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<document id="58979AAB56692E4FB61AEE5DC3D90BF9" ID-DOI="10.15298/rusentj.31.1.15" ID-ISSN="0132-8069" ID-Zenodo-Dep="13179620" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722596549374" checkinUser="felipe" docAuthor="Shamshev, I. V." docDate="2022" docId="F15D0A63FFF47D17FEA1FCBB72CD06C1" docLanguage="en" docName="RussEntomolJ.31.1.76-79.pdf" docOrigin="Russian Entomological Journal 31 (1)" docSource="http://dx.doi.org/10.15298/rusentj.31.1.15" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Anomalempis yakutica Shamshev 2022, sp.n." docType="treatment" docVersion="2" lastPageNumber="79" masterDocId="0D64721BFFF57D14FFAAFFDE70700150" masterDocTitle="First record of the genus Anomalempis Melander (Diptera: Empidoidea: Brachystomatidae) from the Palaearctic region (Russia)" masterLastPageNumber="79" masterPageNumber="76" pageNumber="77" updateTime="1722637638130" updateUser="ExternalLinkService">
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<mods:title id="B131B04858BD0F10FA2DD197F04CC062">First record of the genus Anomalempis Melander (Diptera: Empidoidea: Brachystomatidae) from the Palaearctic region (Russia)</mods:title>
</mods:titleInfo>
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<mods:namePart id="5AD6078DCF094AE87DC53028A54EB165">Shamshev, I. V.</mods:namePart>
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<mods:title id="BC81585FE532AF7565F4A420240FF16A">Russian Entomological Journal</mods:title>
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<mods:part id="566687089F29093742EBE3B7170A9840">
<mods:date id="9A912AB040286FA804D0283FBE71B3A1">2022</mods:date>
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<mods:number id="4FA5EF84FB4AD8916D8816F2F475F275">2022-03-31</mods:number>
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<mods:number id="78EBF77FBABE09E484B8A61E0E6CA09B">1</mods:number>
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<mods:url id="68C5926E05E99ED29D107EDCF8B8B652">http://dx.doi.org/10.15298/rusentj.31.1.15</mods:url>
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<mods:classification id="CE301F2A3F5753C690DC1DC8C217835D">journal article</mods:classification>
<mods:identifier id="0D3F7FAD63899C0D98D41CEC29DD8126" type="DOI">10.15298/rusentj.31.1.15</mods:identifier>
<mods:identifier id="9CA36C0AFE10B08E646EE31CD34AEE8E" type="ISSN">0132-8069</mods:identifier>
<mods:identifier id="A1C1D0DE779BA6582B0710E4F3EA5D92" type="Zenodo-Dep">13179620</mods:identifier>
</mods:mods>
<treatment id="F15D0A63FFF47D17FEA1FCBB72CD06C1" LSID="urn:lsid:plazi:treatment:F15D0A63FFF47D17FEA1FCBB72CD06C1" httpUri="http://treatment.plazi.org/id/F15D0A63FFF47D17FEA1FCBB72CD06C1" lastPageId="3" lastPageNumber="79" pageId="1" pageNumber="77">
<subSubSection id="31EEE8FEFFF47D15FEA1FCBB72C7022D" box="[267,695,869,893]" pageId="1" pageNumber="77" type="nomenclature">
<paragraph id="794BBB75FFF47D15FEA1FCBB72C7022D" blockId="1.[189,775,869,1075]" box="[267,695,869,893]" pageId="1" pageNumber="77">
<heading id="22030C19FFF47D15FEA1FCBB72C7022D" box="[267,695,869,893]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="77" reason="8">
<taxonomicName id="BEF4C0F6FFF47D15FEA1FCBB7207022D" authority="Shamshev" authorityName="Shamshev" authorityYear="2022" box="[267,631,869,893]" class="Insecta" family="Empididae" genus="Anomalempis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="77" phylum="Arthropoda" rank="species" species="yakutica" status="sp. nov.">
<emphasis id="4B806767FFF47D15FEA1FCBB718E022D" box="[267,510,869,893]" italics="true" pageId="1" pageNumber="77">Anomalempis yakutica</emphasis>
Shamshev
</taxonomicName>
,
<emphasis id="4B806767FFF47D15FD28FCBB72C7022D" bold="true" box="[642,695,869,893]" pageId="1" pageNumber="77">
<taxonomicNameLabel id="50B3DA1CFFF47D15FD28FCBB72C7022D" box="[642,695,869,893]" pageId="1" pageNumber="77" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="31EEE8FEFFF47D15FE05FC5D726402CA" box="[431,532,898,923]" pageId="1" pageNumber="77" type="description">
<paragraph id="794BBB75FFF47D15FE05FC5D726402CA" blockId="1.[189,775,869,1075]" box="[431,532,898,923]" pageId="1" pageNumber="77">
<figureCitation id="E1CFA7F0FFF47D15FE05FC5D726002CA" box="[431,528,898,923]" captionStart-0="Fig" captionStart-1="Figs 2–5" captionStartId-0="1.[226,258,2017,2036]" captionStartId-1="2.[226,262,1849,1868]" captionTargetBox-0="[211,1387,1099,1995]" captionTargetBox-1="[189,1398,706,1841]" captionTargetId-0="figure-437@1.[189,1399,1099,1996]" captionTargetId-1="figure-422@2.[189,1398,664,1841]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Anomalempis yakutica sp.n., ♂ holotype, lateral view. Рис. 1. Anomalempis yakutica sp.n., ♂ голотип, вид сбоку." captionText-1="Figs 2–5. Anomalempis yakutica Shamshev sp.n., ♂: 2 — hypopygium, lateral view; 3 — internal structures of hypopygium, lateral view; 4 — hypopygium (left cercus and right surstylus removed, setation of epandrium partly omitted), dorsal view; 5 — epandrium and hypandrium, ventral view (setation of epandrium omitted). Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; hyprct — hypoproct; pgt — postgonite lobe; ph — phallus; ph rd — phallic rod; sur — surstylus. Рис. 2–5. Anomalempis yakutica Shamshev sp.n., ♂: 2 — гипопигий, вид сбоку; 3 — внутренние структуры гипопигиЯ, сбоку; 4 — гипопигий (левый церк и правый сурстиль удалены, Щетинки ЭпандриЯ частично не покаЗаны), сверху; 5 — Эпандрий и гипандрий, сниЗу (Щетинки ЭпандриЯ не покаЗаны). СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; hyprct — гипопрокт; pgt — лопасть постгонита; ph — фаллус; ph rd — стержень фаллуса; sur — сурстиль." figureDoi-0="http://doi.org/10.5281/zenodo.13179622" figureDoi-1="http://doi.org/10.5281/zenodo.13179626" httpUri-0="https://zenodo.org/record/13179622/files/figure.png" httpUri-1="https://zenodo.org/record/13179626/files/figure.png" pageId="1" pageNumber="77">Figs 1–5</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="31EEE8FEFFF47D15FF48FC76753D002E" pageId="1" pageNumber="77" type="materials_examined">
<paragraph id="794BBB75FFF47D15FF48FC7672000553" blockId="1.[189,775,869,1075]" pageId="1" pageNumber="77">
<collectionCode id="1FE523B0FFF47D15FF48FC7671E602EB" box="[226,406,936,955]" pageId="1" pageNumber="77">TYPE MATERIAL</collectionCode>
.
<typeStatus id="A64F05D7FFF47D15FE09FC76718602EB" box="[419,502,936,955]" pageId="1" pageNumber="77" type="holotype">Holotype</typeStatus>
,
<specimenCount id="6FF270FCFFF47D15FDAFFC7B726902EB" box="[517,537,933,955]" count="1" pageId="1" pageNumber="77" type="male">♂</specimenCount>
, [
<collectingCountry id="01E3FBE5FFF47D15FD84FC76721C02EB" box="[558,620,936,955]" name="Russia" pageId="1" pageNumber="77">Russia</collectingCountry>
] Nizhnie Kresty [now Chersky,
<geoCoordinate id="1CC0DDB2FFF47D15FEEDFC1E71E10283" box="[327,401,960,979]" degrees="68" direction="north" minutes="46" orientation="latitude" pageId="1" pageNumber="77" precision="925" value="68.76667">68°46´N</geoCoordinate>
<geoCoordinate id="1CC0DDB2FFF47D15FE33FC1E719E0283" box="[409,494,960,979]" degrees="161" direction="east" minutes="20" orientation="longitude" pageId="1" pageNumber="77" precision="925" value="161.33333">161°20´E</geoCoordinate>
] / Kolyma, Yakutia/ Gorodkov 25.vi.[1]963; shore of Kolyma River;
<taxonomicName id="BEF4C0F6FFF47D15FE57FC06720202BB" authorityName="Melander" authorityYear="1928" box="[509,626,984,1003]" class="Insecta" family="Empididae" genus="Anomalempis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="4B806767FFF47D15FE57FC06720202BB" box="[509,626,984,1003]" italics="true" pageId="1" pageNumber="77">Anomalempis</emphasis>
</taxonomicName>
/
<taxonomicName id="BEF4C0F6FFF47D15FDD6FC0672B502BB" authorityName="Shamshev" authorityYear="2022" box="[636,709,984,1003]" class="Insecta" family="Empididae" genus="Anomalempis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="77" phylum="Arthropoda" rank="species" species="yakutica">
<emphasis id="4B806767FFF47D15FDD6FC0672B502BB" box="[636,709,984,1003]" italics="true" pageId="1" pageNumber="77">yakutica</emphasis>
</taxonomicName>
/ Shamshev
<taxonomicNameLabel id="50B3DA1CFFF47D15FF41FC2E71650553" box="[235,277,1008,1027]" pageId="1" pageNumber="77" rank="species">sp.n.</taxonomicNameLabel>
[red label] (
<collectionCode id="1FE523B0FFF47D15FE28FC2E71D40553" box="[386,420,1008,1027]" pageId="1" pageNumber="77">INS</collectionCode>
_DIP_0000622,
<collectionCode id="1FE523B0FFF47D15FD9CFC2E72160553" box="[566,614,1008,1027]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="77">ZISP</collectionCode>
).
</paragraph>
<paragraph id="794BBB75FFF47D15FF48FBD6753D002E" blockId="1.[189,775,869,1075]" lastBlockId="1.[812,1399,315,1076]" pageId="1" pageNumber="77">
<materialsCitation id="C99CB128FFF47D15FF48FBD673D8001F" collectingDate="1976-06-29" collectionCode="V" country="Russia" county="Republic" location="Indigirka River" municipality="Yakutia" pageId="1" pageNumber="77" specimenCount="1" specimenCount-male="1" stateProvince="Sakha" typeStatus="paratype">
<typeStatus id="A64F05D7FFF47D15FF48FBD67149054B" box="[226,313,1032,1051]" pageId="1" pageNumber="77" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="01E3FBE5FFF47D15FEE8FBD6710B054B" box="[322,379,1032,1051]" name="Russia" pageId="1" pageNumber="77">Russia</collectingCountry>
(
<collectingCounty id="902AC3F9FFF47D15FE2DFBD671A6054B" box="[391,470,1032,1051]" pageId="1" pageNumber="77">Republic</collectingCounty>
of
<collectingRegion id="BB307597FFF47D15FE5EFBD67259054B" box="[500,553,1032,1051]" country="Russia" name="Sakha" pageId="1" pageNumber="77">Sakha</collectingRegion>
(
<collectingMunicipality id="992F210FFFF47D15FD9EFBD6720D054B" box="[564,637,1032,1051]" pageId="1" pageNumber="77">Yakutia</collectingMunicipality>
)):
<specimenCount id="6FF270FCFFF47D15FD3EFBD672C9054B" box="[660,697,1029,1051]" count="1" pageId="1" pageNumber="77" type="male">1 ♂</specimenCount>
,
<location id="7C2BEDAEFFF47D15FD6EFBD6717B0563" LSID="urn:lsid:plazi:treatment:F15D0A63FFF47D17FEA1FCBB72CD06C1:7C2BEDAEFFF47D15FD6EFBD6717B0563" country="Russia" county="Republic" municipality="Yakutia" name="Indigirka River" pageId="1" pageNumber="77" stateProvince="Sakha">Indigirka River</location>
, lower flow of
<location id="7C2BEDAEFFF47D15FE30FBFE722A0563" LSID="urn:lsid:plazi:treatment:F15D0A63FFF47D17FEA1FCBB72CD06C1:7C2BEDAEFFF47D15FE30FBFE722A0563" box="[410,602,1056,1075]" country="Russia" county="Republic" municipality="Yakutia" name="Ystan-Yuryakh River" pageId="1" pageNumber="77" stateProvince="Sakha">Ystan-Yuryakh River</location>
,
<location id="7C2BEDAEFFF47D15FDCFFBFE73730563" LSID="urn:lsid:plazi:treatment:F15D0A63FFF47D17FEA1FCBB72CD06C1:7C2BEDAEFFF47D15FDCFFBFE73730563" box="[613,771,1056,1075]" country="Russia" county="Republic" municipality="Yakutia" name="Momskiy District" pageId="1" pageNumber="77" stateProvince="Sakha">Momskiy District</location>
,
<date id="0D4A9DB5FFF47D15FC86FEE573FB001E" box="[812,907,315,334]" pageId="1" pageNumber="77" value="1976-06-29">
<collectingDate id="1D0E645DFFF47D15FC86FEE573FB001E" box="[812,907,315,334]" pageId="1" pageNumber="77" value="1976-06-29">29.vi.1976</collectingDate>
</date>
,
<collectionCode id="1FE523B0FFF47D15FC3CFEE273D8001F" box="[918,936,316,335]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="77" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="C99CB128FFF47D15FC19FEE574A50037" collectingDate="1976-07-01" collectionCode="V" country="Russia" county="Republic" location="Kovalev" municipality="Yakutia" pageId="1" pageNumber="77" specimenCount="1" specimenCount-female="1" stateProvince="Sakha" typeStatus="paratype">
<location id="7C2BEDAEFFF47D15FC19FEE5738C001E" LSID="urn:lsid:plazi:treatment:F15D0A63FFF47D17FEA1FCBB72CD06C1:7C2BEDAEFFF47D15FC19FEE5738C001E" box="[947,1020,315,334]" country="Russia" name="Kovalev" pageId="1" pageNumber="77">Kovalev</location>
[terminalia dissected, in microvial pinned with specimen];
<specimenCount id="6FF270FCFFF47D15FC16FE8D73A80037" box="[956,984,337,359]" count="1" pageId="1" pageNumber="77" type="female">1 ♀</specimenCount>
, same locality,
<date id="0D4A9DB5FFF47D15FBCAFE8D74C90036" box="[1120,1209,339,358]" pageId="1" pageNumber="77" value="1976-07-01">
<collectingDate id="1D0E645DFFF47D15FBCAFE8D74C90036" box="[1120,1209,339,358]" pageId="1" pageNumber="77" value="1976-07-01">1.vii.1976</collectingDate>
</date>
,
<collectionCode id="1FE523B0FFF47D15FB69FE8A74A50037" box="[1219,1237,340,359]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="77" type="Museum">V</collectionCode>
</materialsCitation>
. Kovalev; slope of hill larch forest with steppe areas (both
<typeStatus id="A64F05D7FFF47D15FB21FEB274AE002F" box="[1163,1246,364,383]" pageId="1" pageNumber="77" type="paratype">paratypes</typeStatus>
in
<collectionCode id="1FE523B0FFF47D15FB56FEB57533002E" box="[1276,1347,363,382]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="77" type="Museum">ZMMU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="31EEE8FEFFF47D15FCF8FE5D739600B9" pageId="1" pageNumber="77" type="diagnosis">
<paragraph id="794BBB75FFF47D15FCF8FE5D739600B9" blockId="1.[812,1399,315,1076]" pageId="1" pageNumber="77">
DIAGNOSIS. Mid-sized (body about
<quantity id="BE0C1690FFF47D15FB7EFE5D756400C8" box="[1236,1300,387,408]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="77" unit="mm" value="4.0">4 mm</quantity>
), greyish, pale setose flies; mesonotum and abdomen greyish pruinose; wings whitish in male and hyaline in female, with inconspicuous pterostigma.
</paragraph>
</subSubSection>
<subSubSection id="31EEE8FEFFF47D17FCF8FE3072D50610" lastPageId="3" lastPageNumber="79" pageId="1" pageNumber="77" type="description">
<paragraph id="794BBB75FFF47D15FCF8FE3075270299" blockId="1.[812,1399,315,1076]" pageId="1" pageNumber="77">
DESCRIPTION.
<emphasis id="4B806767FFF47D15FC50FE30745D0353" bold="true" box="[1018,1069,494,515]" pageId="1" pageNumber="77">Male</emphasis>
(
<figureCitation id="E1CFA7F0FFF47D15FB97FE3074060353" box="[1085,1142,494,516]" captionStart="Fig" captionStartId="1.[226,258,2017,2036]" captionTargetBox="[211,1387,1099,1995]" captionTargetId="figure-437@1.[189,1399,1099,1996]" captionTargetPageId="1" captionText="Fig. 1. Anomalempis yakutica sp.n., ♂ holotype, lateral view. Рис. 1. Anomalempis yakutica sp.n., ♂ голотип, вид сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179622" httpUri="https://zenodo.org/record/13179622/files/figure.png" pageId="1" pageNumber="77">Fig. 1</figureCitation>
). Body 4.0 mm, wing 4.0 mm (
<typeStatus id="A64F05D7FFF47D15FCF4FDD773C6034F" box="[862,950,521,543]" pageId="1" pageNumber="77" type="holotype">holotype</typeStatus>
). Head dark brown in ground-colour, with pale setation, occiput densely greyish pruinose. Eyes holoptic, with upper ommatidia enlarged, border between larger and smaller ommatidia distinct. Frons represented by small triangular space just below ocellar triangle and somewhat larger space above antennae, bare, greyish pruinose. Face parallel-sided, bare, densely greyish pruinose. Ocellar triangle prominent, with 2 pairs of long setae. Upper postocular setae nearly as long as ocellar setae; postvertical setae much shorter than upper postocular setae, fine; otherwise occiput covered with numerous similar fine setae (except above neck). Antenna with scape and pedicel brownish, postpedicel and stylus black; scape short, slightly shorter than globular pedicel, both with short setae; postpedicel conical, nearly 2.5 times longer than basal width; stylus slightly shorter than postpedicel, apex bare, peg-like. Proboscis short; length of labrum nearly half of eye height; palpus dirty yellow, with moderately long, fine pale setae.
</paragraph>
<paragraph id="794BBB75FFF47D16FCF8FC10737703C0" blockId="1.[812,1399,315,1076]" lastBlockId="2.[189,775,315,657]" lastPageId="2" lastPageNumber="78" pageId="1" pageNumber="77">Thorax dark in ground-colour, densely greyish pruinose on prothoracic sclerites and mesopleuron, sparser pruinose on mesonotum, mesoscutum without vittae (greyish viewed anteriorly; faintly pruinose, subshiny viewed dorsally); with pale setation (stronger setae rather pale yellowish). Proepisternum with several moderately long, fine setae on lower section; upper proepisternum in front of spiracle with 3–4 short, fine setae. Prosternum bare. Antepronotum with numerous short to moderately long setae. Postpronotal lobe with 3 long and 2 short setae. Mesonotum with rather hardly differentiat- ed setation; numerous short setae on presutural supra-alar area, 3–4 long and some short to moderately long finer setae on postsutural supra-alar area, notopleuron with 4–5 long and several shorter setae, 1 long and 1 short postalars, 8–10 scutellars; dorsocentrals mostly arranged in 2–3 irregular rows, uniserial just before scutellum, of different lengths on presutural space (intermixed short to moderately long setae),</paragraph>
<caption id="2D8BEBFDFFF47D15FF48F83F728C0941" ID-DOI="http://doi.org/10.5281/zenodo.13179622" ID-Zenodo-Dep="13179622" httpUri="https://zenodo.org/record/13179622/files/figure.png" pageId="1" pageNumber="77" startId="1.[226,258,2017,2036]" targetBox="[211,1387,1099,1995]" targetPageId="1" targetType="figure">
<paragraph id="794BBB75FFF47D15FF48F83F728C0941" blockId="1.[226,774,2015,2065]" pageId="1" pageNumber="77">
Fig. 1.
<taxonomicName id="BEF4C0F6FFF47D15FE88F83F719706A4" authorityName="Shamshev" authorityYear="2022" box="[290,487,2017,2036]" class="Insecta" family="Empididae" genus="Anomalempis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="77" phylum="Arthropoda" rank="species" species="yakutica" status="sp. nov.">
<emphasis id="4B806767FFF47D15FE88F83F719706A4" box="[290,487,2017,2036]" italics="true" pageId="1" pageNumber="77">Anomalempis yakutica</emphasis>
</taxonomicName>
<emphasis id="4B806767FFF47D15FE44F83F726906A4" bold="true" box="[494,537,2017,2036]" pageId="1" pageNumber="77">
<taxonomicNameLabel id="50B3DA1CFFF47D15FE44F83F726906A4" box="[494,537,2017,2036]" pageId="1" pageNumber="77" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂ holotype, lateral view. Рис. 1.
<taxonomicName id="BEF4C0F6FFF47D15FE8DF823719C0940" authorityName="Shamshev" authorityYear="2022" box="[295,492,2045,2064]" class="Insecta" family="Empididae" genus="Anomalempis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="77" phylum="Arthropoda" rank="species" species="yakutica" status="sp. nov.">
<emphasis id="4B806767FFF47D15FE8DF823719C0940" box="[295,492,2045,2064]" italics="true" pageId="1" pageNumber="77">Anomalempis yakutica</emphasis>
</taxonomicName>
<emphasis id="4B806767FFF47D15FE59F823726E0940" bold="true" box="[499,542,2045,2064]" pageId="1" pageNumber="77">
<taxonomicNameLabel id="50B3DA1CFFF47D15FE59F823726E0940" box="[499,542,2045,2064]" pageId="1" pageNumber="77" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂ голотип, вид сбоку.
</paragraph>
</caption>
<paragraph id="794BBB75FFF77D16FDC3FD7C752600F1" blockId="2.[248,666,674,718]" lastBlockId="2.[812,1399,315,657]" pageId="2" pageNumber="78">cerc epand becoming more uniform and longer toward scutellum; acrostichal setae arranged in 2 irregular rows, nearly as long as presutural dorsocentrals, absent on prescutellar depression. Laterotergite bare. Anterior and posterior spiracles dark.</paragraph>
<paragraph id="794BBB75FFF77D16FCF8FE787535039A" blockId="2.[812,1399,315,657]" lastBlockId="2.[945,1349,688,718]" pageId="2" pageNumber="78">Legs almost uniformly brownish, only extreme apex of coxae and “knees” yellowish to brownish yellow; coxae densely greyish pruinose, remaining parts subshiny; all podomeres only pale setose (some stronger setae somewhat darker), unmodified. Fore femur with complete rows of moderately long anteroventral (somewhat stronger on subapical part) and longer, uniformly fine posteroventral setae. Mid femur with complete rows of short, strong anteroventral and somewhat longer posteroventral setae. Hind femur with complete row of hyprct ph rd</paragraph>
<caption id="2D8BEBFDFFF77D16FF48F8E772740940" ID-DOI="http://doi.org/10.5281/zenodo.13179626" ID-Zenodo-Dep="13179626" httpUri="https://zenodo.org/record/13179626/files/figure.png" pageId="2" pageNumber="78" startId="2.[226,262,1849,1868]" targetBox="[189,1398,706,1841]" targetPageId="2" targetType="figure">
<paragraph id="794BBB75FFF77D16FF48F8E7750706C4" blockId="2.[188,1399,1847,2064]" pageId="2" pageNumber="78">
Figs 2–5.
<taxonomicName id="BEF4C0F6FFF77D16FE9CF8E77222061C" authority="Shamshev" authorityName="Shamshev" authorityYear="2022" box="[310,594,1849,1868]" class="Insecta" family="Empididae" genus="Anomalempis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="species" species="yakutica" status="sp. nov.">
<emphasis id="4B806767FFF77D16FE9CF8E77185061C" box="[310,501,1849,1868]" italics="true" pageId="2" pageNumber="78">Anomalempis yakutica</emphasis>
Shamshev
</taxonomicName>
<emphasis id="4B806767FFF77D16FDFDF8E772F1061C" bold="true" box="[599,641,1849,1868]" pageId="2" pageNumber="78">
<taxonomicNameLabel id="50B3DA1CFFF77D16FDFDF8E772F1061C" box="[599,641,1849,1868]" pageId="2" pageNumber="78" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂: 2 — hypopygium, lateral view; 3 — internal structures of hypopygium, lateral view; 4 — hypopygium (left cercus and right surstylus removed, setation of epandrium partly omitted), dorsal view; 5 — epandrium and hypandrium, ventral view (setation of epandrium omitted). Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; hyprct — hypoproct; pgt — postgonite lobe; ph — phallus; ph rd — phallic rod; sur — surstylus.
</paragraph>
<paragraph id="794BBB75FFF77D16FF48F84372740940" blockId="2.[188,1399,1847,2064]" pageId="2" pageNumber="78">
Рис. 2–5.
<taxonomicName id="BEF4C0F6FFF77D16FE96F843721306E0" authority="Shamshev" authorityName="Shamshev" authorityYear="2022" box="[316,611,1949,1968]" class="Insecta" family="Empididae" genus="Anomalempis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="species" species="yakutica" status="sp. nov.">
<emphasis id="4B806767FFF77D16FE96F843727306E0" box="[316,515,1949,1968]" italics="true" pageId="2" pageNumber="78">Anomalempis yakutica</emphasis>
Shamshev
</taxonomicName>
<emphasis id="4B806767FFF77D16FDC3F84372E506E0" bold="true" box="[617,661,1949,1968]" pageId="2" pageNumber="78">
<taxonomicNameLabel id="50B3DA1CFFF77D16FDC3F84372E506E0" box="[617,661,1949,1968]" pageId="2" pageNumber="78" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, ♂: 2 — гипопигий, вид сбоку; 3 — внутренние структуры гипопигиЯ, сбоку; 4 — гипопигий (левый церк и правый сурстиль удалены, Щетинки ЭпандриЯ частично не покаЗаны), сверху; 5 — Эпандрий и гипандрий, сниЗу (Щетинки ЭпандриЯ не покаЗаны). СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; hyprct — гипопрокт; pgt — лопасть постгонита; ph — фаллус; ph rd — стержень фаллуса; sur — сурстиль.
</paragraph>
</caption>
<paragraph id="794BBB75FFF67D17FF17FEE5721400A1" blockId="3.[189,776,315,1937]" pageId="3" pageNumber="79">short, strong anteroventral setae, some longer setae dorsally; all femora with 2–3 short, strong setae near apex anteriorly. Fore tibia with 2–3, mid tibia with 3–4, hind tibia with 5–6 short anterodorsal and posterodorsal setae (number, position and robustness variable). Mid and hind basitarsi with some stronger setae ventrally (longer closer to base), otherwise tarsomeres covered with short, simple setae.</paragraph>
<paragraph id="794BBB75FFF67D17FF48FE2873760397" blockId="3.[189,776,315,1937]" pageId="3" pageNumber="79">
Wing membrane somewhat whitish; basal costal seta absent, veins brownish yellow to yellowish, bare (except costa); pterostigma inconspicuous; anal lobe well developed, axillary incision slightly more than 90°. Costa circumambient; Sc complete; R
<subScript id="E570B930FFF67D17FE84FDAE7133032C" attach="left" box="[302,323,624,636]" fontSize="5" pageId="3" pageNumber="79">4+5</subScript>
unbranched; cell dm moderately long, produced apically; cell cua long, about third of wing length, rounded apically, CuA+CuP absent; cell bm long, only slightly shorter than cell cua. Squama pale coloured and fringed. Halter pale.
</paragraph>
<paragraph id="794BBB75FFF67D17FF48FD1572330237" blockId="3.[189,776,315,1937]" pageId="3" pageNumber="79">Abdomen dark brown in ground-colour, densely uniformly light grey pruinose; covered with numerous, long, pale fine setae; segment 8 with tergite and sternite separated; tergite 8 very short, about 5X shorter than tergite 7, entire, posterior margin slightly concave mid-dorsally; sternite 8 about 1.5X longer than tergite 8, entire.</paragraph>
<paragraph id="794BBB75FFF67D17FF48FCB572A60441" blockId="3.[189,776,315,1937]" pageId="3" pageNumber="79">
Terminalia (
<figureCitation id="E1CFA7F0FFF67D17FEF1FCB571C402D1" box="[347,436,875,897]" captionStart="Figs 2–5" captionStartId="2.[226,262,1849,1868]" captionTargetBox="[189,1398,706,1841]" captionTargetId="figure-422@2.[189,1398,664,1841]" captionTargetPageId="2" captionText="Figs 2–5. Anomalempis yakutica Shamshev sp.n., ♂: 2 — hypopygium, lateral view; 3 — internal structures of hypopygium, lateral view; 4 — hypopygium (left cercus and right surstylus removed, setation of epandrium partly omitted), dorsal view; 5 — epandrium and hypandrium, ventral view (setation of epandrium omitted). Abbreviations: cerc — cercus; ej apod — ejaculatory apodeme; epand — epandrium; goncx apod — gonocoxal apodeme; hypd — hypandrium; hyprct — hypoproct; pgt — postgonite lobe; ph — phallus; ph rd — phallic rod; sur — surstylus. Рис. 2–5. Anomalempis yakutica Shamshev sp.n., ♂: 2 — гипопигий, вид сбоку; 3 — внутренние структуры гипопигиЯ, сбоку; 4 — гипопигий (левый церк и правый сурстиль удалены, Щетинки ЭпандриЯ частично не покаЗаны), сверху; 5 — Эпандрий и гипандрий, сниЗу (Щетинки ЭпандриЯ не покаЗаны). СокраЩениЯ: cerc — церк; ej apod — ЭЯкулЯторнаЯ аподема; epand — Эпандрий; goncx apod — гонококсальнаЯ аподема; hypd — гипандрий; hyprct — гипопрокт; pgt — лопасть постгонита; ph — фаллус; ph rd — стержень фаллуса; sur — сурстиль." figureDoi="http://doi.org/10.5281/zenodo.13179626" httpUri="https://zenodo.org/record/13179626/files/figure.png" pageId="3" pageNumber="79">Figs 2–5</figureCitation>
) with epandrium and hypandrium brown and subshiny, remaining sclerites pale brown; epandrium, hypandrium and cerci pale setose. Epandrium with broad epandrial bridge, fused with hypandrium anteriorly; with numerous moderately long to long setae apically. Surstylus present, trilobate, lobes hook-like, fused at base, bare; upper and middle lobes of subequal lengths, lower lobe longer. Hypandrium extended beyond apex of epandrium, with numerous long setae; upcurved apically, with rounded and truncate apex; gonocoxal apodeme long, narrow. Postgonite slender, digitiform, sinuate. Phallus slender, arched beyond epandrium, with pair of short dorsal rods near middle, evenly arched and partly membranous subapically; ejaculatory apodeme narrow, mostly flattened, articulated at base of phallus. Cerci articulated with epandrium; cercus long, digitiform, rounded apically, covered with short fine setae.
</paragraph>
<paragraph id="794BBB75FFF67D17FF48FAC8720C074C" blockId="3.[189,776,315,1937]" pageId="3" pageNumber="79">
<emphasis id="4B806767FFF67D17FF48FAC87143047B" bold="true" box="[226,307,1302,1323]" pageId="3" pageNumber="79">Female.</emphasis>
Body
<quantity id="BE0C1690FFF67D17FED3FAC871B7047B" box="[377,455,1302,1323]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="79" unit="mm" value="4.1">4.1 mm</quantity>
, wing 4.0 mm. Similar to male, except as follows: eyes dichoptic, ommatidia equally small; frons narrow on upper part (nearly as broad as anterior ocellus), broadened toward antennae, bare, mostly densely greyish pruinose, narrowly shiny laterally; generally shorter setose, especially on occiput, mesonotum and abdomen; wing membrane rather hyaline; abdomen truncate (terminalia not dissected), segments 8, 9+10 and cerci retracted into segment 7; tergite 7 subshiny posteriorly, with fringe of short, fine, dense yellowish setae along posterior margin.
</paragraph>
<paragraph id="794BBB75FFF67D17FF48F9FF72D50610" blockId="3.[189,776,315,1937]" pageId="3" pageNumber="79">
DIFFERENTIAL DIAGNOSIS. The new species differs from both North American species primarily by pale mesonotal setae (
<emphasis id="4B806767FFF67D17FF51F989717F073C" box="[251,271,1623,1644]" italics="true" pageId="3" pageNumber="79">vs</emphasis>
. black). In addition,
<taxonomicName id="BEF4C0F6FFF67D17FE70F9887236073B" authorityName="Shamshev" authorityYear="2022" box="[474,582,1622,1643]" class="Insecta" family="Empididae" genus="Anomalempis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="79" phylum="Arthropoda" rank="species" species="yakutica" status="sp.n.">
<emphasis id="4B806767FFF67D17FE70F9887199073B" box="[474,489,1622,1643]" italics="true" pageId="3" pageNumber="79">A</emphasis>
.
<emphasis id="4B806767FFF67D17FE5CF9887236073B" box="[502,582,1622,1643]" italics="true" pageId="3" pageNumber="79">yakutica</emphasis>
</taxonomicName>
<emphasis id="4B806767FFF67D17FDE7F988720D073B" bold="true" box="[589,637,1622,1643]" pageId="3" pageNumber="79">
<taxonomicNameLabel id="50B3DA1CFFF67D17FDE7F988720D073B" box="[589,637,1622,1643]" pageId="3" pageNumber="79" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="BEF4C0F6FFF67D17FE95F9AF71ED07D6" authorityName="Melander" authorityYear="1945" box="[319,413,1649,1670]" class="Insecta" family="Empididae" genus="Anomalempis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="79" phylum="Arthropoda" rank="species" species="archon">
<emphasis id="4B806767FFF67D17FE95F9AF713E07D6" box="[319,334,1649,1670]" italics="true" pageId="3" pageNumber="79">A</emphasis>
.
<emphasis id="4B806767FFF67D17FEF0F9AF71ED07D6" box="[346,413,1649,1670]" italics="true" pageId="3" pageNumber="79">archon</emphasis>
</taxonomicName>
by the inconspicuous pterostigma (
<emphasis id="4B806767FFF67D17FD47F9AF737107D6" box="[749,769,1649,1670]" italics="true" pageId="3" pageNumber="79">vs</emphasis>
. pale brown), densely pruinose abdomen (
<emphasis id="4B806767FFF67D17FDE9F952722707F1" box="[579,599,1676,1697]" italics="true" pageId="3" pageNumber="79">vs</emphasis>
. subshiny) as well as by some distinctive details of the male terminalia, including primarily trilobate surstylus (
<emphasis id="4B806767FFF67D17FE77F91F71810786" box="[477,497,1729,1750]" italics="true" pageId="3" pageNumber="79">vs</emphasis>
. bilobate) [
<bibRefCitation id="1D65C684FFF67D17FDC9F91F73720786" author="Melander A. L." box="[611,770,1728,1751]" pageId="3" pageNumber="79" pagination="79 - 87" refId="ref3001" refString="Melander A. L. 1945. Ten new species of Empididae (Diptera) // Psyche. Vol. 52. P. 79 - 87. https: // doi. org / 10.1155 / 1945 / 27183" type="book chapter" year="1945">Melander, 1945</bibRefCitation>
;
<bibRefCitation id="1D65C684FFF67D17FF17F905719907A1" author="Sinclair B. J. &amp; Cumming J. M." box="[189,489,1755,1777]" pageId="3" pageNumber="79" pagination="1 - 170" refId="ref3105" refString="Sinclair B. J., Cumming J. M. 2006. The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera) // Zootaxa. Vol. 1180. P. 1 - 170." type="journal article" year="2006">Sinclair, Cumming, 2006: 160</bibRefCitation>
, figs 332, 333; B.J. Sinclair, personal communication].
<taxonomicName id="BEF4C0F6FFF67D17FE16F92872E4065C" authorityName="Melander" authorityYear="1928" box="[444,660,1782,1804]" class="Insecta" family="Empididae" genus="Anomalempis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="79" phylum="Arthropoda" rank="species" species="tacomae">
<emphasis id="4B806767FFF67D17FE16F92872E4065C" box="[444,660,1782,1804]" italics="true" pageId="3" pageNumber="79">Anomalempis tacomae</emphasis>
</taxonomicName>
(known after a female only) is smaller (body length
<quantity id="BE0C1690FFF67D17FDE5F8CE72EC0676" box="[591,668,1808,1830]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="79" unit="mm" value="2.5">2.5 mm</quantity>
), has black antennae and polished abdomen [
<bibRefCitation id="1D65C684FFF67D17FE56F8F572EB0610" author="Melander A. L." box="[508,667,1835,1857]" pageId="3" pageNumber="79" pagination="1 - 434" refId="ref2958" refString="Melander A. L. 1928. Diptera, Fam. Empididae // P. Wytsman (ed.). Genera Insectorum. Fasc. 185 (1927). Bruxelles: Louis Desmet- Verteneuil. P. 1 - 434." type="book chapter" year="1928">Melander, 1928</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="31EEE8FEFFF67D17FF48F89872880627" pageId="3" pageNumber="79" type="etymology">
<paragraph id="794BBB75FFF67D17FF48F89872880627" blockId="3.[189,776,315,1937]" pageId="3" pageNumber="79">
ETYMOLOGY. The specific epithet refers to the region of origin of the new species, Republic of
<collectingRegion id="BB307597FFF67D17FDF9F8BF72FE0627" box="[595,654,1889,1911]" country="Russia" name="Sakha" pageId="3" pageNumber="79">Sakha</collectingRegion>
(Yakutia).
</paragraph>
</subSubSection>
<subSubSection id="31EEE8FEFFF67D17FF48F8A572CD06C1" box="[226,701,1915,1937]" pageId="3" pageNumber="79" type="distribution">
<paragraph id="794BBB75FFF67D17FF48F8A572CD06C1" blockId="3.[189,776,315,1937]" box="[226,701,1915,1937]" pageId="3" pageNumber="79">
DISTRIBUTION. Palaearctic:
<collectingCountry id="01E3FBE5FFF67D17FDBBF8A5722106C1" box="[529,593,1915,1937]" name="Russia" pageId="3" pageNumber="79">Russia</collectingCountry>
(Yakutia).
</paragraph>
</subSubSection>
</treatment>
</document>
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