270 lines
30 KiB
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270 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1110.85399" ID-Pensoft-Pub="1313-2970-1110-39" ID-Pensoft-UUID="766F9A94793D5793A9BE1E70F575798B" ID-ZooBank="4681CC6DE5F347C6B1D052DEA78BE7C3" ModsDocID="1313-2970-1110-39" checkinTime="1657006630715" checkinUser="pensoft" docAuthor="Sian Man, Nem, Siriboon, Thanit, Lin, Aung, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="E0ADEBC87C2B57F5817010371DA2C99E" docLanguage="en" docName="ZooKeys 1110: 39-102" docOrigin="ZooKeys 1110" docPubDate="2022-07-04" docSource="http://dx.doi.org/10.3897/zookeys.1110.85399" docTitle="Haploptychius blanfordi" docType="treatment" docVersion="1" id="766F9A94793D5793A9BE1E70F575798B" lastPageNumber="39" masterDocId="766F9A94793D5793A9BE1E70F575798B" masterDocTitle="Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species" masterLastPageNumber="102" masterPageNumber="39" pageNumber="39" updateTime="1657006630715" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the carnivorous land snail family Streptaxidae (Stylommatophora, Achatinina) in Myanmar, with description of four new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sian Man, Nem</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4453-734X</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Siriboon, Thanit</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Srinakharinwirot University, Bangkok, 10110, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lin, Aung</mods:namePart>
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<mods:affiliation>Fauna and Flora International, No. 35, 3</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-07-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1110</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>39</mods:start>
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<mods:end>102</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1110.85399</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1110.85399</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1110-39</mods:identifier>
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<mods:identifier type="ZooBank">4681CC6DE5F347C6B1D052DEA78BE7C3</mods:identifier>
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<mods:identifier type="Pensoft-UUID">766F9A94793D5793A9BE1E70F575798B</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:E0ADEBC87C2B57F5817010371DA2C99E" httpUri="http://treatment.plazi.org/id/E0ADEBC87C2B57F5817010371DA2C99E" lastPageNumber="39" pageId="0" pageNumber="39">
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<subSubSection pageId="0" pageNumber="39" type="nomenclature">
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName LSID="E0ADEBC8-7C2B-57F5-8170-10371DA2C99E" authority="(Theobald, 1864)" baseAuthorityName="Theobald" baseAuthorityYear="1864" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius blanfordi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordi">Haploptychius blanfordi (Theobald, 1864)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="description">
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<paragraph pageId="0" pageNumber="39">
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-C Haploptychius blanfordi A syntype NHMUK 1888.12.4.777 - 779 from Pegu with apertural dentition B specimen NHMUK 1906.2.2.338 from Akouktoung, Pegu C specimen NHMUK 1909.3.15.71 from Pegu D Haploptychius burmanicus, syntype NHMUK 1906.2.2.199 from Tongoop, Arakan with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure10" httpUri="https://binary.pensoft.net/fig/710475" pageId="0" pageNumber="39">Fig. 10A-C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="reference_group">
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis blanfordi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordi">Streptaxis blanfordi</taxonomicName>
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Theobald, 1864: 245. Type locality: Montibus Arakanensibus provincia Pegu [Arakan Mountains in Bago Region, Myanmar].
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Indian Museum, Calcutta" pageId="0" pageNumber="39" refId="B48" refString="Nevill, G, 1878. Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Indian Museum, Calcutta, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
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: 2.
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis birmanica" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="birmanica">Streptaxis birmanica</taxonomicName>
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Blanford, (in MSS.) Theobald, 1864: 245, 246 (in part). Type locality: Pegu [Bago Region, Myanmar].
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<bibRefCitation author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="273 - 279" refId="B88" refString="Theobald, W, 1865. Notes on a collection of land and freshwater shells from the Shan States, collected by F. Fedden. Journal of the Asiatic Society of Bengal 34: 273 - 279" title="Notes on a collection of land and freshwater shells from the Shan States, collected by F. Fedden." volume="34" year="1865">Theobald 1865</bibRefCitation>
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: 277.
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis blanfordianus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordianus">Streptaxis blanfordianus</taxonomicName>
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<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="143 - 177" refId="B80" refString="Stoliczka, F, 1871. Notes on the terrestrial Mollusca from the neighbourhood of Moulmein, with descriptions of new species. Journal of the Asiatic Society of Bengal 40: 143 - 177" title="Notes on the terrestrial Mollusca from the neighbourhood of Moulmein, with descriptions of new species." volume="40" year="1871">Stoliczka 1871</bibRefCitation>
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: 163, pl. 7, figs 8, 9 (unjustified emendation).
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="39" refId="B64" refString="Pfeiffer, L, 1871. Novitates Conchologicae. Series Prima. Mollusca Extramarina. Band 4. Verlag von Theodor Fischer Cassel 1-171 pls 109-137. [pp 25-40 pls 118-125 (1871) ; pp 145-171, pls 135-137 (1876). Published in parts, dates follow Johnson (1969)]" title="Novitates Conchologicae. Series Prima. Mollusca Extramarina. Band 4. Verlag von Theodor Fischer Cassel 1 - 171 pls 109 - 137. [pp 25 - 40 pls 118 - 125 (1871); pp 145 - 171, pls 135 - 137 (1876). Published in parts, dates follow Johnson (1969)]" year="1871">Pfeiffer 1876</bibRefCitation>
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: 494.
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis birmanica" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="birmanica">Streptaxis birmanica</taxonomicName>
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-
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<bibRefCitation author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="238 - 250" refId="B87" refString="Theobald, W, 1864. Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda. Journal of the Asiatic Society of Bengal 33: 238 - 250" title="Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda." volume="33" year="1864">Hanley and Theobald 1876</bibRefCitation>
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: 4, pl. 8, fig. 5 (non
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<bibRefCitation author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="238 - 250" refId="B87" refString="Theobald, W, 1864. Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda. Journal of the Asiatic Society of Bengal 33: 238 - 250" title="Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda." volume="33" year="1864">Theobald 1864</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis blanfordi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordi">Streptaxis blanfordi</taxonomicName>
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var. - Godwin-Austen 1895: 443.
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius blanfordianus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordianus">Haploptychius blanfordianus</taxonomicName>
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-
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Journal of the Society for Bibliography of Natural History" pageId="0" pageNumber="39" refId="B37" refString="Kobelt, W, 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1(12b) (2): 1-211 pls42-71 [pp. 1-96 pls 42-59 (1905) ; 97-211, pls 60-71 (1906)]." title="Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (12 b) (2): 1 - 211 pls 42 - 71 [pp. 1 - 96 pls 42 - 59 (1905); 97 - 211, pls 60 - 71 (1906)]." year="1905">Kobelt 1906</bibRefCitation>
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: 143, 144, pl. 57, fig. 15, pl. 62, figs 4, 5.
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName baseAuthorityName="Theobald" baseAuthorityYear="1864" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius blanfordi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordi">Haploptychius blanfordi</taxonomicName>
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-
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<bibRefCitation author="Richardson, L" journalOrPublisher="Tryonia" pageId="0" pageNumber="39" pagination="1 - 326" refId="B70" refString="Richardson, L, 1988. Streptaxacea: Catalog of species, Part I, Streptaxidae. Tryonia 16: 1 - 326" title="Streptaxacea: Catalog of species, Part I, Streptaxidae." volume="16" year="1988">Richardson 1988</bibRefCitation>
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: 212.
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis blanfordi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordi">Streptaxis blanfordi</taxonomicName>
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-
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<bibRefCitation author="Hanley, SCT" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="39" refId="B33" refString="Hanley, SCT, Theobald, W, 1870. Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. London[i-xviii+] 1-65 pp.[160 pls.] [pp 1-18 pls 1-40(1870) ; pp 19-28, pls 41-60 (1872); pp 29-34, pls 61-80 (1873); pp 35-48, pls 81-120 (1874); pp 49-56, pls 121-140 (1875); pp 57-65, pls 141-160 (1876)] [Published in parts, dates after Prashad (1927)]." title="Conchologia Indica: Illustrations of the Land and Freshwater Shells of British India. London [i-xviii +] 1 - 65 pp. [160 pls.] [pp 1 - 18 pls 1 - 40 (1870); pp 19 - 28, pls 41 - 60 (1872); pp 29 - 34, pls 61 - 80 (1873); pp 35 - 48, pls 81 - 120 (1874); pp 49 - 56, pls 121 - 140 (1875); pp 57 - 65, pls 141 - 160 (1876)] [Published in parts, dates after Prashad (1927)]." year="1870">Hanley and Theobald 1870</bibRefCitation>
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: 4, pl. 8, fig. 10 (non
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<bibRefCitation author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="238 - 250" refId="B87" refString="Theobald, W, 1864. Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda. Journal of the Asiatic Society of Bengal 33: 238 - 250" title="Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda." volume="33" year="1864">Theobald 1864</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Streptaxis birmanicus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="birmanicus">Streptaxis birmanicus</taxonomicName>
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[sic] -
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<bibRefCitation author="Richardson, L" journalOrPublisher="Tryonia" pageId="0" pageNumber="39" pagination="1 - 326" refId="B70" refString="Richardson, L, 1988. Streptaxacea: Catalog of species, Part I, Streptaxidae. Tryonia 16: 1 - 326" title="Streptaxacea: Catalog of species, Part I, Streptaxidae." volume="16" year="1988">Richardson 1988</bibRefCitation>
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: 251.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="materials_examined">
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<paragraph pageId="0" pageNumber="39">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="39">
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<typeStatus>Syntype</typeStatus>
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</emphasis>
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NHMUK 1888.12.4.777-779 (3 shells; Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-C Haploptychius blanfordi A syntype NHMUK 1888.12.4.777 - 779 from Pegu with apertural dentition B specimen NHMUK 1906.2.2.338 from Akouktoung, Pegu C specimen NHMUK 1909.3.15.71 from Pegu D Haploptychius burmanicus, syntype NHMUK 1906.2.2.199 from Tongoop, Arakan with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure10" httpUri="https://binary.pensoft.net/fig/710475" pageId="0" pageNumber="39">10A</figureCitation>
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) from Pegu. NHMUK 1909.3.15.71 (1 shell; Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-C Haploptychius blanfordi A syntype NHMUK 1888.12.4.777 - 779 from Pegu with apertural dentition B specimen NHMUK 1906.2.2.338 from Akouktoung, Pegu C specimen NHMUK 1909.3.15.71 from Pegu D Haploptychius burmanicus, syntype NHMUK 1906.2.2.199 from Tongoop, Arakan with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure10" httpUri="https://binary.pensoft.net/fig/710475" pageId="0" pageNumber="39">10C</figureCitation>
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) ex. Godwin-Austen collection. Pegu, Arakan: NHMUK 1909.3.15.93 (3 shells) ex. H. Blanford collection. Akouktoung, Pegu: NHMUK 1906.2.2.338 (4 shells; Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-C Haploptychius blanfordi A syntype NHMUK 1888.12.4.777 - 779 from Pegu with apertural dentition B specimen NHMUK 1906.2.2.338 from Akouktoung, Pegu C specimen NHMUK 1909.3.15.71 from Pegu D Haploptychius burmanicus, syntype NHMUK 1906.2.2.199 from Tongoop, Arakan with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure10" httpUri="https://binary.pensoft.net/fig/710475" pageId="0" pageNumber="39">10B</figureCitation>
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) ex. W.T. Blanford collection. Pegu, Arakan: NHMUK 1909.3.15.93 (3 shells) ex. H. Blanford collection.
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</paragraph>
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<caption doi="10.3897/zookeys.1110.85399.figure10" httpUri="https://binary.pensoft.net/fig/710475" pageId="0" pageNumber="39" start="Figure 10" startId="F10">
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<paragraph pageId="0" pageNumber="39">
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<emphasis bold="true" pageId="0" pageNumber="39">Figure 10.</emphasis>
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<emphasis bold="true" pageId="0" pageNumber="39">A-C</emphasis>
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<taxonomicName baseAuthorityName="Theobald" baseAuthorityYear="1864" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius blanfordi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordi">
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<emphasis italics="true" pageId="0" pageNumber="39">Haploptychius blanfordi</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="39">A</emphasis>
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<typeStatus>syntype</typeStatus>
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NHMUK 1888.12.4.777-779 from Pegu with apertural dentition
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<emphasis bold="true" pageId="0" pageNumber="39">B</emphasis>
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specimen NHMUK 1906.2.2.338 from Akouktoung, Pegu
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<emphasis bold="true" pageId="0" pageNumber="39">C</emphasis>
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specimen NHMUK 1909.3.15.71 from Pegu
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<emphasis bold="true" pageId="0" pageNumber="39">D</emphasis>
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<taxonomicName baseAuthorityName="Blanford" baseAuthorityYear="1865" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius burmanicus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="burmanicus">
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<emphasis italics="true" pageId="0" pageNumber="39">Haploptychius burmanicus</emphasis>
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</taxonomicName>
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,
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<typeStatus>syntype</typeStatus>
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NHMUK 1906.2.2.199 from Tongoop, Arakan with apertural dentition.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="diagnosis">
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<paragraph pageId="0" pageNumber="39">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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<taxonomicName baseAuthorityName="Morlet" baseAuthorityYear="1886" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius fischeri" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="fischeri">
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<emphasis italics="true" pageId="0" pageNumber="39">Haploptychius fischeri</emphasis>
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</taxonomicName>
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(Morlet, 1887) almost shares the same shell form as
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<taxonomicName genus="H." lsidName="H. blanfordi" pageId="0" pageNumber="39" rank="species" species="blanfordi">
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<emphasis italics="true" pageId="0" pageNumber="39">H. blanfordi</emphasis>
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</taxonomicName>
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, but the latter possesses a more depressed shell, penultimate whorl more bulging, aperture wider, longer and subquadrangular shape.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="description">
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<paragraph pageId="0" pageNumber="39">Description.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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Shell oblique-heliciform, white, and translucent; whorls 6; spire convex with distinct suture. Shell surface glossy with fine transverse ridges that diminish below periphery. Embryonic shell ~
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<normalizedToken originalValue="2½">21/2</normalizedToken>
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whorls with smooth surface; following whorls regularly coiled. Shell periphery rounded; last whorl axially deflected. Aperture semi-ovate; peristome discontinuous, thickened, expanded, and reflected. Apertural dentition with one strong parietal and small palatal lamellae. Umbilicus open and deep (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-C Haploptychius blanfordi A syntype NHMUK 1888.12.4.777 - 779 from Pegu with apertural dentition B specimen NHMUK 1906.2.2.338 from Akouktoung, Pegu C specimen NHMUK 1909.3.15.71 from Pegu D Haploptychius burmanicus, syntype NHMUK 1906.2.2.199 from Tongoop, Arakan with apertural dentition." figureDoi="10.3897/zookeys.1110.85399.figure10" httpUri="https://binary.pensoft.net/fig/710475" pageId="0" pageNumber="39">10A-C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="distribution">
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<paragraph pageId="0" pageNumber="39">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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This species has been recorded with a wide distribution range from Arakan [Rakhine State], Pegu [Bago Region], and Shan State in Myanmar. However, the record from Cocos Islands in the southern Indian Ocean by
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="39" refId="B10" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, Including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, Including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen (1908)</bibRefCitation>
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needs to be verified with modern evidence rather than by shell morphology alone.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="39" type="remarks">
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<paragraph pageId="0" pageNumber="39">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="39">
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The taxonomic status of this species is still ambiguous because it is identical in shell morphology to
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" kingdom="Animalia" lsidName="Streptaxis birmanica" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="birmanica">
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|
<emphasis italics="true" pageId="0" pageNumber="39">Streptaxis birmanica</emphasis>
|
|
</taxonomicName>
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|
.
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|
<bibRefCitation author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="238 - 250" refId="B87" refString="Theobald, W, 1864. Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda. Journal of the Asiatic Society of Bengal 33: 238 - 250" title="Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda." volume="33" year="1864">Theobald (1864)</bibRefCitation>
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|
consecutively described
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|
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" kingdom="Animalia" lsidName="Streptaxis blanfordi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordi">
|
|
<emphasis italics="true" pageId="0" pageNumber="39">Streptaxis blanfordi</emphasis>
|
|
</taxonomicName>
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|
and
|
|
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" kingdom="Animalia" lsidName="Streptaxis birmanica" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="birmanica">
|
|
<emphasis italics="true" pageId="0" pageNumber="39">Streptaxis birmanica</emphasis>
|
|
</taxonomicName>
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|
based on specimens from the same geographical area as Pegu. These two nominal species resemble each other in having one parietal and one palatal lamella, whereas they differ in only minor characters of shell shape and size. In this revision, we consider these two nominal species as synonyms. Only the type specimens of
|
|
<taxonomicName genus="H." lsidName="H. blanfordi" pageId="0" pageNumber="39" rank="species" species="blanfordi">
|
|
<emphasis italics="true" pageId="0" pageNumber="39">H. blanfordi</emphasis>
|
|
</taxonomicName>
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|
could be located and examined; we therefore consider
|
|
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" kingdom="Animalia" lsidName="Streptaxis birmanica" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="birmanica">
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|
<emphasis italics="true" pageId="0" pageNumber="39">Streptaxis birmanica</emphasis>
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|
</taxonomicName>
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|
as the junior synonym.
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|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="39">
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|
The original spelling of the species was
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|
<taxonomicName baseAuthorityName="Theobald" baseAuthorityYear="1864" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius blanfordi" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordi">
|
|
<emphasis italics="true" pageId="0" pageNumber="39">Haploptychius blanfordi</emphasis>
|
|
</taxonomicName>
|
|
, which was intentionally modified to
|
|
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius blanfordianus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordianus">
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|
<emphasis italics="true" pageId="0" pageNumber="39">Haploptychius blanfordianus</emphasis>
|
|
</taxonomicName>
|
|
by
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|
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="143 - 177" refId="B80" refString="Stoliczka, F, 1871. Notes on the terrestrial Mollusca from the neighbourhood of Moulmein, with descriptions of new species. Journal of the Asiatic Society of Bengal 40: 143 - 177" title="Notes on the terrestrial Mollusca from the neighbourhood of Moulmein, with descriptions of new species." volume="40" year="1871">Stoliczka (1871)</bibRefCitation>
|
|
but for an unclear reason. This subsequent spelling of
|
|
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius blanfordianus" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="blanfordianus">
|
|
<emphasis italics="true" pageId="0" pageNumber="39">Haploptychius blanfordianus</emphasis>
|
|
</taxonomicName>
|
|
is an unjustified emendation made available with its own authorship and date, and so became a junior objective synonym (ICZN 1999: Arts 32.3, 33.2.3, 50.5).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="39">
|
|
<bibRefCitation author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="238 - 250" refId="B87" refString="Theobald, W, 1864. Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda. Journal of the Asiatic Society of Bengal 33: 238 - 250" title="Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda." volume="33" year="1864">Theobald (1864)</bibRefCitation>
|
|
clearly stated that
|
|
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" kingdom="Animalia" lsidName="Streptaxis birmanica" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="birmanica">
|
|
<emphasis italics="true" pageId="0" pageNumber="39">Streptaxis birmanica</emphasis>
|
|
</taxonomicName>
|
|
W. Blanford, (in MSS.) was described based on a single specimen received from W.T. Blanford. However, we could not locate this single shell as the holotype fixed by monotypy.
|
|
<bibRefCitation author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="39" pagination="238 - 250" refId="B87" refString="Theobald, W, 1864. Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda. Journal of the Asiatic Society of Bengal 33: 238 - 250" title="Notes on the variation of Indian and Burmese Helicidae, with an attempt at their re-arrangement, together with descriptions of new Burmese Gastropoda." volume="33" year="1864">Theobald (1864)</bibRefCitation>
|
|
also stated that the other two shells lacked the palatal lamella and were of a relatively smaller size, which are recognized as distinct varietal entities. Therefore, these two specimens are excluded from the type series of
|
|
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Streptaxis" kingdom="Animalia" lsidName="Streptaxis birmanica" order="Stylommatophora" pageId="0" pageNumber="39" phylum="Mollusca" rank="species" species="birmanica">
|
|
<emphasis italics="true" pageId="0" pageNumber="39">Streptaxis birmanica</emphasis>
|
|
</taxonomicName>
|
|
. The authorship was originally attributed to W.T. Blanford as the manuscript name. However, since Blanford did not write the description or credit the description to W.T. Blanford, the taxon is attributed to Theobald only (ICZN 1999: Art. 50.1.1).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
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</document> |