIF-Dataset="6f1d30f9-d20f-415b-a70c-a65074943adf" ID-ISSN="0024-4082" ID-Zenodo-Dep="5358180" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630223637443" checkinUser="marcus" docAuthor="Hosoishi, Shingo &amp; Ogata, Kazuo" docDate="2016" docId="1E64E12B3E4CFF857C65B88FD934FBFD" docLanguage="en" docName="zoj12330.pdf" docOrigin="Zoological Journal of the Linnean Society 176 (3)" docSource="http://dx.doi.org/10.1111/zoj.12330" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Crematogaster suehiro Terayama 1999" docType="treatment" docVersion="8" lastPageNumber="600" masterDocId="E25D99533E7FFFB07F0BBA1BD82FFFC9" masterDocTitle="Systematics and biogeography of the ant genus Crematogaster Lund subgenus Orthocrema Santschi in Asia (Hymenoptera: Formicidae)" masterLastPageNumber="606" masterPageNumber="547" pageNumber="598" updateTime="1699216389104" updateUser="ExternalLinkService">
<mods:mods id="15051AC5BF8E3E2BF598BCD60795C7D8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="50C0F88C33D8A0D6FCBC8ADBB4AEF50D">Systematics and biogeography of the ant genus Crematogaster Lund subgenus Orthocrema Santschi in Asia (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="4499F2B341BDC4565AE08D73A30F4871">Hosoishi, Shingo</mods:namePart>
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<mods:namePart id="3910F9C9987CE1EB5960862AA051BA29">Ogata, Kazuo</mods:namePart>
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<mods:title id="7710B4138BBA27B0D5BDB63E4E7B4DE0">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="507A7E78F469DD58420F8DA025EE56AF">
<mods:date id="618B92163E2FE6E1451741D4D6881656">2016</mods:date>
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<mods:number id="461A7A2C4EDFADC98CC2FC3444870396">2016-02-10</mods:number>
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<mods:detail id="DDF8EF74DFEB6B99F0853892DBBB436D" type="volume">
<mods:number id="3B0F88A7E4B7674A5579D69355050D7D">176</mods:number>
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<mods:start id="6D15602F01DE0AFF29A1B6A8E459898C">547</mods:start>
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<mods:classification id="DEF87884A364FB72500CC731CE55CD6D">journal article</mods:classification>
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<mods:identifier id="CFC26459453420210965C9E96466F0FE" type="DOI">10.1111/zoj.12330</mods:identifier>
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<treatment id="1E64E12B3E4CFF857C65B88FD934FBFD" ID-DOI="http://doi.org/10.5281/zenodo.7015441" ID-GBIF-Taxon="199534335" ID-Zenodo-Dep="7015441" LSID="urn:lsid:plazi:treatment:1E64E12B3E4CFF857C65B88FD934FBFD" httpUri="http://treatment.plazi.org/id/1E64E12B3E4CFF857C65B88FD934FBFD" lastPageId="53" lastPageNumber="600" pageId="51" pageNumber="598">
<subSubSection id="DED703B63E4CFF837C65B88FDD42FD60" box="[878,1389,660,681]" pageId="51" pageNumber="598" type="nomenclature">
<paragraph id="9672503D3E4CFF837C65B88FDD42FD60" blockId="51.[878,1389,660,681]" box="[878,1389,660,681]" pageId="51" pageNumber="598">
<heading id="CD3AE7513E4CFF837C65B88FDD42FD60" box="[878,1389,660,681]" centered="true" fontSize="9" level="2" pageId="51" pageNumber="598" reason="4">
<taxonomicName id="51CD2BBE3E4CFF837C65B88FDD42FD60" authority="TERAYAMA, 1999" authorityName="Terayama" authorityYear="1999" box="[878,1389,660,681]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="598" phylum="Arthropoda" rank="species" species="suehiro">
<emphasis id="A4B98C2F3E4CFF837C65B88FDC86FD61" bold="true" box="[878,1193,660,681]" italics="true" pageId="51" pageNumber="598">CREMATOGASTER SUEHIRO</emphasis>
<bibRefCitation id="F25C2DCC3E4CFF837BBAB88FDD42FD60" author="Terayama M" box="[1201,1389,660,681]" pageId="51" pageNumber="598" pagination="726 - 727" refId="ref40436" refString="Terayama M. 1999. Description and record of a new and a little known ants from Japan (Formicidae). In: Yamane SK, Ikudome S, Terayama M, eds. Identification guide to the aculeata of the Nansei Islands, Japan. Sapporo: Hokkaido University Press, 726 - 727." type="book chapter" year="1999">TERAYAMA, 1999</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E4CFF837B3EB8A0DC89FD18" box="[1077,1190,699,721]" pageId="51" pageNumber="598" type="description">
<paragraph id="9672503D3E4CFF837B3EB8A0DC89FD18" blockId="51.[1077,1190,699,721]" box="[1077,1190,699,721]" pageId="51" pageNumber="598">
(
<figureCitation id="0EF64CB83E4CFF837B36B8A0DCB0FD18" box="[1085,1183,699,721]" captionStart="Figures 120–123" captionStartId="52.[144,234,953,972]" captionTargetBox="[144,1424,194,923]" captionTargetId="figure-382@52.[144,1424,194,923]" captionTargetPageId="52" captionText="Figures 120–123. Body in lateral view. 120, Crematogaster suehiro, worker [Mt. Omoto, Ishigaki Island, Japan]; 121, Crematogaster sundalandensis, worker [Maliau Basin (riparian forest), Sabah, Borneo, E. Malaysia]; 122, Crematogaster vieti, worker [near Forestry Station, Khe Kem, Pu Mat National Park, Nghe An, Vietnam]; 123, Crematogaster vieti, intermediate worker [near Forestry Station, Khe Kem, Pu Mat National Park, Nghe An, Vietnam]." figureDoi="http://doi.org/10.5281/zenodo.5358226" httpUri="https://zenodo.org/record/5358226/files/figure.png" pageId="51" pageNumber="598">FIG. 120</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E4CFF837C31B8FADD8EFD3F" box="[826,1441,737,759]" pageId="51" pageNumber="598" type="reference_group">
<paragraph id="9672503D3E4CFF837C31B8FADD8EFD3F" blockId="51.[826,1441,737,1065]" box="[826,1441,737,759]" pageId="51" pageNumber="598">
<treatmentCitationGroup id="B6DD77133E4CFF837C31B8FADD8EFD3F" box="[826,1441,737,759]" pageId="51" pageNumber="598">
<treatmentCitation id="176C762C3E4CFF837C31B8FADD00FD3E" author="Terayama M" box="[826,1327,737,759]" page="726" pageId="51" pageNumber="598" year="1999">
<taxonomicName id="51CD2BBE3E4CFF837C31B8FADD00FD3E" authority="Terayama, 1999: 726" authorityName="Terayama" authorityPageNumber="726" authorityYear="1999" box="[826,1327,737,759]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="598" phylum="Arthropoda" rank="species" species="suehiro">
<emphasis id="A4B98C2F3E4CFF837C31B8FADC16FD3F" box="[826,1081,737,758]" italics="true" pageId="51" pageNumber="598">Crematogaster suehiro</emphasis>
<bibRefCitation id="F25C2DCC3E4CFF837B4BB8FADD00FD3E" author="Terayama M" box="[1088,1327,737,759]" pageId="51" pageNumber="598" pagination="726 - 727" refId="ref40436" refString="Terayama M. 1999. Description and record of a new and a little known ants from Japan (Formicidae). In: Yamane SK, Ikudome S, Terayama M, eds. Identification guide to the aculeata of the Nansei Islands, Japan. Sapporo: Hokkaido University Press, 726 - 727." type="book chapter" year="1999">Terayama, 1999: 726</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 1–4;
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E4CFF837C31B91BDC8EFC9A" pageId="51" pageNumber="598" type="reference_group">
<paragraph id="9672503D3E4CFF837C31B91BDC8EFC9A" blockId="51.[826,1441,737,1065]" pageId="51" pageNumber="598">
<materialsCitation id="26A55A603E4CFF837C31B91BDCB1FC9A" ID-GBIF-Occurrence="3891597423" collectionCode="MNHA" country="Japan" location="Ishigaki Island" pageId="51" pageNumber="598" specimenCount="4" specimenCount-worker="2" typeStatus="holotype">
<typeStatus id="4976EE9F3E4CFF837C31B91BDBBAFCDF" box="[826,917,768,790]" pageId="51" pageNumber="598" type="holotype">holotype</typeStatus>
<specimenCount id="80CB9BB43E4CFF837C90B91BDBC8FCDF" box="[923,999,768,790]" pageId="51" pageNumber="598" type="worker">worker</specimenCount>
and
<specimenCount id="80CB9BB43E4CFF837B15B91BDC6AFCDC" box="[1054,1093,768,789]" pageId="51" pageNumber="598">two</specimenCount>
paratype
<specimenType id="C663DB313E4CFF837BB9B91BDD24FCDF" box="[1202,1291,768,790]" pageId="51" pageNumber="598">workers</specimenType>
,
<specimenCount id="80CB9BB43E4CFF837A1EB91BDD14FCDC" box="[1301,1339,768,789]" pageId="51" pageNumber="598">one</specimenCount>
paratype
<specimenType id="C663DB313E4CFF837C31B904DB51FCFD" box="[826,894,799,820]" pageId="51" pageNumber="598">queen</specimenType>
and one intermediate
<specimenCount id="80CB9BB43E4CFF837B87B905DCF2FCFD" box="[1164,1245,798,820]" pageId="51" pageNumber="598" type="worker">worker</specimenCount>
,
<location id="931206E63E4CFF837BE0B905DDB3FCFD" LSID="urn:lsid:plazi:treatment:1E64E12B3E4CFF857C65B88FD934FBFD:931206E63E4CFF837BE0B905DDB3FCFD" box="[1259,1436,798,820]" country="Japan" name="Ishigaki Island" pageId="51" pageNumber="598">Ishigaki Island</location>
,
<collectingCountry id="EEDA10AD3E4CFF837C31B926DBAFFC9B" box="[826,896,829,850]" name="Japan" pageId="51" pageNumber="598">Japan</collectingCountry>
(
<collectionCode id="F0DCC8F83E4CFF837C9AB926DBC9FC9B" box="[913,998,829,850]" pageId="51" pageNumber="598">MNHA</collectionCode>
, not examined)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E4CFF837C58B947DB74FC59" pageId="51" pageNumber="598" type="reference_group">
<paragraph id="9672503D3E4CFF837C58B947DB74FC59" blockId="51.[826,1441,737,1065]" pageId="51" pageNumber="598">
<treatmentCitationGroup id="B6DD77133E4CFF837C58B947DB74FC59" pageId="51" pageNumber="598">
<treatmentCitation id="176C762C3E4CFF837C58B947DD14FCB8" author="Terayama M" box="[851,1339,859,881]" page="12" pageId="51" pageNumber="598" year="2013">
<taxonomicName id="51CD2BBE3E4CFF837C58B947DD14FCB8" authority="Terayama, 2013: 12" authorityName=", Terayama" authorityPageNumber="12" authorityYear="2013" box="[851,1339,859,881]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="598" phylum="Arthropoda" rank="species" species="miroku">
<emphasis id="A4B98C2F3E4CFF837C58B947DC7EFCB9" box="[851,1105,859,881]" italics="true" pageId="51" pageNumber="598">Crematogaster miroku</emphasis>
<bibRefCitation id="F25C2DCC3E4CFF837B52B947DD14FCB8" author="Terayama M" box="[1113,1339,860,881]" pageId="51" pageNumber="598" pagination="1 - 24" refId="ref40527" refString="Terayama M. 2013. Additions to knowledge of the ant fauna of Japan (Hymenoptera: Formicidae). Memoirs of the Myrmecological Society of Japan 3: 1 - 24." type="journal article" year="2013">Terayama, 2013: 12</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 23– 25;
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E4CFF837C69B961DD8EFC22" pageId="51" pageNumber="598" type="reference_group">
<paragraph id="9672503D3E4CFF837C69B961DD8EFC22" blockId="51.[826,1441,737,1065]" pageId="51" pageNumber="598">
<materialsCitation id="26A55A603E4CFF837C69B961DD8EFC22" ID-GBIF-Occurrence="3891597360" collectingDate="2012-11-03" collectionCode="ITLJ" collectorName="H. Takamine" country="Japan" location="Higashison" municipality="Okinawa Pref." pageId="51" pageNumber="598" specimenCount="1" specimenCount-worker="1" typeStatus="holotype">
<typeStatus id="4976EE9F3E4CFF837C69B961DBEBFC59" box="[866,964,890,912]" pageId="51" pageNumber="598" type="holotype">Holotype</typeStatus>
<specimenCount id="80CB9BB43E4CFF837CC0B961DC37FC59" box="[971,1048,890,912]" pageId="51" pageNumber="598" type="worker">worker</specimenCount>
and six paratype workers,
<location id="931206E63E4CFF837A4FB961DB4BFC67" LSID="urn:lsid:plazi:treatment:1E64E12B3E4CFF857C65B88FD934FBFD:931206E63E4CFF837A4FB961DB4BFC67" country="Japan" municipality="Okinawa Pref." name="Higashison" pageId="51" pageNumber="598">Higashison</location>
,
<location id="931206E63E4CFF837C7EB982DC0EFC66" LSID="urn:lsid:plazi:treatment:1E64E12B3E4CFF857C65B88FD934FBFD:931206E63E4CFF837C7EB982DC0EFC66" box="[885,1057,921,943]" country="Japan" municipality="Okinawa Pref." name="Kunigami-gun" pageId="51" pageNumber="598">Kunigami-gun</location>
,
<location id="931206E63E4CFF837B39B982DCF4FC66" LSID="urn:lsid:plazi:treatment:1E64E12B3E4CFF857C65B88FD934FBFD:931206E63E4CFF837B39B982DCF4FC66" box="[1074,1243,921,943]" country="Japan" municipality="Okinawa Pref." name="Okinawa-jima" pageId="51" pageNumber="598">Okinawa-jima</location>
,
<collectingMunicipality id="7616CA473E4CFF837BE7B982DDB2FC66" box="[1260,1437,921,943]" pageId="51" pageNumber="598">Okinawa Pref.</collectingMunicipality>
,
<collectingCountry id="EEDA10AD3E4CFF837C31B9A3DB50FC04" box="[826,895,952,973]" name="Japan" pageId="51" pageNumber="598">Japan</collectingCountry>
,
<date id="E27376FD3E4CFF837C81B9ACDBC5FC04" box="[906,1002,951,973]" pageId="51" pageNumber="598" value="2012-11-03">
<collectingDate id="F2378F153E4CFF837C81B9ACDBC5FC04" box="[906,1002,951,973]" pageId="51" pageNumber="598" value="2012-11-03">3.xi.2012</collectingDate>
</date>
(
<collectorName id="3B3835EB3E4CFF837CF1B9ACDCA8FC04" box="[1018,1159,951,973]" pageId="51" pageNumber="598">H. Takamine</collectorName>
) (
<collectionCode id="F0DCC8F83E4CFF837B97B9ACDCFBFC05" box="[1180,1236,951,972]" pageId="51" pageNumber="598">ITLJ</collectionCode>
, examined; images also examined from
<bibRefCitation id="F25C2DCC3E4CFF837B2FB9CDDD34FC25" author="Yoshitake H &amp; Nakatani Y &amp; Yoshimatsu S &amp; Yasuda K" box="[1060,1307,982,1004]" pageId="51" pageNumber="598" refId="ref41008" refString="Yoshitake H, Nakatani Y, Yoshimatsu S, Yasuda K. 2011. Insect Inventory Search Engine. Available from: http: // insect. niaes. affrc. go. jp / (accessed March, 2015). National Institute for Agro-Environmental Sciences, Tsukuba, Japan." type="url" year="2011">
Yoshitake
<emphasis id="A4B98C2F3E4CFF837B90B9CDDCE3FC22" box="[1179,1228,982,1003]" italics="true" pageId="51" pageNumber="598">et al</emphasis>
., 2011
</bibRefCitation>
).
<emphasis id="A4B98C2F3E4CFF837A25B9CEDD8EFC22" bold="true" box="[1326,1441,981,1003]" pageId="51" pageNumber="598">Syn. nov.</emphasis>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E4CFF837C58B9EEDC59FBE0" pageId="51" pageNumber="598" type="reference_group">
<paragraph id="9672503D3E4CFF837C58B9EEDC59FBE0" blockId="51.[826,1441,737,1065]" pageId="51" pageNumber="598">
<treatmentCitationGroup id="B6DD77133E4CFF837C58B9EEDC59FBE0" pageId="51" pageNumber="598">
<taxonomicName id="51CD2BBE3E4CFF837C58B9EEDC7AFBC0" authorityName="Terayama" authorityYear="1999" box="[851,1109,1012,1034]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="598" phylum="Arthropoda" rank="species" species="suehiro">
<emphasis id="A4B98C2F3E4CFF837C58B9EEDC7AFBC0" box="[851,1109,1012,1034]" italics="true" pageId="51" pageNumber="598">Crematogaster suehiro</emphasis>
</taxonomicName>
;
<treatmentCitation id="176C762C3E4CFF837B6EB9EFDD12FBC3" author="Blaimer BB" box="[1125,1341,1012,1034]" page="55" pageId="51" pageNumber="598" year="2012">
<bibRefCitation id="F25C2DCC3E4CFF837B6EB9EFDD12FBC3" author="Blaimer BB" box="[1125,1341,1012,1034]" pageId="51" pageNumber="598" pagination="47 - 67" refId="ref38014" refString="Blaimer BB. 2012 c. A subgeneric revision of Crematogaster and discussion of regional species groups (Hymenoptera: Formicidae). Zootaxa 3482: 47 - 67." type="journal article" year="2012">Blaimer, 2012c: 55</bibRefCitation>
</treatmentCitation>
[Combination in
<emphasis id="A4B98C2F3E4CFF837CA5BE08DBEFFBE1" box="[942,960,1043,1064]" italics="true" pageId="51" pageNumber="598">C</emphasis>
. (
<taxonomicName id="51CD2BBE3E4CFF837CD2BE08DC4FFBE1" authorityName="Santschi" baseAuthorityName="Hosoishi &amp; Ogata" baseAuthorityYear="2009" box="[985,1120,1043,1064]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="605" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">
<emphasis id="A4B98C2F3E4CFF837CD2BE08DC4FFBE1" box="[985,1120,1043,1064]" italics="true" pageId="51" pageNumber="598">Orthocrema</emphasis>
</taxonomicName>
)].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E4CFF857C31BE4ED934FBFD" lastPageId="53" lastPageNumber="600" pageId="51" pageNumber="598" type="description">
<paragraph id="9672503D3E4CFF837C31BE4EDC13FACD" blockId="51.[826,1441,1109,1284]" pageId="51" pageNumber="598">
<emphasis id="A4B98C2F3E4CFF837C31BE4EDC11FBA2" box="[826,1086,1109,1131]" italics="true" pageId="51" pageNumber="598">Worker measurements (</emphasis>
n
<emphasis id="A4B98C2F3E4CFF837B58BE4DDCAAFBA3" box="[1107,1157,1109,1131]" italics="true" pageId="51" pageNumber="598">= 6):</emphasis>
HW 0.46–0.52; HL 0.47– 0.51; CI 96–102; SL 0.46–0.50; SI 94–100; EL 0.12– 0.14; PW 0.30–0.33; WL 0.54–0.6; PSL 0.11–0.13; PtL 0.18–0.20; PtW 0.16–0.18; PtH 0.14–0.15; PpL 0.12– 0.14; PpW 0.17–0.19; PtHI 74–79; PtWI 85–90; PpWI 123–158; WI 106–112.
</paragraph>
<paragraph id="9672503D3E4CFF837C31BF2ADD8DFA8E" blockId="51.[826,1442,1329,1902]" box="[826,1442,1329,1351]" pageId="51" pageNumber="598">
<emphasis id="A4B98C2F3E4CFF837C31BF2ADCBEFA8F" box="[826,1169,1329,1350]" italics="true" pageId="51" pageNumber="598">General description of worker:</emphasis>
Workers monomorphic.
</paragraph>
<paragraph id="9672503D3E4CFF837C58BF4BDBDAF9D4" blockId="51.[826,1442,1329,1902]" pageId="51" pageNumber="598">Head subquadratic in full-face view. Mandibles with four teeth arranged at an equal distance, apical and subapical teeth large, basal two teeth smaller. Anterior clypeal margin convex in medial portion. Compound eyes distinctly projecting beyond lateral margins of head in full-face view. Scapes reaching posterolateral corners of head.</paragraph>
<paragraph id="9672503D3E4CFF847C58BC3DD92EFB67" blockId="51.[826,1442,1329,1902]" lastBlockId="52.[144,759,1115,1902]" lastPageId="52" lastPageNumber="599" pageId="51" pageNumber="598">Pronotal collar with weakly concave anterior margin in dorsal view, distinctly lower than pronotum in lateral view. Pronotal dorsum without distinct ridges laterally. Mesonotal dorsum with lateral ridges. Pronotum and mesonotum in lateral view not clearly forming continuous dorsal outline. Metanotal groove in dorsal view transverse, almost straight in median portion, forming deep concavity that is laterally margined by lamellate ridges. Propodeal spiracles oval, situated at posterolateral corners of propodeum, touching metapleural gland bullae. Propodeal spines developed, longer than diameter of propodeal spiracles, in dorsal view directed posteriad.</paragraph>
<caption id="C2B200B53E4BFF847F9BB9A2DC92FBED" ID-DOI="http://doi.org/10.5281/zenodo.5358226" ID-Zenodo-Dep="5358226" httpUri="https://zenodo.org/record/5358226/files/figure.png" pageId="52" pageNumber="599" startId="52.[144,234,953,972]" targetBox="[144,1424,194,923]" targetPageId="52">
<paragraph id="9672503D3E4BFF847F9BB9A2DC92FBED" blockId="52.[144,1421,953,1060]" pageId="52" pageNumber="599">
<emphasis id="A4B98C2F3E4BFF847F9BB9A2D97FFC05" bold="true" box="[144,336,953,972]" pageId="52" pageNumber="599">Figures 120–123.</emphasis>
Body in lateral view. 120,
<taxonomicName id="51CD2BBE3E4BFF847D70B9A2DB45FC05" authorityName="Terayama" authorityYear="1999" box="[635,874,953,972]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="599" phylum="Arthropoda" rank="species" species="suehiro">
<emphasis id="A4B98C2F3E4BFF847D70B9A2DB45FC05" box="[635,874,953,972]" italics="true" pageId="52" pageNumber="599">Crematogaster suehiro</emphasis>
</taxonomicName>
, worker [Mt. Omoto, Ishigaki Island, Japan]; 121,
<taxonomicName id="51CD2BBE3E4BFF847F9BB9CCD9FCFC20" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[144,467,982,1002]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="599" phylum="Arthropoda" rank="species" species="sundalandensis">
<emphasis id="A4B98C2F3E4BFF847F9BB9CCD9FCFC20" box="[144,467,982,1002]" italics="true" pageId="52" pageNumber="599">Crematogaster sundalandensis</emphasis>
</taxonomicName>
, worker [Maliau Basin (riparian forest), Sabah, Borneo, E. Malaysia]; 122,
<taxonomicName id="51CD2BBE3E4BFF847BFEB9CCD892FBCF" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="599" phylum="Arthropoda" rank="species" species="vieti">
<emphasis id="A4B98C2F3E4BFF847BFEB9CCD892FBCF" italics="true" pageId="52" pageNumber="599">Crematogaster vieti</emphasis>
</taxonomicName>
, worker [near Forestry Station, Khe Kem, Pu Mat National Park, Nghe An, Vietnam]; 123,
<taxonomicName id="51CD2BBE3E4BFF847BB3B9EFDDA8FBCF" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[1208,1415,1012,1031]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="599" phylum="Arthropoda" rank="species" species="vieti">
<emphasis id="A4B98C2F3E4BFF847BB3B9EFDDA8FBCF" box="[1208,1415,1012,1031]" italics="true" pageId="52" pageNumber="599">Crematogaster vieti</emphasis>
</taxonomicName>
, intermediate worker [near Forestry Station, Khe Kem, Pu Mat National Park, Nghe An, Vietnam].
</paragraph>
</caption>
<paragraph id="9672503D3E4BFF847FA3BEACDA24FA08" blockId="52.[144,759,1115,1902]" pageId="52" pageNumber="599">Petiole in dorsal view with subparallel sides and narrow anteriorly, longer than wide. Posterior portion of petiole without distinct process in lateral view. Subpetiolar process weakly developed as small process. Postpetiole in lateral view with weakly convex dorsum, as high as petiole, in dorsal view slightly wider than petiole, weakly bilobed posteriorly but without longitudinal sulcus. Subpostpetiolar process undeveloped, but venter of postpetiole convex.</paragraph>
<paragraph id="9672503D3E4BFF847FA3BFD1D952F8A7" blockId="52.[144,759,1115,1902]" pageId="52" pageNumber="599">Integument essentially smooth and shining. Dorsal surface of head smooth and shining. Mandibles with feeble rugulae and smooth interspaces. Clypeus generally smooth and shining, but with one distinct pair of longitudinal longer rugulae and one pair of shorter rugulae laterally; longer rugulae extending to posteri- or clypeal margin. Dorsal and lateral surfaces of pronotum smooth and shining; anterolateral shoulders of pronotum without rugulae. Mesopleura smooth and shining. Dorsal surface of propodeum generally smooth and shining, but with one longitudinal rugulae anteriorly. Dorsal and lateral surfaces of petiole smooth and shining. Dorsal and lateral surfaces of postpetiole smooth and shining.</paragraph>
<paragraph id="9672503D3E4BFF847C34BE40DBA5F9D4" blockId="52.[806,1422,1115,1596]" pageId="52" pageNumber="599">Standing pilosity sparse. Dorsal face of head with erect setae sparsely. Clypeus with two pairs of long setae in anterior portion, one directed upward and the other downward. Anterior clypeal margin with one pair of long setae medially and short setae laterally. Scapes with suberect setae. Mesosoma with five pairs of long erect and stout setae [ps1PN, ps2PN, psaMN, pspMN (also one pair of short setae medially), and ps1PS] that are much longer than other erect setae. Posterolateral tubercles of petiole posteriorly with two pairs of stout long setae. Postpetiole with four pairs of setae on disc anterodorsally, anterodorsally and posteriorly. Fourth abdominal tergite with erect setae abundantly, and short decumbent setae sparsely.</paragraph>
<paragraph id="9672503D3E4BFF847C34BC3DDD01F9F5" blockId="52.[806,1422,1115,1596]" box="[831,1326,1574,1596]" pageId="52" pageNumber="599">Body yellow. All flagellar segments yellow.</paragraph>
<paragraph id="9672503D3E4BFF857C2DBC78D935FED1" blockId="52.[806,1422,1635,1902]" lastBlockId="53.[164,779,197,800]" lastPageId="53" lastPageNumber="600" pageId="52" pageNumber="599">
<emphasis id="A4B98C2F3E4BFF847C2DBC78DB88F9B1" box="[806,935,1635,1656]" italics="true" pageId="52" pageNumber="599">Comments:</emphasis>
We have not examined the
<typeStatus id="4976EE9F3E4BFF847A03BC78DD69F9B1" box="[1288,1350,1635,1656]" pageId="52" pageNumber="599">types</typeStatus>
of
<taxonomicName id="51CD2BBE3E4BFF847A7FBC78DB58F95F" authorityName="Terayama" authorityYear="1999" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="599" phylum="Arthropoda" rank="species" species="suehiro">
<emphasis id="A4B98C2F3E4BFF847A7FBC78DDA9F9B1" box="[1396,1414,1635,1656]" italics="true" pageId="52" pageNumber="599">C</emphasis>
.
<emphasis id="A4B98C2F3E4BFF847C2DBC9ADB58F95F" box="[806,887,1665,1686]" italics="true" pageId="52" pageNumber="599">suehiro</emphasis>
</taxonomicName>
, but the original description and figures closely match the specimens examined. In the worker this species can be distinguished from all other members of the
<taxonomicName id="51CD2BBE3E4BFF847C7ABCC5DC3AF93B" authorityName="Forel" authorityYear="1911" box="[881,1045,1757,1779]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="599" phylum="Arthropoda" rank="species" species="quadriruga">
<emphasis id="A4B98C2F3E4BFF847C7ABCC5DBACF93A" box="[881,899,1758,1779]" italics="true" pageId="52" pageNumber="599">C</emphasis>
.
<emphasis id="A4B98C2F3E4BFF847C98BCC6DC3AF93B" box="[915,1045,1757,1778]" italics="true" pageId="52" pageNumber="599">quadriruga</emphasis>
</taxonomicName>
group by the distinct compound eyes, propodeal spiracles small and apart from metapleural gland bulla, and petiole tapering anteriorly in dorsal view. This species is similar to
<taxonomicName id="51CD2BBE3E4BFF847A78BD22DB87F8A4" authorityName="Forel" authorityYear="1911" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="599" phylum="Arthropoda" rank="species" species="quadriruga">
<emphasis id="A4B98C2F3E4BFF847A78BD22DDAAF887" box="[1395,1413,1849,1870]" italics="true" pageId="52" pageNumber="599">C</emphasis>
.
<emphasis id="A4B98C2F3E4BFF847C2DBD43DB87F8A4" box="[806,936,1880,1901]" italics="true" pageId="52" pageNumber="599">quadriruga</emphasis>
</taxonomicName>
, but can be distinguished from it by the longitudinal rugulae on the clypeus extending to the bottoms between frontal carinae and petiole tapering anteriorly.
</paragraph>
<paragraph id="9672503D3E4AFF857FB7BB3AD96DFCE9" blockId="53.[164,779,197,800]" pageId="53" pageNumber="600">
In the original description of
<taxonomicName id="51CD2BBE3E4AFF857D1EBB3AD8C4FE9C" authority=", Terayama (2013)" authorityName=", Terayama" authorityYear="2013" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="600" phylum="Arthropoda" rank="species" species="miroku">
<emphasis id="A4B98C2F3E4AFF857D1EBB3ADA08FEFF" box="[533,551,289,310]" italics="true" pageId="53" pageNumber="600">C</emphasis>
.
<emphasis id="A4B98C2F3E4AFF857D3CBB3ADAA6FEFF" box="[567,649,289,310]" italics="true" pageId="53" pageNumber="600">miroku</emphasis>
,
<bibRefCitation id="F25C2DCC3E4AFF857D92BB3AD8C4FE9C" author="Terayama M" pageId="53" pageNumber="600" pagination="1 - 24" refId="ref40527" refString="Terayama M. 2013. Additions to knowledge of the ant fauna of Japan (Hymenoptera: Formicidae). Memoirs of the Myrmecological Society of Japan 3: 1 - 24." type="journal article" year="2013">Terayama (2013)</bibRefCitation>
</taxonomicName>
separated the species from
<taxonomicName id="51CD2BBE3E4AFF857D25BB5BDA95FE9D" authorityName="Forel" authorityYear="1900" box="[558,698,319,341]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="600" phylum="Arthropoda" rank="species" species="osakensis">
<emphasis id="A4B98C2F3E4AFF857D25BB5BDA6FFE9C" box="[558,576,320,341]" italics="true" pageId="53" pageNumber="600">C</emphasis>
.
<emphasis id="A4B98C2F3E4AFF857D44BB24DA95FE9D" box="[591,698,319,340]" italics="true" pageId="53" pageNumber="600">osakensis</emphasis>
</taxonomicName>
by the carinate dorsolateral corners of mesonotum and smooth and shining mesopleuron. However, the original description and character states mentioned by him match well with
<taxonomicName id="51CD2BBE3E4AFF857E1EBBA1D9A6FE06" authorityName="Terayama" authorityYear="1999" box="[277,393,442,463]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="600" phylum="Arthropoda" rank="species" species="suehiro">
<emphasis id="A4B98C2F3E4AFF857E1EBBA1D908FE06" box="[277,295,442,463]" italics="true" pageId="53" pageNumber="600">C</emphasis>
.
<emphasis id="A4B98C2F3E4AFF857E3DBBA1D9A6FE06" box="[310,393,442,463]" italics="true" pageId="53" pageNumber="600">suehiro</emphasis>
</taxonomicName>
. Additionally longer scape (SI 96) in
<typeStatus id="4976EE9F3E4AFF857FC9BBC3D90FFE27" box="[194,288,472,494]" pageId="53" pageNumber="600" type="holotype">holotype</typeStatus>
of
<taxonomicName id="51CD2BBE3E4AFF857E48BBC2D99CFE24" authorityName=", Terayama" authorityYear="2013" box="[323,435,472,494]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="600" phylum="Arthropoda" rank="species" species="miroku">
<emphasis id="A4B98C2F3E4AFF857E48BBC2D97BFE27" box="[323,340,473,494]" italics="true" pageId="53" pageNumber="600">C</emphasis>
.
<emphasis id="A4B98C2F3E4AFF857E68BBC3D99CFE24" box="[355,435,472,493]" italics="true" pageId="53" pageNumber="600">miroku</emphasis>
</taxonomicName>
also suggests the affinity with
<taxonomicName id="51CD2BBE3E4AFF857FAFBBECD934FDC5" authorityName="Terayama" authorityYear="1999" box="[164,283,503,524]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="600" phylum="Arthropoda" rank="species" species="suehiro">
<emphasis id="A4B98C2F3E4AFF857FAFBBECD899FDC5" box="[164,182,503,524]" italics="true" pageId="53" pageNumber="600">C</emphasis>
.
<emphasis id="A4B98C2F3E4AFF857FC3BBECD934FDC5" box="[200,283,503,524]" italics="true" pageId="53" pageNumber="600">suehiro</emphasis>
</taxonomicName>
(SI 94–100) rather than
<taxonomicName id="51CD2BBE3E4AFF857D45BBECDAF2FDC5" authorityName="Forel" authorityYear="1900" box="[590,733,503,524]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="600" phylum="Arthropoda" rank="species" species="osakensis">
<emphasis id="A4B98C2F3E4AFF857D45BBECDA4FFDC5" box="[590,608,503,524]" italics="true" pageId="53" pageNumber="600">C</emphasis>
.
<emphasis id="A4B98C2F3E4AFF857D79BBECDAF2FDC5" box="[626,733,503,524]" italics="true" pageId="53" pageNumber="600">osakensis</emphasis>
</taxonomicName>
(SI 77–84). In a key to Japanese
<taxonomicName id="51CD2BBE3E4AFF857D01B80DDB25FDE2" box="[522,778,533,555]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="600" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A4B98C2F3E4AFF857D01B80DDA80FDE2" box="[522,687,534,555]" italics="true" pageId="53" pageNumber="600">Crematogaster</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="F25C2DCC3E4AFF857FA0B82FDA67FD83" author="Terayama M &amp; Kubota S &amp; Eguchi K" box="[171,584,564,586]" pageId="53" pageNumber="600" refId="ref40594" refString="Terayama M, Kubota S, Eguchi K. 2014. Encyclopedia of Japanese ants. Tokyo: Asakura Publishing. 336 pp." type="book" year="2014">Terayama, Kubota &amp; Eguchi, 2014</bibRefCitation>
),
<taxonomicName id="51CD2BBE3E4AFF857D55B82FDAFBFD80" authorityName=", Terayama" authorityYear="2013" box="[606,724,564,585]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="600" phylum="Arthropoda" rank="species" species="miroku">
<emphasis id="A4B98C2F3E4AFF857D55B82FDA5FFD80" box="[606,624,564,585]" italics="true" pageId="53" pageNumber="600">C</emphasis>
.
<emphasis id="A4B98C2F3E4AFF857D89B82FDAFBFD80" box="[642,724,564,585]" italics="true" pageId="53" pageNumber="600">miroku</emphasis>
</taxonomicName>
was separated from
<taxonomicName id="51CD2BBE3E4AFF857E57B848D9FFFDA1" authorityName="Terayama" authorityYear="1999" box="[348,464,595,616]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="600" phylum="Arthropoda" rank="species" species="suehiro">
<emphasis id="A4B98C2F3E4AFF857E57B848D941FDA1" box="[348,366,595,616]" italics="true" pageId="53" pageNumber="600">C</emphasis>
.
<emphasis id="A4B98C2F3E4AFF857E76B848D9FFFDA1" box="[381,464,595,616]" italics="true" pageId="53" pageNumber="600">suehiro</emphasis>
</taxonomicName>
by the petiole with convex sides and slightly broader posteriorly, but he mentioned ‘
<emphasis id="A4B98C2F3E4AFF857FF1B88BD969FD6C" box="[250,326,656,677]" italics="true" pageId="53" pageNumber="600">Petiole</emphasis>
..,
<emphasis id="A4B98C2F3E4AFF857E63B88BDA5DFD6C" box="[360,626,656,677]" italics="true" pageId="53" pageNumber="600">widest at posterior end</emphasis>
’ in the original description (
<bibRefCitation id="F25C2DCC3E4AFF857E5BB8B4D9D0FD0C" author="Terayama M" box="[336,511,687,709]" pageId="53" pageNumber="600" pagination="1 - 24" refId="ref40527" refString="Terayama M. 2013. Additions to knowledge of the ant fauna of Japan (Hymenoptera: Formicidae). Memoirs of the Myrmecological Society of Japan 3: 1 - 24." type="journal article" year="2013">Terayama, 2013</bibRefCitation>
). Those slight differences are treated as variation within one species,
<taxonomicName id="51CD2BBE3E4AFF857D93B8D6DB24FD2B" authorityName="Terayama" authorityYear="1999" box="[664,779,717,738]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="600" phylum="Arthropoda" rank="species" species="suehiro">
<emphasis id="A4B98C2F3E4AFF857D93B8D6DA85FD2B" box="[664,682,717,738]" italics="true" pageId="53" pageNumber="600">C</emphasis>
.
<emphasis id="A4B98C2F3E4AFF857DB3B8D6DB24FD2B" box="[696,779,717,738]" italics="true" pageId="53" pageNumber="600">suehiro</emphasis>
</taxonomicName>
until additional characters including molecular data are available.
</paragraph>
<paragraph id="9672503D3E4AFF857FAFB953DA04FC70" blockId="53.[164,779,840,953]" pageId="53" pageNumber="600">
<emphasis id="A4B98C2F3E4AFF857FAFB953D9EAFC94" box="[164,453,840,861]" italics="true" pageId="53" pageNumber="600">Distribution and biology:</emphasis>
This species is known from Southern parts of
<collectingCountry id="EEDA10AD3E4AFF857E89B97CD9E6FCB5" box="[386,457,871,892]" name="Japan" pageId="53" pageNumber="600">Japan</collectingCountry>
(Ishigaki Island,
<collectingRegion id="54099EDF3E4AFF857DA8B97DDB24FCB5" box="[675,779,870,892]" country="Japan" name="Okinawa" pageId="53" pageNumber="600">Okinawa</collectingRegion>
Island) (
<figureCitation id="0EF64CB83E4AFF857FF4B99ED963FC52" box="[255,332,901,923]" captionStart="Figure 52" captionStartId="18.[144,223,1277,1296]" captionTargetBox="[147,1422,198,1246]" captionTargetId="figure-211@18.[144,1424,195,1248]" captionTargetPageId="18" captionText="Figure 52. Geographical distribution of the species of the C. quadriruga group." figureDoi="http://doi.org/10.5281/zenodo.5358206" httpUri="https://zenodo.org/record/5358206/files/figure.png" pageId="53" pageNumber="600">Fig. 52</figureCitation>
). This species inhabits developed forests, and nests in dead twigs on trees.
</paragraph>
<paragraph id="9672503D3E4AFF857FAFB9FAD934FBFD" blockId="53.[164,779,993,1076]" pageId="53" pageNumber="600">
<emphasis id="A4B98C2F3E4AFF857FAFB9FAD9ACFC3F" box="[164,387,993,1014]" italics="true" pageId="53" pageNumber="600">Material examined:</emphasis>
<materialsCitation id="26A55A603E4AFF857E86B9FAD938FBFD" ID-GBIF-Occurrence="3891597421" collectingDate="2008-10-10" collectionCode="T" collectorName="M. Maruyama &amp; Komatsu" country="Japan" location="Mt. Omoto" municipality="Ishigaki Island" pageId="53" pageNumber="600" specimenCount="1" specimenCount-worker="1">
<collectingCountry id="EEDA10AD3E4AFF857E86B9FAD9CAFC3F" box="[397,485,993,1014]" name="Japan" pageId="53" pageNumber="600">JAPAN</collectingCountry>
:
<specimenCount id="80CB9BB43E4AFF857EFAB9FADA56FC3E" box="[497,633,993,1015]" pageId="53" pageNumber="600" type="worker">six workers</specimenCount>
,
<location id="931206E63E4AFF857D8CB9FADB28FC3F" LSID="urn:lsid:plazi:treatment:1E64E12B3E4CFF857C65B88FD934FBFD:931206E63E4AFF857D8CB9FADB28FC3F" box="[647,775,993,1014]" country="Japan" municipality="Ishigaki Island" name="Mt. Omoto" pageId="53" pageNumber="600">Mt. Omoto</location>
,
<collectingMunicipality id="7616CA473E4AFF857FAFB9E4D944FBDC" box="[164,363,1023,1045]" pageId="53" pageNumber="600">Ishigaki Island</collectingMunicipality>
,
<date id="E27376FD3E4AFF857E75BE1BD9D9FBDC" box="[382,502,1024,1045]" pageId="53" pageNumber="600" value="2008-10-10">
<collectingDate id="F2378F153E4AFF857E75BE1BD9D9FBDC" box="[382,502,1024,1045]" pageId="53" pageNumber="600" value="2008-10-10">10.x.2008</collectingDate>
</date>
(
<collectorName id="3B3835EB3E4AFF857D06BE1BDAEDFBDC" box="[525,706,1024,1045]" pageId="53" pageNumber="600">M. Maruyama</collectorName>
&amp; T.
<collectorName id="3B3835EB3E4AFF857FAFBE04D920FBFD" box="[164,271,1055,1076]" pageId="53" pageNumber="600">Komatsu</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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