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<mods:title id="612DDF453F96BFAEFA802EED9A5AD8E4">Redescription of two ghost shrimps (Decapoda: Axiidea: Callianassidae) from the Middle Miocene of the Central Paratethys: systematics, intraspecific variation, and in situ preservation</mods:title>
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<paragraph id="582C36E6FFB6FFD4FF5E83F21B366BBC" blockId="3.[151,566,221,247]" box="[151,566,221,247]" pageId="3" pageNumber="4">
<heading id="0364818AFFB6FFD4FF5E83F21B366BBC" bold="true" box="[151,566,221,247]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="6AE7EAF4FFB6FFD4FF5E83F21B366BBC" bold="true" box="[151,566,221,247]" pageId="3" pageNumber="4">
Genus
<taxonomicName id="9F934D65FFB6FFD4FF2783F21B366BBC" authority="Sakai, 1988" authorityName="Sakai" authorityYear="1988" box="[238,566,221,247]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FF2783F218A06BBC" bold="true" box="[238,416,221,247]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
<bibRefCitation id="3C024B17FFB6FFD4FE6E83F21B366BBC" author="Sakai" box="[423,566,221,247]" pageId="3" pageNumber="4" refString="Sakai, K. (1988) A new genus and five new species of Callianassidae (Crustacea: Decapoda: Thalassinidea) from Northern Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 5 (1), 51 - 69." type="journal article" year="1988">Sakai, 1988</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="582C36E6FFB6FFD4FF5E820B1A856A72" blockId="3.[151,901,291,314]" box="[151,901,291,314]" pageId="3" pageNumber="4">
<emphasis id="6AE7EAF4FFB6FFD4FF5E820B18226A71" bold="true" box="[151,290,291,314]" pageId="3" pageNumber="4">
<typeStatus id="87288844FFB6FFD4FF5E820B19CC6A72" box="[151,204,292,313]" pageId="3" pageNumber="4">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="9F934D65FFB6FFD4FEE0820B1B8F6A72" authority="Sakai, 1988" authorityName="Sakai" authorityYear="1988" box="[297,655,291,314]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="horneri">
<emphasis id="6AE7EAF4FFB6FFD4FEE0820B1B116A71" box="[297,529,292,314]" italics="true" pageId="3" pageNumber="4">Neocallichirus horneri</emphasis>
<bibRefCitation id="3C024B17FFB6FFD4FDDE820C1B8F6A72" author="Sakai" box="[535,655,291,313]" pageId="3" pageNumber="4" refString="Sakai, K. (1988) A new genus and five new species of Callianassidae (Crustacea: Decapoda: Thalassinidea) from Northern Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 5 (1), 51 - 69." type="journal article" year="1988">Sakai, 1988</bibRefCitation>
</taxonomicName>
by original designation.
</paragraph>
</subSubSection>
<subSubSection id="1089656DFFB6FFD4FF5E8248180B6DFA" pageId="3" pageNumber="4" type="discussion">
<paragraph id="582C36E6FFB6FFD4FF5E824819F7683D" blockId="3.[151,1437,358,1570]" pageId="3" pageNumber="4">
<emphasis id="6AE7EAF4FFB6FFD4FF5E8248180B6A34" bold="true" box="[151,267,359,383]" pageId="3" pageNumber="4">Remarks.</emphasis>
The genus was erected by
<bibRefCitation id="3C024B17FFB6FFD4FDF182491BC76A35" author="Sakai" box="[568,711,358,382]" pageId="3" pageNumber="4" refString="Sakai, K. (1988) A new genus and five new species of Callianassidae (Crustacea: Decapoda: Thalassinidea) from Northern Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 5 (1), 51 - 69." type="journal article" year="1988">Sakai (1988)</bibRefCitation>
to accomodate callianassid forms characterized by the following features: carapace with well-defined dorsal oval, but without rostral carina and cardiac prominence; 3rd maxilliped without exopod, endopod subpediform, propodus expanded, over 3 times broader than dactylus; uropodal endopod broadened posteriorly; telson about as long as broad, about as long as uropods. These characters are, however, of very poor fossilization potential, and as
<bibRefCitation id="3C024B17FFB6FFD4FD9382D91BEA6945" author="Sakai" box="[602,746,502,526]" pageId="3" pageNumber="4" refString="Sakai, K. (1988) A new genus and five new species of Callianassidae (Crustacea: Decapoda: Thalassinidea) from Northern Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 5 (1), 51 - 69." type="journal article" year="1988">Sakai (1988)</bibRefCitation>
stated, the form of the 3rd maxilliped alone is not specific to
<taxonomicName id="9F934D65FFB6FFD4FF5E8134183C6978" box="[151,316,539,563]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FF5E8134183C6978" box="[151,316,539,563]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
</taxonomicName>
, but is similar in several other genera, and therefore always a combination of characters has to be used for generic assignment. Later,
<bibRefCitation id="3C024B17FFB6FFD4FDFC81111A60691D" author="Manning" box="[565,864,574,598]" pageId="3" pageNumber="4" refString="Manning, R. B. &amp; Felder, D. L. (1991) Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington, 104 (4), 764 - 792." type="journal article" year="1991">Manning &amp; Felder (1991)</bibRefCitation>
reconsidered the diagnosis of
<taxonomicName id="9F934D65FFB6FFD4FB0D81111C69691D" box="[1220,1385,574,598]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FB0D81111C69691D" box="[1220,1385,574,598]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
</taxonomicName>
and added also the characters on the chelipeds, which are common in the fossil record and therefore usable for palaeontologists too. They argued for the taxonomic importance of the shape of the merus (e.g. presence/absence of meral hook), which is significant for many callianassid genera.
<bibRefCitation id="3C024B17FFB6FFD4FCFB81851D9D6989" author="Schweitzer" box="[818,1181,682,706]" pageId="3" pageNumber="4" refString="Schweitzer, C. E. &amp; Feldmann, R. M. (2002) New Eocene decapods (Thalassinidea and Brachyura) from Southern California. Journal of Crustacean Biology, 22 (4), 938 - 967." type="journal article" year="2002">Schweitzer &amp; Feldmann (2002)</bibRefCitation>
and
<bibRefCitation id="3C024B17FFB6FFD4FB13818519EF69AD" pageId="3" pageNumber="4" refString="Schweitzer, C. E., Gonzalez-Barba, G., Feldmann, R. M. &amp; Waugh, D. A. (2006 a) Decapoda (Thalassinidea and Paguroidea) from the Eocene Bateque and Tepetate Formations, Baja California Sur, Mexico: systematics, cuticle microstructure, and paleoecology. Annals of Carnegie Museum, 74 (4), 275 - 293." type="journal article">
Schweitzer
<emphasis id="6AE7EAF4FFB6FFD4FAA881831C956988" box="[1377,1429,683,707]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
. (2006
<emphasis id="6AE7EAF4FFB6FFD4FF1181FF19E669AC" box="[216,230,720,743]" italics="true" pageId="3" pageNumber="4">a</emphasis>
)
</bibRefCitation>
discussed usage of the characters on the major cheliped in the fossil record for assignment to the genus
<taxonomicName id="9F934D65FFB6FFD4FF5E81DC183C6840" box="[151,316,755,779]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FF5E81DC183C6840" box="[151,316,755,779]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
</taxonomicName>
. They noted that the carpus is variable in shape; however, it is not rectangular and typically has a rounded lower margin. The propodus has often serrated distal margin. The fixed finger tends to be edentulous, whereas the dactylus is stout and heavily armed with a triangular or rectangular tooth near the proximal end of the element.
</paragraph>
<paragraph id="582C36E6FFB6FFD4FF0080AD1A6C6F4D" blockId="3.[151,1437,358,1570]" pageId="3" pageNumber="4">
The merus of members of
<taxonomicName id="9F934D65FFB6FFD4FE3480AC1BA268D0" box="[509,674,899,923]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FE3480AC1BA268D0" box="[509,674,899,923]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
</taxonomicName>
is variable in shape but is always serrate along the lower margin. According to
<bibRefCitation id="3C024B17FFB6FFD4FEE680891B5268F5" author="Manning" box="[303,594,934,958]" pageId="3" pageNumber="4" refString="Manning, R. B. &amp; Felder, D. L. (1991) Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington, 104 (4), 764 - 792." type="journal article" year="1991">Manning &amp; Felder (1991)</bibRefCitation>
, it lacks a meral spine or hook; according to
<bibRefCitation id="3C024B17FFB6FFD4FB8A80891DCB68F5" author="Sakai" box="[1091,1227,934,958]" pageId="3" pageNumber="4" refString="Sakai, K. (1999) Synopsis of the family Callianassidae, with keys to subfamilies, genera and species, and the description of new taxa (Crustacea: Decapoda: Thalassinidea). Zoologische Verhandelingen (Leiden), 326, 1 - 152." type="journal article" year="1999">Sakai (1999</bibRefCitation>
,
<bibRefCitation id="3C024B17FFB6FFD4FB1F80891C1068F5" author="Sakai" box="[1238,1296,934,958]" pageId="3" pageNumber="4" refString="Sakai, K. (2005) Callianassoidea of the world (Decapoda: Thalassinidea). Crustaceana Monographs, 4, 1 - 285." type="journal article" year="2005">2005</bibRefCitation>
) it is with or without meral hook (mainly due to the synonymisation with the genus
<taxonomicName id="9F934D65FFB6FFD4FC7480E41C4968A9" authority="Manning &amp; Lemaitre, 1994" authorityName="Manning &amp; Lemaitre" authorityYear="1994" box="[957,1353,970,994]" class="Malacostraca" family="Callianassidae" genus="Sergio" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FC7480E41D0568A9" box="[957,1029,971,994]" italics="true" pageId="3" pageNumber="4">Sergio</emphasis>
<bibRefCitation id="3C024B17FFB6FFD4FBC480E51C4968A9" author="Manning" box="[1037,1353,970,994]" pageId="3" pageNumber="4" refString="Manning, R. B. &amp; Lemaitre, R. (1994) Sergio, a new genus of ghost shrimp from the Americas (Crustacea: Decapoda: Callianassidae). Nauplius, 1, 39 - 43." type="journal article" year="1994">Manning &amp; Lemaitre, 1994</bibRefCitation>
</taxonomicName>
).
<bibRefCitation id="3C024B17FFB6FFD4FA9780E519E06F4D" author="Sakai" pageId="3" pageNumber="4" refString="Sakai, K. (2011) Axioidea of the World and a Reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 520." type="journal article" year="2011">Sakai (2011)</bibRefCitation>
stated that the merus is with or without ventral convexity.
</paragraph>
<paragraph id="582C36E6FFB6FFD4FF00873D1C436E6D" blockId="3.[151,1437,358,1570]" pageId="3" pageNumber="4">
There are several genera different from
<taxonomicName id="9F934D65FFB6FFD4FD6F873C1A4B6F60" box="[678,843,1043,1067]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FD6F873C1A4B6F60" box="[678,843,1043,1067]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
</taxonomicName>
, namely
<taxonomicName id="9F934D65FFB6FFD4FC75873C1D046F61" box="[956,1028,1043,1066]" class="Malacostraca" family="Callianassidae" genus="Sergio" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FC75873C1D046F61" box="[956,1028,1043,1066]" italics="true" pageId="3" pageNumber="4">Sergio</emphasis>
</taxonomicName>
,
<taxonomicName id="9F934D65FFB6FFD4FBD1873C1C636F61" authority="Sakai, 1999" authorityName="Sakai" authorityYear="1999" box="[1048,1379,1042,1067]" class="Malacostraca" family="Callianassidae" genus="Podocallichirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FBD1873C1DCB6F60" box="[1048,1227,1043,1067]" italics="true" pageId="3" pageNumber="4">Podocallichirus</emphasis>
<bibRefCitation id="3C024B17FFB6FFD4FB1E873D1C636F61" author="Sakai" box="[1239,1379,1042,1066]" pageId="3" pageNumber="4" refString="Sakai, K. (1999) Synopsis of the family Callianassidae, with keys to subfamilies, genera and species, and the description of new taxa (Crustacea: Decapoda: Thalassinidea). Zoologische Verhandelingen (Leiden), 326, 1 - 152." type="journal article" year="1999">Sakai, 1999</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="9F934D65FFB6FFD4FF5E871818BA6F05" authority="Poore, 2000" authorityName="Poore" authorityYear="2000" box="[151,442,1078,1102]" class="Malacostraca" family="Callianassidae" genus="Grynaminna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FF5E871818256F05" box="[151,293,1079,1102]" italics="true" pageId="3" pageNumber="4">Grynaminna</emphasis>
<bibRefCitation id="3C024B17FFB6FFD4FEE7871818BA6F05" author="Poore" box="[302,442,1078,1102]" pageId="3" pageNumber="4" refString="Poore, G. C. B. (2000) A new genus and species of callianassid ghost shrimp from Kyushu, Japan (Decapoda: Thalassinidea)." type="book" year="2000">Poore, 2000</bibRefCitation>
</taxonomicName>
, which in general share characters on the chelipeds mentioned above. It is typical that such a morphology, i.e. generalized cheliped morphology embracing all above mentioned genera, is in palaeontological literature connected virtually with
<taxonomicName id="9F934D65FFB6FFD4FDB287511A206FDD" box="[635,800,1150,1174]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FDB287511A206FDD" box="[635,800,1150,1174]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
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only. This can be documented by currently recognized fossil taxa assigned to respective genera. According to
<bibRefCitation id="3C024B17FFB6FFD4FD32878C1AF56FF1" box="[763,1013,1186,1211]" pageId="3" pageNumber="4" refString="De Grave, S., Pentcheff, N. D., Ahyong, S. T., Chan, T. - Y., Crandall, K. A., Dworschak, P. C., Felder, D. L., Feldmann, R. M., Fransen, C. H. J. M., Goulding, L. Y. D., Lemaitre, R., Low, M. E. Y., Martin, J. W., Ng, P. K. L., Schweitzer, C. E., Tan, S. H., Tshudy, D. &amp; Wetzer, R. (2009) A classification of living and fossil genera of decapod crustaceans. The Raffles Bulletin of Zoology Supplements, 21, 1 - 109." type="journal article">
<collectingCountry id="20847676FFB6FFD4FD32878C1A1C6FF1" box="[763,796,1187,1210]" name="South Georgia and the South Sandwich Islands" pageId="3" pageNumber="4">De</collectingCountry>
Grave
<emphasis id="6AE7EAF4FFB6FFD4FCA5878B1A9D6FF0" box="[876,925,1187,1211]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
. (2009)
</bibRefCitation>
and
<bibRefCitation id="3C024B17FFB6FFD4FBE3878D1C356FF1" box="[1066,1333,1186,1211]" pageId="3" pageNumber="4" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, 1 - 222." type="journal article">
Schweitzer
<emphasis id="6AE7EAF4FFB6FFD4FB65878B1DDD6FF0" box="[1196,1245,1187,1211]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
. (2010)
</bibRefCitation>
there are 18 fossil species of
<taxonomicName id="9F934D65FFB6FFD4FEB687E91B246F95" box="[383,548,1222,1246]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FEB687E91B246F95" box="[383,548,1222,1246]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
</taxonomicName>
, but only one fossil species of
<taxonomicName id="9F934D65FFB6FFD4FC5D87E91D476F95" box="[916,1095,1222,1246]" class="Malacostraca" family="Callianassidae" genus="Podocallichirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FC5D87E91D476F95" box="[916,1095,1222,1246]" italics="true" pageId="3" pageNumber="4">Podocallichirus</emphasis>
</taxonomicName>
(known from the Pleistocene strata of
<collectingCountry id="20847676FFB6FFD4FECC87C418486E49" box="[261,328,1259,1282]" name="Japan" pageId="3" pageNumber="4">Japan</collectingCountry>
; see
<bibRefCitation id="3C024B17FFB6FFD4FE4387C41B8B6E49" author="Karasawa" box="[394,651,1258,1283]" pageId="3" pageNumber="4" refString="Karasawa, H., Tanaka, T., Kobayashi, N., Goda, T., Ohira, N. &amp; Shinya, J. (2006) Podocallichirus grandis (Crustacea: Decapoda: Thalassinidea) preserved within burrows from the middle Pleistocene Atsumi Group of Aichi Prefecture, Japan. Bulletin of the Mizunami Fossil Museum, 33, 127 - 133. [In Japanese]" type="journal article" year="2006">
Karasawa
<emphasis id="6AE7EAF4FFB6FFD4FDCC87C31B3C6E48" box="[517,572,1259,1283]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
. 2006
</bibRefCitation>
) and no fossil
<taxonomicName id="9F934D65FFB6FFD4FC8E87C41AD56E49" box="[839,981,1259,1282]" class="Malacostraca" family="Callianassidae" genus="Grynaminna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FC8E87C41AD56E49" box="[839,981,1259,1282]" italics="true" pageId="3" pageNumber="4">Grynaminna</emphasis>
</taxonomicName>
(see Hyžný &amp; Karasawa in press for a different view). One extant species of
<taxonomicName id="9F934D65FFB6FFD4FD9F86201B9E6E6D" box="[598,670,1295,1318]" class="Malacostraca" family="Callianassidae" genus="Sergio" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FD9F86201B9E6E6D" box="[598,670,1295,1318]" italics="true" pageId="3" pageNumber="4">Sergio</emphasis>
</taxonomicName>
is also known from the fossil state (
<bibRefCitation id="3C024B17FFB6FFD4FBFB86211C376E6D" author="Portell" box="[1074,1335,1294,1319]" pageId="3" pageNumber="4" refString="Portell, R. W. &amp; Agnew, J. G. (2004) Pliocene and Pleistocene decapod crustaceans. Florida Fossil Invertebrates, 4, 1 - 29." type="journal article" year="2004">Portell &amp; Agnew 2004</bibRefCitation>
).
</paragraph>
<paragraph id="582C36E6FFB6FFD4FF00861D189A6D69" blockId="3.[151,1437,358,1570]" pageId="3" pageNumber="4">
Much confusion has been created by
<bibRefCitation id="3C024B17FFB6FFD4FDAF861D1BEF6E01" author="Sakai" box="[614,751,1330,1354]" pageId="3" pageNumber="4" refString="Sakai, K. (1999) Synopsis of the family Callianassidae, with keys to subfamilies, genera and species, and the description of new taxa (Crustacea: Decapoda: Thalassinidea). Zoologische Verhandelingen (Leiden), 326, 1 - 152." type="journal article" year="1999">Sakai (1999</bibRefCitation>
,
<bibRefCitation id="3C024B17FFB6FFD4FD33861D1A336E01" author="Sakai" box="[762,819,1330,1354]" pageId="3" pageNumber="4" refString="Sakai, K. (2005) Callianassoidea of the world (Decapoda: Thalassinidea). Crustaceana Monographs, 4, 1 - 285." type="journal article" year="2005">2005</bibRefCitation>
) when he synonymized
<taxonomicName id="9F934D65FFB6FFD4FB88861C1D896E01" box="[1089,1161,1331,1354]" class="Malacostraca" family="Callianassidae" genus="Sergio" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FB88861C1D896E01" box="[1089,1161,1331,1354]" italics="true" pageId="3" pageNumber="4">Sergio</emphasis>
</taxonomicName>
with
<taxonomicName id="9F934D65FFB6FFD4FB01861C1C6D6E00" box="[1224,1389,1331,1355]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FB01861C1C6D6E00" box="[1224,1389,1331,1355]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
</taxonomicName>
and
<taxonomicName id="9F934D65FFB6FFD4FF5E867818256E25" box="[151,293,1367,1390]" class="Malacostraca" family="Callianassidae" genus="Grynaminna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FF5E867818256E25" box="[151,293,1367,1390]" italics="true" pageId="3" pageNumber="4">Grynaminna</emphasis>
</taxonomicName>
with
<taxonomicName id="9F934D65FFB6FFD4FEA386791B1D6E25" box="[362,541,1366,1390]" class="Malacostraca" family="Callianassidae" genus="Podocallichirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FEA386791B1D6E25" box="[362,541,1366,1390]" italics="true" pageId="3" pageNumber="4">Podocallichirus</emphasis>
</taxonomicName>
(see also
<bibRefCitation id="3C024B17FFB6FFD4FD5C86791A146E25" author="Sakai" box="[661,788,1366,1390]" pageId="3" pageNumber="4" refString="Sakai, K. (2011) Axioidea of the World and a Reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 520." type="journal article" year="2011">Sakai 2011</bibRefCitation>
for
<bibRefCitation id="3C024B17FFB6FFD4FC8086781A556E25" author="Sakai" box="[841,853,1367,1390]" pageId="3" pageNumber="4" refString="Sakai, K. (2011) Axioidea of the World and a Reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 520." type="journal article" year="2011" yearSuffix="a">a</bibRefCitation>
different view on the taxonomy of these genera). Thus, we can talk about several different taxonomic concepts of
<taxonomicName id="9F934D65FFB6FFD4FCB486541D226ED8" box="[893,1058,1403,1427]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FCB486541D226ED8" box="[893,1058,1403,1427]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
</taxonomicName>
presented by
<bibRefCitation id="3C024B17FFB6FFD4FB0C86551C4F6ED9" author="Sakai" box="[1221,1359,1402,1426]" pageId="3" pageNumber="4" refString="Sakai, K. (1988) A new genus and five new species of Callianassidae (Crustacea: Decapoda: Thalassinidea) from Northern Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 5 (1), 51 - 69." type="journal article" year="1988">Sakai (1988</bibRefCitation>
,
<bibRefCitation id="3C024B17FFB6FFD4FA9486551C986ED9" author="Sakai" box="[1373,1432,1402,1426]" pageId="3" pageNumber="4" refString="Sakai, K. (1999) Synopsis of the family Callianassidae, with keys to subfamilies, genera and species, and the description of new taxa (Crustacea: Decapoda: Thalassinidea). Zoologische Verhandelingen (Leiden), 326, 1 - 152." type="journal article" year="1999">1999</bibRefCitation>
,
<bibRefCitation id="3C024B17FFB6FFD4FF5E86B119D26EFD" author="Sakai" box="[151,210,1438,1462]" pageId="3" pageNumber="4" refString="Sakai, K. (2005) Callianassoidea of the world (Decapoda: Thalassinidea). Crustaceana Monographs, 4, 1 - 285." type="journal article" year="2005">2005</bibRefCitation>
,
<bibRefCitation id="3C024B17FFB6FFD4FF1786B118176EFD" author="Sakai" box="[222,279,1438,1462]" pageId="3" pageNumber="4" refString="Sakai, K. (2011) Axioidea of the World and a Reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 520." type="journal article" year="2011">2011</bibRefCitation>
) and
<bibRefCitation id="3C024B17FFB6FFD4FE9E86B11B816EFD" author="Manning" box="[343,641,1438,1462]" pageId="3" pageNumber="4" refString="Manning, R. B. &amp; Felder, D. L. (1991) Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington, 104 (4), 764 - 792." type="journal article" year="1991">Manning &amp; Felder (1991)</bibRefCitation>
. Identifying the fossil material as a member of any of the above mentioned genera strongly depends on taxonomy adopted. Therefore it is possible that the genus
<taxonomicName id="9F934D65FFB6FFD4FB6B86EC1C476E90" box="[1186,1351,1475,1499]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FB6B86EC1C476E90" box="[1186,1351,1475,1499]" italics="true" pageId="3" pageNumber="4">Neocallichirus</emphasis>
</taxonomicName>
as usually recognized in the fossil record is a mixture of closely allied genera. The issue was recently discussed by Hyžný &amp; Karasawa (in press).
</paragraph>
<paragraph id="582C36E6FFB6FFD4FF5E8559180B6DFA" blockId="3.[151,884,1654,1713]" pageId="3" pageNumber="4">
<emphasis id="6AE7EAF4FFB6FFD4FF5E855918D76DDB" bold="true" box="[151,471,1654,1680]" pageId="3" pageNumber="4">
<taxonomicName id="9F934D65FFB6FFD4FF5E855918B36DDB" box="[151,435,1654,1680]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFB6FFD4FF5E855918B36DDB" bold="true" box="[151,435,1654,1680]" italics="true" pageId="3" pageNumber="4">Neocallichirus brocchii</emphasis>
</taxonomicName>
(L
</emphasis>
ő
<emphasis id="6AE7EAF4FFB6FFD4FE2F85591A746DDB" bold="true" box="[486,884,1654,1680]" pageId="3" pageNumber="4">renthey, 1897) new combination</emphasis>
(
<figureCitation id="C0A82A63FFB6FFD4FF5685B618036DFA" box="[159,259,1689,1713]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="4.[151,250,1113,1136]" captionStartId-1="5.[151,250,1674,1697]" captionStartId-2="6.[151,250,1006,1029]" captionStartId-3="7.[151,250,1431,1454]" captionStartId-4="8.[151,250,1740,1763]" captionTargetBox-0="[159,1428,205,1070]" captionTargetBox-1="[279,1308,201,1641]" captionTargetBox-2="[160,1426,202,978]" captionTargetBox-3="[170,1427,388,1389]" captionTargetBox-4="[159,1424,199,1700]" captionTargetId-0="figure@4.[151,1436,195,1091]" captionTargetId-1="figure@5.[272,1314,193,1645]" captionTargetId-2="figure@6.[151,1436,193,985]" captionTargetId-3="figure@7.[151,1436,373,1410]" captionTargetId-4="figure@8.[151,1436,193,1719]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." captionText-1="FIGURE 3. Neocallichirus brocchii (Lőrenthey, 1897) new combination, major chelae (propodi and dactyli): A, left chela (robust morphotype), KGP-MH DH 001; B, right chela (slender morphotype), PCMH- 001; C D, right chela from inner and outer view (robust morphotype), PCRB-DH 008; E, left dactylus, PCRB-DH 001; F, left dactylus, PCRB-DH 002; G, right dactylus, KGP-MH DH 012; H, right dactylus, PCRB-DH 004; I, right dactylus, PCRB-DH 003; J, left chela (robust morphotype), KGP-MH DH 064, note serration on the lower margin (K); L M, fragmentary right chela (robust morphotype), PCRB-DH 005; N, left chela (robust morphotype), SNM Z- 37534, note serration on the lower margin (O); P, right chela (slender morphotype), PCRB-DH 010, note short carpus; Q, fragmented right chela (slender morphotype), PCMH- 003, note occlusal keel on the fixed finger positioned medially; R, right chela (slender morphotype), PCRB-DH 007, note occlusal keel on the fixed finger positioned medially; S, fragmented right chela (robust morphotype), PCMH- 002, note occlusal keel on the fixed finger positioned laterally; T, right chela, KGP-MH DH- 66, note the unusual length / height ratio. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." captionText-2="FIGURE 4. Neocallichirus brocchii (Lőrenthey, 1897) new combination, major chelae (carpi and meri): A, left carpus, KGP- MH DH 065; B, right carpus, KGP-MH DH 060; C, left carpus, PCRB-DH 009, note serration on the lower margin (D); E, left articulated merus and carpus, SNM Z- 37549; F, left merus, PCRB-DH 014; G, right merus, KGP-MH DH 052; H, left merus, KGP-MH DH 010; I, right merus, PCRB-DH 016; J, left merus, PCRB-DH 015; K, right merus, SNM Z- 37542. All specimens belong to robust morphotype, except „ I “ being a slender morphotype. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." captionText-3="FIGURE 5. Neocallichirus brocchii (Lőrenthey, 1897) new combination, minor chelae (propodi and dactyli): A B, left chela from outer and inner view, KGP-MH DH 067; C, right chela, PCRB-DH 013; D, fragmented right chela, PCRB-DH 011; E, left dactylus, PCRB-DH 012. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." captionText-4="FIGURE 6. Neocallichirus brocchii (Lőrenthey, 1897) new combination, in situ preservation of the specimen KGP-MH DH- 0 58 (Dúbravská hlavica locality): A, view of the individual (slender morphotype) preserved in a presumed burrow structure; B, minor cheliped; C, major cheliped; D, view from below where both chelipeds can be recognized. Note also the partial preservation of thoracopods P 2 P 4. All specimens are to scale and were covered with ammonium chloride prior to photography." httpUri-0="https://zenodo.org/record/209486/files/figure.png" httpUri-1="https://zenodo.org/record/209487/files/figure.png" httpUri-2="https://zenodo.org/record/209488/files/figure.png" httpUri-3="https://zenodo.org/record/209489/files/figure.png" httpUri-4="https://zenodo.org/record/209490/files/figure.png" pageId="3" pageNumber="4">Figs 26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1089656DFFB6FFDEFF5E85CE1B166A74" lastPageId="9" lastPageNumber="10" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="582C36E6FFB6FFD4FF5E85CE1C026C5E" blockId="3.[151,1436,1761,1967]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="788311C8FFB6FFD4FF5E85CE1C026C5E" pageId="3" pageNumber="4">
<taxonomicName id="9F934D65FFB6FFD4FF5E85CE1B486DBC" authority="Brocchii Lorenthey, 1897: 161" authorityName="Brocchii Lorenthey" authorityPageNumber="161" authorityYear="1897" box="[151,584,1761,1783]" class="Malacostraca" family="Callianassidae" genus="Calianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FF5E85CE18656DBC" box="[151,357,1761,1783]" italics="true" pageId="3" pageNumber="4">Calianassa Brocchii</emphasis>
<treatmentCitation id="D93210F7FFB6FFD4FEA585CE1B486DBC" author="Lorenthey" box="[364,584,1761,1783]" page="161" pageId="3" pageNumber="4" year="1897">
<bibRefCitation id="3C024B17FFB6FFD4FEA585CE1B166DBC" author="Lorenthey" box="[364,534,1761,1783]" pageId="3" pageNumber="4" refString="Lorenthey, E. (1897) Adatok Magyarorszag harmadkoru rak faunajahoz. Mathematikai es Termeszettudomanyi Ertesito, 15, 149 - 169." type="journal article" year="1897">Lőrenthey, 1897</bibRefCitation>
: 161
</treatmentCitation>
</taxonomicName>
, 168, 169;
<treatmentCitation id="D93210F7FFB6FFD4FD7085CE1AA06DBC" author="Lorenthey" box="[697,928,1761,1783]" page="106" pageId="3" pageNumber="4" year="1898">
<bibRefCitation id="3C024B17FFB6FFD4FD7085CE1A6E6DBC" author="Lorenthey" box="[697,878,1761,1783]" pageId="3" pageNumber="4" refString="Lorenthey, E. (1898 a) Beitrage zur Decapodenfauna des ungarischen Tertiars. Mathematischen und Naturwissenschaftlichen Berichte aus Ungarn, 14, 92 - 115." type="journal article" year="1898" yearSuffix="a">Lőrenthey, 1898a</bibRefCitation>
: 106
</treatmentCitation>
, 114, 115;
<treatmentCitation id="D93210F7FFB6FFD4FBD985CE1DF76DBC" author="Lorenthey" box="[1040,1271,1761,1783]" page="132" pageId="3" pageNumber="4" year="1898">
<bibRefCitation id="3C024B17FFB6FFD4FBD985CE1DC66DBC" author="Lorenthey" box="[1040,1222,1761,1783]" pageId="3" pageNumber="4" refString="Lorenthey, E. (1898 b) Adatok Magyarorszag harmadkoru rak-faunajahoz. Mathematikai es Termeszettudomanyi Kozllemenyek, 27 (2), 103 - 271." type="journal article" year="1898" yearSuffix="b">Lőrenthey, 1898b</bibRefCitation>
: 132
</treatmentCitation>
134, 155, pl. 9, fig. 5;
<treatmentCitation id="D93210F7FFB6FFD4FEC3842F18F06C5D" author="Lorenthey" box="[266,496,1792,1814]" page="104" pageId="3" pageNumber="4" year="1898">
<bibRefCitation id="3C024B17FFB6FFD4FEC3842F18C06C5D" author="Lorenthey" box="[266,448,1792,1814]" pageId="3" pageNumber="4" refString="Lorenthey, E. (1898 b) Adatok Magyarorszag harmadkoru rak-faunajahoz. Mathematikai es Termeszettudomanyi Kozllemenyek, 27 (2), 103 - 271." type="journal article" year="1898" yearSuffix="b">Lőrenthey, 1898b</bibRefCitation>
: 104
</treatmentCitation>
105, pl. 9, fig. 5; Lőrenthey in
<treatmentCitation id="D93210F7FFB6FFD4FCFA842F1D746C5D" author="Lorenthey" box="[819,1140,1792,1814]" page="64" pageId="3" pageNumber="4" year="1929">
<bibRefCitation id="3C024B17FFB6FFD4FCFA842F1D4F6C5E" author="Lorenthey" box="[819,1103,1792,1814]" pageId="3" pageNumber="4" refString="Lorenthey, E. &amp; Beurlen, K. (1929) Die fossilen Dekapoden der Lander der Ungarischen Krone. Geologica Hungarica, Series Palaeontologica, 3, 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen, 1929</bibRefCitation>
: 64
</treatmentCitation>
, pl. 2, fig. 13;
</treatmentCitationGroup>
</paragraph>
<paragraph id="582C36E6FFB6FFD4FF5E84301B686C7F" blockId="3.[151,1436,1761,1967]" box="[151,616,1822,1845]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="788311C8FFB6FFD4FF5E84301B686C7F" box="[151,616,1822,1845]" pageId="3" pageNumber="4">
<taxonomicName id="9F934D65FFB6FFD4FF5E843018736C7E" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" authority="Brocchii." authorityName="Brocchii." box="[151,371,1823,1845]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FF5E843018736C7E" box="[151,371,1823,1845]" italics="true" pageId="3" pageNumber="4">Callianassa Brocchii.</emphasis>
</taxonomicName>
<treatmentCitation id="D93210F7FFB6FFD4FE5084311B646C7F" author="Glaessner" box="[409,612,1822,1844]" page="77" pageId="3" pageNumber="4" year="1929">
<bibRefCitation id="3C024B17FFB6FFD4FE5084311B3E6C7F" author="Glaessner" box="[409,574,1822,1844]" pageId="3" pageNumber="4" refString="Glaessner, M. F. (1929) Crustacea Decapoda. In. Pompeckj, F. J. (Ed.), Fossilium catalogus, 1: Animalium, 41, 1 - 464. W. Junk, Berlin." type="journal article" year="1929">Glaessner, 1929</bibRefCitation>
: 77
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="582C36E6FFB6FFD4FF5E84111A8B6C18" blockId="3.[151,1436,1761,1967]" box="[151,907,1853,1875]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="788311C8FFB6FFD4FF5E84111A8B6C18" box="[151,907,1853,1875]" pageId="3" pageNumber="4">
?
<taxonomicName id="9F934D65FFB6FFD4FF6B841218146C18" box="[162,276,1853,1875]" class="Malacostraca" family="Callianassidae" genus="Calianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FF6B841218146C18" box="[162,276,1853,1875]" italics="true" pageId="3" pageNumber="4">Calianassa</emphasis>
</taxonomicName>
<taxonomicName id="9F934D65FFB6FFD4FED3841218956C18" box="[282,405,1853,1875]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
cf.
<emphasis id="6AE7EAF4FFB6FFD4FEF2841218956C18" box="[315,405,1853,1875]" italics="true" pageId="3" pageNumber="4">Brocchii</emphasis>
</taxonomicName>
. — Lőrenthey in
<treatmentCitation id="D93210F7FFB6FFD4FD8C84121A876C18" author="Lorenthey" box="[581,903,1853,1875]" page="69" pageId="3" pageNumber="4" year="1929">
<bibRefCitation id="3C024B17FFB6FFD4FD8C84121A616C18" author="Lorenthey" box="[581,865,1853,1875]" pageId="3" pageNumber="4" refString="Lorenthey, E. &amp; Beurlen, K. (1929) Die fossilen Dekapoden der Lander der Ungarischen Krone. Geologica Hungarica, Series Palaeontologica, 3, 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen, 1929</bibRefCitation>
: 69
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="582C36E6FFB6FFD4FF5E84731A6D6C3A" blockId="3.[151,1436,1761,1967]" box="[151,877,1883,1906]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="788311C8FFB6FFD4FF5E84731A6D6C3A" box="[151,877,1883,1906]" pageId="3" pageNumber="4">
<emphasis id="6AE7EAF4FFB6FFD4FF5E8473188D6C39" box="[151,397,1884,1906]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="9F934D65FFB6FFD4FF5E847318716C39" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" authority="Brocchii." authorityName="Brocchii." box="[151,369,1884,1906]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Callianassa brocchii.</taxonomicName>
</emphasis>
<treatmentCitation id="D93210F7FFB6FFD4FE5A84741B4C6C3A" author="Muller" box="[403,588,1883,1905]" page="274" pageId="3" pageNumber="4" year="1979">
<bibRefCitation id="3C024B17FFB6FFD4FE5A84741B196C3A" author="Muller" box="[403,537,1883,1905]" pageId="3" pageNumber="4" refString="Muller, P. (1979) Decapoda (Crustacea) fauna a budapesti miocenbol (5). Foldtani Kozlony, Bulletin of the Hungarian Geological Society, 108, 272 - 318." type="journal article" year="1979">Müller, 1979</bibRefCitation>
: 274
</treatmentCitation>
;
<treatmentCitation id="D93210F7FFB6FFD4FD9F84741A696C3A" author="Schweitzer" box="[598,873,1883,1906]" page="34" pageId="3" pageNumber="4" year="2010">
<bibRefCitation id="3C024B17FFB6FFD4FD9F84741A436C3A" author="Schweitzer" box="[598,835,1883,1906]" pageId="3" pageNumber="4" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, 1 - 222." type="journal article" year="2010">
Schweitzer
<emphasis id="6AE7EAF4FFB6FFD4FD0484721BFA6C39" box="[717,762,1884,1906]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
., 2010
</bibRefCitation>
: 34
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="582C36E6FFB6FFD4FF5E84541BCA6CDB" blockId="3.[151,1436,1761,1967]" box="[151,714,1914,1936]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="788311C8FFB6FFD4FF5E84541BCA6CDB" box="[151,714,1914,1936]" pageId="3" pageNumber="4">
<taxonomicName id="9F934D65FFB6FFD4FF56845518186CDB" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" box="[159,280,1914,1936]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FF56845518186CDB" box="[159,280,1914,1936]" italics="true" pageId="3" pageNumber="4">Callianassa</emphasis>
</taxonomicName>
<emphasis id="6AE7EAF4FFB6FFD4FEEE8455189D6CDB" box="[295,413,1914,1936]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="9F934D65FFB6FFD4FEEE8455187E6CDB" box="[295,382,1914,1936]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="brocchii">brocchii</taxonomicName>
. —
</emphasis>
<treatmentCitation id="D93210F7FFB6FFD4FE6384551B566CDB" author="Muller" box="[426,598,1914,1936]" page="51" pageId="3" pageNumber="4" year="1984">
<bibRefCitation id="3C024B17FFB6FFD4FE6384551B326CDB" author="Muller" box="[426,562,1914,1936]" pageId="3" pageNumber="4" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller, 1984</bibRefCitation>
: 51
</treatmentCitation>
, pl. 3 fig.3.
</treatmentCitationGroup>
</paragraph>
<paragraph id="582C36E6FFB6FFD4FF5E84B51D156CE4" blockId="3.[151,1436,1761,1967]" box="[151,1045,1945,1967]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="788311C8FFB6FFD4FF5E84B51D156CE4" box="[151,1045,1945,1967]" pageId="3" pageNumber="4">
&quot;
<taxonomicName id="9F934D65FFB6FFD4FF6884B6181A6CE4" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" box="[161,282,1945,1967]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB6FFD4FF6884B6181A6CE4" box="[161,282,1945,1967]" italics="true" pageId="3" pageNumber="4">Callianassa</emphasis>
</taxonomicName>
&quot;
<emphasis id="6AE7EAF4FFB6FFD4FEE284B618776CE4" box="[299,375,1945,1967]" italics="true" pageId="3" pageNumber="4">brocchi</emphasis>
(sic). —
<treatmentCitation id="D93210F7FFB6FFD4FE1E84B61B8C6CE4" author="Hyzny" box="[471,652,1945,1967]" page="41" pageId="3" pageNumber="4" year="2011">
<bibRefCitation id="3C024B17FFB6FFD4FE1E84B61B686CE4" author="Hyzny" box="[471,616,1945,1967]" pageId="3" pageNumber="4" refString="Hyzny, M. (2011 a) In situ mud shrimps (Decapoda: Axiidea: Callianassidae) preserved within their burrows from the middle Miocene of the Central Paratethys. Bulletin of the Mizunami Fossil Museum, 37, 37 - 46." type="journal article" year="2011" yearSuffix="a">Hyžný, 2011a</bibRefCitation>
: 41
</treatmentCitation>
,
<tableCitation id="1511035DFFB6FFD4FD5E84B61BE66CE4" box="[663,742,1945,1967]" captionStart="TABLE 2" captionStartId="18.[151,235,152,173]" captionTargetBox="[151,1436,304,1979]" captionTargetPageId="18" captionText="TABLE 2. Measurements of selected well preserved specimens of Eucalliax pseudorakosensis (Lőrenthey in Lőrenthey &amp; Beurlen, 1929) comb. nov. Handedness: L = left; R = right; measurements: L = max. length; H = max. height; ~ = approximate value;&gt; = &quot; more than &quot; (in case the approximation is not possible due to the state of preservation); values are in mm." httpUri="http://table.plazi.org/id/0CEC666EFFA7FFC5FF5E83B71C326BA0" pageId="3" pageNumber="4" tableUuid="0CEC666EFFA7FFC5FF5E83B71C326BA0">Table 2</tableCitation>
.;
<treatmentCitation id="D93210F7FFB6FFD4FD3F84B61AB96CE4" author="Hyzny" box="[758,953,1945,1967]" page="167" pageId="3" pageNumber="4" year="2011">
<bibRefCitation id="3C024B17FFB6FFD4FD3F84B61A886CE4" author="Hyzny" box="[758,904,1945,1967]" pageId="3" pageNumber="4" refString="Hyzny M. (2011 b) Synopsis of fossil decapod crustaceans from Slovakia (Western Carpathians). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 260 (2), 165 - 171." type="journal article" year="2011" yearSuffix="b">Hyžný, 2011b</bibRefCitation>
: 167
</treatmentCitation>
,
<tableCitation id="1511035DFFB6FFD4FC0D84B61D106CE4" box="[964,1040,1945,1967]" captionStart="TABLE 1" captionStartId="9.[151,235,506,527]" captionTargetBox="[151,1436,657,1332]" captionTargetPageId="9" captionText="TABLE 1. Measurements of selected well preserved specimens of Neocallichirus brocchii (Lőrenthey, 1897) comb. nov. Handedness: L = left; R = right; measurements: L = max. length; H = max. height; ~ = approximate value;&gt; = &quot; more than &quot; (in case the approximation is not possible due to the state of preservation); values are in mm." httpUri="http://table.plazi.org/id/0CEC666EFFBCFFDEFF5E82D51AEF6905" pageId="3" pageNumber="4" tableUuid="0CEC666EFFBCFFDEFF5E82D51AEF6905">Table 1</tableCitation>
.
</treatmentCitationGroup>
</paragraph>
<caption id="0CEC666EFFB1FFD3FF5E87761C346FA2" httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="4" pageNumber="5" targetBox="[159,1428,205,1070]" targetPageId="4">
<paragraph id="582C36E6FFB1FFD3FF5E87761C346FA2" blockId="4.[151,1436,1113,1258]" pageId="4" pageNumber="5">
<emphasis id="6AE7EAF4FFB1FFD3FF5E877618126F25" bold="true" box="[151,274,1113,1136]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
<taxonomicName id="9F934D65FFB1FFD3FED187761BC26F3B" authority="Lorenthey, 1897" authorityName="Lorenthey" authorityYear="1897" box="[280,706,1113,1136]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="brocchii" status="comb. nov.">
<emphasis id="6AE7EAF4FFB1FFD3FED187761B076F24" box="[280,519,1113,1135]" italics="true" pageId="4" pageNumber="5">Neocallichirus brocchii</emphasis>
(Lőrenthey, 1897)
</taxonomicName>
<taxonomicNameLabel id="71D4578FFFB1FFD3FD0187751A796F24" box="[712,889,1113,1135]" pageId="4" pageNumber="5">new combination</taxonomicNameLabel>
, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); DE, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; GH, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures AF are in scale.
</paragraph>
</caption>
<paragraph id="582C36E6FFB1FFD3FF5E86381C406D61" blockId="4.[151,1437,1302,1974]" pageId="4" pageNumber="5">
<emphasis id="6AE7EAF4FFB1FFD3FF5E863818856E64" bold="true" box="[151,389,1302,1327]" pageId="4" pageNumber="5">Emended diagnosis.</emphasis>
Strongly heterochelous callichirine ghost shrimp with chelipeds without pronounced tuberculation. Merus of major cheliped ovoid with lower margin possesssing a toothed blade consisting of up to 12 teeth; carpus broad with rounded and serrated lower margin; propodus tapering distally with serrated lower margin, the palm subequal in length and height, distal margin rounded and serrated, occlusal margin of fixed finger armed with small teeth; dactylus long and slender with occlusal margin forming a well developed keel with long, quadrate, blunt tooth proximally and one indented, sharp tooth distally followed with several tiny teeth, tip forming sharp hook. Carpus of minor cheliped slightly higher than propodus; propodus slender, tapering distally, fixed finger about as long as manus; fingers without armature, occlusal margin of dactylus forming weak serrated keel.
</paragraph>
<paragraph id="582C36E6FFB1FFD3FF0E85181A7A6DF1" blockId="4.[151,1437,1302,1974]" pageId="4" pageNumber="5">
<emphasis id="6AE7EAF4FFB1FFD3FF0E851818CE6D04" bold="true" box="[199,462,1590,1615]" pageId="4" pageNumber="5">Emended description.</emphasis>
Major cheliped massive, located on either right or left side of body, smooth, without pronounced tuberculation on lateral surfaces; outer surface usually vaulted. Two different morphotypes were identified in the nature of major cheliped; robust morphotype is relatively larger than slender morphotype. Ischium is slender, longer than high, tapering proximally, not well preserved.
</paragraph>
<paragraph id="582C36E6FFB1FFD1FF0085E81D326EE4" blockId="4.[151,1437,1302,1974]" lastBlockId="6.[151,1436,1251,2032]" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<emphasis id="6AE7EAF4FFB1FFD3FF0085E81B2E6D95" box="[201,558,1734,1758]" italics="true" pageId="4" pageNumber="5">Major chela robust morphotype</emphasis>
: Merus ovoid in shape, longer than high (L/H = 1.51.7), highest proximally; lower margin with toothed blade (up to 12 teeth present); upper margin slightly convex (
<figureCitation id="C0A82A63FFB1FFD3FBBA85C51DB96C49" box="[1139,1209,1770,1794]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
G, 4FH, J, K). Carpus broad, massive, approximately as long as high (L/H = 0.81.1); upper margin straight; lower margin rounded forming one edge with proximal margin (
<figureCitation id="C0A82A63FFB1FFD3FDA1841D1BAF6C01" box="[616,687,1842,1866]" captionStart="FIGURE 4" captionStartId="6.[151,250,1006,1029]" captionTargetBox="[160,1426,202,978]" captionTargetId="figure@6.[151,1436,193,985]" captionTargetPageId="6" captionText="FIGURE 4. Neocallichirus brocchii (Lőrenthey, 1897) new combination, major chelae (carpi and meri): A, left carpus, KGP- MH DH 065; B, right carpus, KGP-MH DH 060; C, left carpus, PCRB-DH 009, note serration on the lower margin (D); E, left articulated merus and carpus, SNM Z- 37549; F, left merus, PCRB-DH 014; G, right merus, KGP-MH DH 052; H, left merus, KGP-MH DH 010; I, right merus, PCRB-DH 016; J, left merus, PCRB-DH 015; K, right merus, SNM Z- 37542. All specimens belong to robust morphotype, except „ I “ being a slender morphotype. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/209488/files/figure.png" pageId="4" pageNumber="5">Figs 4</figureCitation>
AE), serrated with tiny denticles (
<figureCitation id="C0A82A63FFB1FFD3FBF3841D1D7E6C01" box="[1082,1150,1842,1866]" captionStart="FIGURE 4" captionStartId="6.[151,250,1006,1029]" captionTargetBox="[160,1426,202,978]" captionTargetId="figure@6.[151,1436,193,985]" captionTargetPageId="6" captionText="FIGURE 4. Neocallichirus brocchii (Lőrenthey, 1897) new combination, major chelae (carpi and meri): A, left carpus, KGP- MH DH 065; B, right carpus, KGP-MH DH 060; C, left carpus, PCRB-DH 009, note serration on the lower margin (D); E, left articulated merus and carpus, SNM Z- 37549; F, left merus, PCRB-DH 014; G, right merus, KGP-MH DH 052; H, left merus, KGP-MH DH 010; I, right merus, PCRB-DH 016; J, left merus, PCRB-DH 015; K, right merus, SNM Z- 37542. All specimens belong to robust morphotype, except „ I “ being a slender morphotype. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/209488/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
D); upper and lower margin forming prominent keel curved inward. Propodus massive, longer than high, slightly longer than carpus, tapering distally (
<figureCitation id="C0A82A63FFB1FFD3FEEA8455186A6CD9" box="[291,362,1914,1938]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
A, 3); palm of the propodus subequal in length and height; upper margin slightly convex; lower margin straight, serrated (
<figureCitation id="C0A82A63FFB1FFD3FE7F84B118FC6CFD" box="[438,508,1950,1974]" captionStart="FIGURE 3" captionStartId="5.[151,250,1674,1697]" captionTargetBox="[279,1308,201,1641]" captionTargetId="figure@5.[272,1314,193,1645]" captionTargetPageId="5" captionText="FIGURE 3. Neocallichirus brocchii (Lőrenthey, 1897) new combination, major chelae (propodi and dactyli): A, left chela (robust morphotype), KGP-MH DH 001; B, right chela (slender morphotype), PCMH- 001; C D, right chela from inner and outer view (robust morphotype), PCRB-DH 008; E, left dactylus, PCRB-DH 001; F, left dactylus, PCRB-DH 002; G, right dactylus, KGP-MH DH 012; H, right dactylus, PCRB-DH 004; I, right dactylus, PCRB-DH 003; J, left chela (robust morphotype), KGP-MH DH 064, note serration on the lower margin (K); L M, fragmentary right chela (robust morphotype), PCRB-DH 005; N, left chela (robust morphotype), SNM Z- 37534, note serration on the lower margin (O); P, right chela (slender morphotype), PCRB-DH 010, note short carpus; Q, fragmented right chela (slender morphotype), PCMH- 003, note occlusal keel on the fixed finger positioned medially; R, right chela (slender morphotype), PCRB-DH 007, note occlusal keel on the fixed finger positioned medially; S, fragmented right chela (robust morphotype), PCMH- 002, note occlusal keel on the fixed finger positioned laterally; T, right chela, KGP-MH DH- 66, note the unusual length / height ratio. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/209487/files/figure.png" pageId="4" pageNumber="5">Figs 3</figureCitation>
A,
<collectingCountry id="20847676FFB1FFD3FDD684B11B366CFD" box="[543,566,1950,1974]" name="Iceland" pageId="4" pageNumber="5">O</collectingCountry>
); upper and lower margins keeled, keel missing on fixed finger; distal margin rounded and serrated with up to 10 small denticles on each side of articulation with dactylus, serrated incision present just below the articulation with dactylus (
<figureCitation id="C0A82A63FFB3FFD1FD5986281BD66E54" box="[656,726,1287,1311]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
A, 3D, J, M). Fixed finger slender, triangular in shape; occlusal margin armed with small teeth, forming serrated keel on the outer side of fixed finger from distal view (
<figureCitation id="C0A82A63FFB3FFD1FAF586041C856E08" box="[1340,1413,1323,1347]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
E, 3S); tip curved slightly upward; lower margin serrated. Dactylus long and slender, occlusal margin forming sharp keel with long quadrate blunt tooth proximally and one indented sharp tooth distally followed with several tiny teeth (
<figureCitation id="C0A82A63FFB3FFD1FF1586B818236EE4" box="[220,291,1431,1455]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
F, 3EI); tip bent slightly downward or forming sharp hook (
<figureCitation id="C0A82A63FFB3FFD1FC0286B81D0D6EE4" box="[971,1037,1431,1455]" captionStart="FIGURE 3" captionStartId="5.[151,250,1674,1697]" captionTargetBox="[279,1308,201,1641]" captionTargetId="figure@5.[272,1314,193,1645]" captionTargetPageId="5" captionText="FIGURE 3. Neocallichirus brocchii (Lőrenthey, 1897) new combination, major chelae (propodi and dactyli): A, left chela (robust morphotype), KGP-MH DH 001; B, right chela (slender morphotype), PCMH- 001; C D, right chela from inner and outer view (robust morphotype), PCRB-DH 008; E, left dactylus, PCRB-DH 001; F, left dactylus, PCRB-DH 002; G, right dactylus, KGP-MH DH 012; H, right dactylus, PCRB-DH 004; I, right dactylus, PCRB-DH 003; J, left chela (robust morphotype), KGP-MH DH 064, note serration on the lower margin (K); L M, fragmentary right chela (robust morphotype), PCRB-DH 005; N, left chela (robust morphotype), SNM Z- 37534, note serration on the lower margin (O); P, right chela (slender morphotype), PCRB-DH 010, note short carpus; Q, fragmented right chela (slender morphotype), PCMH- 003, note occlusal keel on the fixed finger positioned medially; R, right chela (slender morphotype), PCRB-DH 007, note occlusal keel on the fixed finger positioned medially; S, fragmented right chela (robust morphotype), PCMH- 002, note occlusal keel on the fixed finger positioned laterally; T, right chela, KGP-MH DH- 66, note the unusual length / height ratio. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/209487/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
G).
</paragraph>
<caption id="0CEC666EFFB0FFD2FF5E85A5186E6CB9" httpUri="https://zenodo.org/record/209487/files/figure.png" pageId="5" pageNumber="6" targetBox="[279,1308,201,1641]" targetPageId="5">
<paragraph id="582C36E6FFB0FFD2FF5E85A5186E6CB9" blockId="5.[151,1436,1674,2034]" pageId="5" pageNumber="6">
<emphasis id="6AE7EAF4FFB0FFD2FF5E85A518156DEB" bold="true" box="[151,277,1674,1697]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
<taxonomicName id="9F934D65FFB0FFD2FED685A51BD36DEA" authority="Lorenthey, 1897" authorityName="Lorenthey" authorityYear="1897" box="[287,723,1674,1697]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brocchii" status="comb. nov.">
<emphasis id="6AE7EAF4FFB0FFD2FED685A51B106DEB" box="[287,528,1674,1696]" italics="true" pageId="5" pageNumber="6">Neocallichirus brocchii</emphasis>
(Lőrenthey, 1897)
</taxonomicName>
<taxonomicNameLabel id="71D4578FFFB0FFD2FD1485A41A916DEB" box="[733,913,1674,1696]" pageId="5" pageNumber="6">new combination</taxonomicNameLabel>
, major chelae (propodi and dactyli): A, left chela (robust morphotype), KGP-MH DH001; B, right chela (slender morphotype), PCMH-001; CD, right chela from inner and outer view (robust morphotype), PCRB-DH008; E, left dactylus, PCRB-DH001; F, left dactylus, PCRB-DH002; G, right dactylus, KGP-MH DH012; H, right dactylus, PCRB-DH004; I, right dactylus, PCRB-DH003; J, left chela (robust morphotype), KGP-MH DH064, note serration on the lower margin (K); LM, fragmentary right chela (robust morphotype), PCRB-DH005; N, left chela (robust morphotype), SNM Z-37534, note serration on the lower margin (O); P, right chela (slender morphotype), PCRB-DH010, note short carpus; Q, fragmented right chela (slender morphotype), PCMH-003, note occlusal keel on the fixed finger positioned medially; R, right chela (slender morphotype), PCRB-DH007, note occlusal keel on the fixed finger positioned medially; S, fragmented right chela (robust morphotype), PCMH-002, note occlusal keel on the fixed finger positioned laterally; T, right chela, KGP-MH DH-66, note the unusual length/height ratio. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography.
</paragraph>
</caption>
<caption id="0CEC666EFFB3FFD1FF5E80C119F66FF7" httpUri="https://zenodo.org/record/209488/files/figure.png" pageId="6" pageNumber="7" targetBox="[160,1426,202,978]" targetPageId="6">
<paragraph id="582C36E6FFB3FFD1FF5E80C119F66FF7" blockId="6.[151,1436,1006,1212]" pageId="6" pageNumber="7">
<emphasis id="6AE7EAF4FFB3FFD1FF5E80C118136F48" bold="true" box="[151,275,1006,1029]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
<taxonomicName id="9F934D65FFB3FFD1FED380C11BC86F4E" authority="Lorenthey, 1897" authorityName="Lorenthey" authorityYear="1897" box="[282,712,1006,1029]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="brocchii" status="comb. nov.">
<emphasis id="6AE7EAF4FFB3FFD1FED380C11B0A6F4F" box="[282,522,1006,1028]" italics="true" pageId="6" pageNumber="7">Neocallichirus brocchii</emphasis>
(Lőrenthey, 1897)
</taxonomicName>
<taxonomicNameLabel id="71D4578FFFB3FFD1FD0680C01A826F4F" box="[719,898,1006,1028]" pageId="6" pageNumber="7">new combination</taxonomicNameLabel>
, major chelae (carpi and meri): A, left carpus, KGP- MH DH065; B, right carpus, KGP-MH DH060; C, left carpus, PCRB-DH009, note serration on the lower margin (D); E, left articulated merus and carpus, SNM Z-37549; F, left merus, PCRB-DH014; G, right merus, KGP-MH DH052; H, left merus, KGP-MH DH010; I, right merus, PCRB-DH016; J, left merus, PCRB-DH015; K, right merus, SNM Z-37542. All specimens belong to robust morphotype, except „I“ being a slender morphotype. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography.
</paragraph>
</caption>
<paragraph id="582C36E6FFB3FFD1FF0086921D276DCC" blockId="6.[151,1436,1251,2032]" pageId="6" pageNumber="7">
<emphasis id="6AE7EAF4FFB3FFD1FF0086921B396E9F" box="[201,569,1468,1492]" italics="true" pageId="6" pageNumber="7">Major chela slender morphotype</emphasis>
: Merus ovoid in shape, longer than high, highest at midlength; lower margin finely serrated without distinct tooth blade (
<figureCitation id="C0A82A63FFB3FFD1FD4D86F01BCC6EBC" box="[644,716,1503,1527]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
H,
<date id="2C2D1026FFB3FFD1FD2786F01A086EBC" box="[750,776,1503,1527]" pageId="6" pageNumber="7">4I</date>
). Carpus higher than long (L/H = 0.700.85) (
<figureCitation id="C0A82A63FFB3FFD1FADB86F01C5A6EBC" box="[1298,1370,1503,1527]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
B, 3P, 6C), keels on upper and lower margins not as pronounced as in robust morphotype; lower margin serrated. Propodus distinctly longer than carpus, general shape much the same as in robust morphotype (
<figureCitation id="C0A82A63FFB3FFD1FB4285081DD36D74" box="[1163,1235,1575,1599]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
B, 3B, 6C). Fixed finger with serrated keel on occlusal margin positioned in the middle (
<figureCitation id="C0A82A63FFB3FFD1FC6E85641AEF6D28" box="[935,1007,1611,1635]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
D, 3Q, R) instead of lateral position of robust morphotype. Dactylus similarly shaped as in robust morphotype (
<figureCitation id="C0A82A63FFB3FFD1FC0885401D046DCC" box="[961,1028,1647,1671]" captionStart="FIGURE 3" captionStartId="5.[151,250,1674,1697]" captionTargetBox="[279,1308,201,1641]" captionTargetId="figure@5.[272,1314,193,1645]" captionTargetPageId="5" captionText="FIGURE 3. Neocallichirus brocchii (Lőrenthey, 1897) new combination, major chelae (propodi and dactyli): A, left chela (robust morphotype), KGP-MH DH 001; B, right chela (slender morphotype), PCMH- 001; C D, right chela from inner and outer view (robust morphotype), PCRB-DH 008; E, left dactylus, PCRB-DH 001; F, left dactylus, PCRB-DH 002; G, right dactylus, KGP-MH DH 012; H, right dactylus, PCRB-DH 004; I, right dactylus, PCRB-DH 003; J, left chela (robust morphotype), KGP-MH DH 064, note serration on the lower margin (K); L M, fragmentary right chela (robust morphotype), PCRB-DH 005; N, left chela (robust morphotype), SNM Z- 37534, note serration on the lower margin (O); P, right chela (slender morphotype), PCRB-DH 010, note short carpus; Q, fragmented right chela (slender morphotype), PCMH- 003, note occlusal keel on the fixed finger positioned medially; R, right chela (slender morphotype), PCRB-DH 007, note occlusal keel on the fixed finger positioned medially; S, fragmented right chela (robust morphotype), PCMH- 002, note occlusal keel on the fixed finger positioned laterally; T, right chela, KGP-MH DH- 66, note the unusual length / height ratio. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/209487/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
B).
</paragraph>
<paragraph id="582C36E6FFB3FFD1FF0085BC182C6C14" blockId="6.[151,1436,1251,2032]" pageId="6" pageNumber="7">
Minor cheliped more slender than major one, smooth (
<figureCitation id="C0A82A63FFB3FFD1FCE385BC1A6D6DE0" box="[810,877,1683,1707]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
C). Ischium and merus not well preserved. Carpus slightly longer than high, higher than propodus (
<figureCitation id="C0A82A63FFB3FFD1FD7785981A076D84" box="[702,775,1719,1743]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
C, 6B); upper and lower margins convex. The palm of the propodus as long as high; distal margin without serration (
<figureCitation id="C0A82A63FFB3FFD1FCE285F41A716DB8" box="[811,881,1755,1779]" captionStart="FIGURE 5" captionStartId="7.[151,250,1431,1454]" captionTargetBox="[170,1427,388,1389]" captionTargetId="figure@7.[151,1436,373,1410]" captionTargetPageId="7" captionText="FIGURE 5. Neocallichirus brocchii (Lőrenthey, 1897) new combination, minor chelae (propodi and dactyli): A B, left chela from outer and inner view, KGP-MH DH 067; C, right chela, PCRB-DH 013; D, fragmented right chela, PCRB-DH 011; E, left dactylus, PCRB-DH 012. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/209489/files/figure.png" pageId="6" pageNumber="7">Figs 5</figureCitation>
AD); lower margin of propodus slightly convex at the point where fixed finger beggins. Fixed finger about as long as palm, slender, forming sharp tip, occlusal margin not armed. Dactylus long, slender; occlusal margin forming faint finely serrated keel, no bigger teeth present (
<figureCitation id="C0A82A63FFB3FFD1FF018468180B6C14" box="[200,267,1863,1887]" captionStart="FIGURE 5" captionStartId="7.[151,250,1431,1454]" captionTargetBox="[170,1427,388,1389]" captionTargetId="figure@7.[151,1436,373,1410]" captionTargetPageId="7" captionText="FIGURE 5. Neocallichirus brocchii (Lőrenthey, 1897) new combination, minor chelae (propodi and dactyli): A B, left chela from outer and inner view, KGP-MH DH 067; C, right chela, PCRB-DH 013; D, fragmented right chela, PCRB-DH 011; E, left dactylus, PCRB-DH 012. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/209489/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
E).
</paragraph>
<paragraph id="582C36E6FFB3FFD0FF0084441DC86A74" blockId="6.[151,1436,1251,2032]" lastBlockId="7.[151,1436,151,319]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="6AE7EAF4FFB3FFD1FF00844418496CCF" bold="true" box="[201,329,1899,1924]" pageId="6" pageNumber="7">Variations.</emphasis>
The studied material consists of the two morphotypes described above. The overall shape of the propodus and morphology of the dactylus and minor chela clearly demonstrate that both morphs belong to the same species. Both morphotypes occur together in the same strata, so they represent the same population (or sequence of populations). The recognized morphs may mirror sexual dimorphism (slender morphotype being a female one); however, it is very difficult to determine sex in the fossil state, especially when dealing with
<taxonomicName id="9F934D65FFB2FFD0FB0C83B71C6A6BFB" box="[1221,1386,152,176]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB2FFD0FB0C83B71C6A6BFB" box="[1221,1386,152,176]" italics="true" pageId="7" pageNumber="8">Neocallichirus</emphasis>
</taxonomicName>
. No obvious sexual dimorphism has been observed in several extant species (see
<bibRefCitation id="3C024B17FFB2FFD0FC3A83931DBF6B9F" author="Dworschak" box="[1011,1215,188,212]" pageId="7" pageNumber="8" refString="Dworschak, P. C. (2011 a) Redescription of Callianassa jousseaumei Nobili, 1904, a junior (sic!) subjective synonym of Callianassa indica de Man, 1905 with description of a new species of Neocallichirus (Decapoda: Axiidea: Callianassidae). Zootaxa, 2746, 1 - 19." type="journal article" year="2011" yearSuffix="a">Dworschak 2011a</bibRefCitation>
,
<bibRefCitation id="3C024B17FFB2FFD0FB0283931DDC6B9F" author="Dworschak" box="[1227,1244,188,212]" pageId="7" pageNumber="8" refString="Dworschak, P. C. (2011 b) Redescription of Callianassa vigilax de Man, 1916, a subjective synonym of Neocallichirus denticulatus Ngoc-Ho, 1994 (Crustacea: Decapoda: Callianassidae). Annalen des Naturhistorischen Museums in Wien, B 112, 137 - 151." type="journal article" year="2011" yearSuffix="b">b</bibRefCitation>
). Thus, the interpretation of morphs as sexual dimorphism should be considered as a preliminary one. Possible attribution of the slender morphotype to female individuals is based mostly on other callianassid taxa in which mature males usually possess more robust chelipeds than females (e.g.,
<bibRefCitation id="3C024B17FFB2FFD0FD0C82081AB86A74" author="Felder" box="[709,952,295,319]" pageId="7" pageNumber="8" refString="Felder, D. L. &amp; Lovett, D. L. (1989) Relative growth and sexual maturation in the estuarine ghost shrimp Callianassa louisianensis Schmitt, 1935. Journal of Crustacean Biology, 9 (4), 540 - 553." type="journal article" year="1989">Felder &amp; Lovett 1989</bibRefCitation>
and references therein).
</paragraph>
<caption id="0CEC666EFFB2FFD0FF5E86B81D3C6D41" httpUri="https://zenodo.org/record/209489/files/figure.png" pageId="7" pageNumber="8" targetBox="[170,1427,388,1389]" targetPageId="7">
<paragraph id="582C36E6FFB2FFD0FF5E86B81D3C6D41" blockId="7.[151,1437,1431,1546]" pageId="7" pageNumber="8">
<emphasis id="6AE7EAF4FFB2FFD0FF5E86B818136EE6" bold="true" box="[151,275,1431,1454]" pageId="7" pageNumber="8">FIGURE 5.</emphasis>
<taxonomicName id="9F934D65FFB2FFD0FED286B71BC96EE5" authority="Lorenthey, 1897" authorityName="Lorenthey" authorityYear="1897" box="[283,713,1432,1454]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="brocchii" status="comb. nov.">
<emphasis id="6AE7EAF4FFB2FFD0FED286B71B0B6EE5" box="[283,523,1432,1454]" italics="true" pageId="7" pageNumber="8">Neocallichirus brocchii</emphasis>
(Lőrenthey, 1897)
</taxonomicName>
<taxonomicNameLabel id="71D4578FFFB2FFD0FD1886B61A836EE5" box="[721,899,1432,1454]" pageId="7" pageNumber="8">new combination</taxonomicNameLabel>
, minor chelae (propodi and dactyli): AB, left chela from outer and inner view, KGP-MH DH067; C, right chela, PCRB-DH013; D, fragmented right chela, PCRB-DH011; E, left dactylus, PCRB-DH012. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography.
</paragraph>
</caption>
<paragraph id="582C36E6FFB2FFD0FF0085181D7B6C24" blockId="7.[151,1437,1591,2010]" pageId="7" pageNumber="8">
The merus shows variations in the number of teeth on its lower margin which seems to be a consequence of growth (more teeth are present on larger meri) rather than real variability. The variation in meral shape, notably the development of the meral blade, can be documented in several extant
<taxonomicName id="9F934D65FFB2FFD0FC0A85501D686DDC" box="[963,1128,1663,1687]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFB2FFD0FC0A85501D686DDC" box="[963,1128,1663,1687]" italics="true" pageId="7" pageNumber="8">Neocallichirus</emphasis>
</taxonomicName>
species, e.g. in
<taxonomicName id="9F934D65FFB2FFD0FAED85AF188F6DF1" authority="Gibbes, 1850" authorityName="Gibbes" authorityYear="1850" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="grandimanus">
<emphasis id="6AE7EAF4FFB2FFD0FAED85AF19E06DF0" italics="true" pageId="7" pageNumber="8">N. grandimanus</emphasis>
(
<bibRefCitation id="3C024B17FFB2FFD0FF26858D18876DF1" author="Gibbes" box="[239,391,1698,1722]" pageId="7" pageNumber="8" refString="Gibbes, L. R. (1850) On the carcinological collections of the United States, and an enumeration of the species contained in them, with notes on the most remarkable, and descriptions of new species. Proceedings of the American Association for the Advancement of Science, 3 rd meeting, 167 - 201." type="proceedings paper" year="1850">Gibbes, 1850</bibRefCitation>
)
</taxonomicName>
(compare e.g.
<bibRefCitation id="3C024B17FFB2FFD0FDFE858D1BE06DF1" author="Manning" box="[567,736,1698,1722]" pageId="7" pageNumber="8" refString="Manning, R. B. (1987) Notes on western Atlantic Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington, 100 (2), 386 - 401." type="journal article" year="1987">Manning 1987</bibRefCitation>
: fig. 2;
<bibRefCitation id="3C024B17FFB2FFD0FCFC858D1D4F6DF1" author="Lemaitre" box="[821,1103,1698,1722]" pageId="7" pageNumber="8" refString="Lemaitre, R. &amp; Ramos, G. E. (1992) A collectio of Thalassinidea (Crustacea: Decapoda) from the Pacific coast of Colombia, with description of a new species and a checklist of Eastern Pacific species. Proceedings of the Biological Society of Washington, 105, 343 - 358." type="journal article" year="1992">Lemaitre &amp; Ramos 1992</bibRefCitation>
: fig. 5;
<bibRefCitation id="3C024B17FFB2FFD0FB6C858D1C986DF1" author="Blanco-Rambla" box="[1189,1432,1698,1722]" pageId="7" pageNumber="8" refString="Blanco-Rambla, J. P. (2000) Cuatro nuevos registros de especies del genero Neocallichirus (Crustacea: Decapoda: Callianassidae) para aguas venezolanas. Boletin del Instituto Oceanographico de Venezuela, Universidad de Oriente 39 (1 - 2), 71 - 78." type="journal article" year="2000">Blanco-Rambla 2000</bibRefCitation>
: fig. 2) and
<taxonomicName id="9F934D65FFB2FFD0FED485E71B7A6D94" authority="Nobili, 1904" authorityName="Nobili" authorityYear="1904" box="[285,634,1735,1759]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="jousseaumei">
<emphasis id="6AE7EAF4FFB2FFD0FED485E718CC6D94" box="[285,460,1736,1759]" italics="true" pageId="7" pageNumber="8">N. jousseaumei</emphasis>
(
<bibRefCitation id="3C024B17FFB2FFD0FE1785E81B726D94" author="Nobili" box="[478,626,1735,1759]" pageId="7" pageNumber="8" refString="Nobili, G. (1904) Diagnoses preliminaires de vingt-huit especes nouvelles de Stomatopodes et Decapodes Macroures de la Mer Rouge. Bulletin du Museum d'Histoire Naturelle de Paris, 10, 230 - 238." type="journal article" year="1904">Nobili, 1904</bibRefCitation>
)
</taxonomicName>
(compare e.g.
<bibRefCitation id="3C024B17FFB2FFD0FCE585E81ABA6D94" author="Nobili" box="[812,954,1735,1759]" pageId="7" pageNumber="8" refString="Nobili, G. (1906) Faune carcinologique de la Mer Rouge: Decapodes et Stomatopodes. Annales des Sciences Naturelles, Zoologie, (9) 4, 1 - 347." type="journal article" year="1906">Nobili 1906</bibRefCitation>
: fig. 2;
<bibRefCitation id="3C024B17FFB2FFD0FBD385E81DBD6D94" author="Kensley" box="[1050,1213,1735,1759]" pageId="7" pageNumber="8" refString="Kensley, B. (1976) Records of mud-prawns (genus Callianassa) from South Africa and Mauritius (Crustacea, Decapoda, Thalassinidea). Annals of the South African Museum, 69, 47 - 58." type="journal article" year="1976">Kensley 1976</bibRefCitation>
: fig. 2;
<bibRefCitation id="3C024B17FFB2FFD0FAD485E819DC6C48" author="Dworschak" pageId="7" pageNumber="8" refString="Dworschak, P. C. (2011 a) Redescription of Callianassa jousseaumei Nobili, 1904, a junior (sic!) subjective synonym of Callianassa indica de Man, 1905 with description of a new species of Neocallichirus (Decapoda: Axiidea: Callianassidae). Zootaxa, 2746, 1 - 19." type="journal article" year="2011" yearSuffix="a">
Dworschak 2011
<emphasis id="6AE7EAF4FFB2FFD0FF0785C319DC6C48" box="[206,220,1772,1795]" italics="true" pageId="7" pageNumber="8">a</emphasis>
</bibRefCitation>
: figs. 14). In this respect the merus of the slender morphotype of
<taxonomicName id="9F934D65FFB2FFD0FC0185C41DD06C48" box="[968,1232,1771,1795]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="brocchii" status="comb. nov.">
<emphasis id="6AE7EAF4FFB2FFD0FC0185C41DD06C48" box="[968,1232,1771,1795]" italics="true" pageId="7" pageNumber="8">Neocallichirus brocchii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71D4578FFFB2FFD0FB1E85C51C566C49" box="[1239,1366,1770,1794]" pageId="7" pageNumber="8">comb. nov.</taxonomicNameLabel>
, without a pronounced meral blade (
<figureCitation id="C0A82A63FFB2FFD0FE3A84201B376C6C" box="[499,567,1807,1831]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
H), is similar to the merus of
<taxonomicName id="9F934D65FFB2FFD0FC49843F1D356C6C" box="[896,1077,1807,1831]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="grandimanus">
<emphasis id="6AE7EAF4FFB2FFD0FC49843F1D356C6C" box="[896,1077,1807,1831]" italics="true" pageId="7" pageNumber="8">N. grandimanus</emphasis>
</taxonomicName>
depicted by
<bibRefCitation id="3C024B17FFB2FFD0FB0C8420183E6C01" author="Lemaitre" pageId="7" pageNumber="8" refString="Lemaitre, R. &amp; Ramos, G. E. (1992) A collectio of Thalassinidea (Crustacea: Decapoda) from the Pacific coast of Colombia, with description of a new species and a checklist of Eastern Pacific species. Proceedings of the Biological Society of Washington, 105, 343 - 358." type="journal article" year="1992" yearSuffix="d">Lemaitre &amp; Ramos (1992: fig. 5d)</bibRefCitation>
, whereas the merus of the robust morphotype with a well developed meral blade (
<figureCitation id="C0A82A63FFB2FFD0FB21841D1C2C6C01" box="[1256,1324,1842,1866]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
G) can be compared with the merus of
<taxonomicName id="9F934D65FFB2FFD0FE1184771B8D6C24" box="[472,653,1879,1903]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="grandimanus">
<emphasis id="6AE7EAF4FFB2FFD0FE1184771B8D6C24" box="[472,653,1879,1903]" italics="true" pageId="7" pageNumber="8">N. grandimanus</emphasis>
</taxonomicName>
depicted by
<bibRefCitation id="3C024B17FFB2FFD0FCD584781D776C24" author="Blanco-Rambla" box="[796,1143,1879,1903]" pageId="7" pageNumber="8" refString="Blanco-Rambla, J. P. (2000) Cuatro nuevos registros de especies del genero Neocallichirus (Crustacea: Decapoda: Callianassidae) para aguas venezolanas. Boletin del Instituto Oceanographico de Venezuela, Universidad de Oriente 39 (1 - 2), 71 - 78." type="journal article" year="2000" yearSuffix="c">Blanco-Rambla (2000: fig. 2c)</bibRefCitation>
.
</paragraph>
<paragraph id="582C36E6FFB2FFD0FF0084551C0A6C91" blockId="7.[151,1437,1591,2010]" pageId="7" pageNumber="8">
There are slight variations in the outline of the distal margin of the propodus. Variations exist mainly in the nature of the incision under dactylus articulation. In the robust morphotype the incision is usually deeper and more rounded (e.g.
<figureCitation id="C0A82A63FFB2FFD0FEFB84ED18766C91" box="[306,374,1986,2010]" captionStart="FIGURE 3" captionStartId="5.[151,250,1674,1697]" captionTargetBox="[279,1308,201,1641]" captionTargetId="figure@5.[272,1314,193,1645]" captionTargetPageId="5" captionText="FIGURE 3. Neocallichirus brocchii (Lőrenthey, 1897) new combination, major chelae (propodi and dactyli): A, left chela (robust morphotype), KGP-MH DH 001; B, right chela (slender morphotype), PCMH- 001; C D, right chela from inner and outer view (robust morphotype), PCRB-DH 008; E, left dactylus, PCRB-DH 001; F, left dactylus, PCRB-DH 002; G, right dactylus, KGP-MH DH 012; H, right dactylus, PCRB-DH 004; I, right dactylus, PCRB-DH 003; J, left chela (robust morphotype), KGP-MH DH 064, note serration on the lower margin (K); L M, fragmentary right chela (robust morphotype), PCRB-DH 005; N, left chela (robust morphotype), SNM Z- 37534, note serration on the lower margin (O); P, right chela (slender morphotype), PCRB-DH 010, note short carpus; Q, fragmented right chela (slender morphotype), PCMH- 003, note occlusal keel on the fixed finger positioned medially; R, right chela (slender morphotype), PCRB-DH 007, note occlusal keel on the fixed finger positioned medially; S, fragmented right chela (robust morphotype), PCMH- 002, note occlusal keel on the fixed finger positioned laterally; T, right chela, KGP-MH DH- 66, note the unusual length / height ratio. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/209487/files/figure.png" pageId="7" pageNumber="8">Figs 3</figureCitation>
D, J, M), whereas in the slender morphotype, it is sometimes completely missing.
</paragraph>
<caption id="0CEC666EFFBDFFDFFF5E85E31C556C74" httpUri="https://zenodo.org/record/209490/files/figure.png" pageId="8" pageNumber="9" targetBox="[159,1424,199,1700]" targetPageId="8">
<paragraph id="582C36E6FFBDFFDFFF5E85E31C556C74" blockId="8.[151,1436,1740,1855]" pageId="8" pageNumber="9">
<emphasis id="6AE7EAF4FFBDFFDFFF5E85E318136DA9" bold="true" box="[151,275,1740,1763]" pageId="8" pageNumber="9">FIGURE 6.</emphasis>
<taxonomicName id="9F934D65FFBDFFDFFED285E21BC96DA8" authority="Lorenthey, 1897" authorityName="Lorenthey" authorityYear="1897" box="[283,713,1741,1763]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="brocchii" status="comb. nov.">
<emphasis id="6AE7EAF4FFBDFFDFFED285E21B0A6DA8" box="[283,522,1741,1763]" italics="true" pageId="8" pageNumber="9">Neocallichirus brocchii</emphasis>
(Lőrenthey, 1897)
</taxonomicName>
<taxonomicNameLabel id="71D4578FFFBDFFDFFD1885E11A846DA8" box="[721,900,1741,1763]" pageId="8" pageNumber="9">new combination</taxonomicNameLabel>
,
<emphasis id="6AE7EAF4FFBDFFDFFC4685E11ACF6DA8" box="[911,975,1742,1763]" italics="true" pageId="8" pageNumber="9">in situ</emphasis>
preservation of the specimen KGP-MH DH- 0 58 (Dúbravská hlavica locality): A, view of the individual (slender morphotype) preserved in a presumed burrow structure; B, minor cheliped; C, major cheliped; D, view from below where both chelipeds can be recognized. Note also the partial preservation of thoracopods P2P4. All specimens are to scale and were covered with ammonium chloride prior to photography.
</paragraph>
</caption>
<paragraph id="582C36E6FFBDFFDFFF0084A01BF36CA4" blockId="8.[151,1436,1935,2031]" pageId="8" pageNumber="9">
Other variations concern the cutting edge of the dactylus. The proximal quadrate tooth is usually divided in two secondary blunt teeth, the distal one being broader. The subdivision is, however, not always distinct and also the shape of the subdivided teeth is variable (
<figureCitation id="C0A82A63FFBDFFDFFD5884F81BD46CA4" box="[657,724,2007,2031]" captionStart="FIGURE 2" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[159,1428,205,1070]" captionTargetId="figure@4.[151,1436,195,1091]" captionTargetPageId="4" captionText="FIGURE 2. Neocallichirus brocchii (Lőrenthey, 1897) new combination, reconstructions: A, major cheliped of robust morphotype (outer view); B, major cheliped of slender morphotype (outer view); C, minor cheliped (outer view); D E, distal view of propodus of slender (D) and robust (E) morphotype, note differences in the development of medial keel on the fixed finger; dactyli and fixed finger tips are not depicted; F, variation in the armature of fixed finger; G H, outer view on merus of robust (G) and slender (H) morphotype. Reconstructions are based on several different specimens; figures A F are in scale." httpUri="https://zenodo.org/record/209486/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
F).
</paragraph>
<paragraph id="582C36E6FFBCFFDEFF0083B71B166A74" blockId="9.[151,1436,151,463]" pageId="9" pageNumber="10">
No obvious allometric growth in the nature of any cheliped element has been recognized, although there is one isolated propodus in the present collection (KGP-MH DH066;
<figureCitation id="C0A82A63FFBCFFDEFC9D83931A946B9F" box="[852,916,188,212]" captionStart="FIGURE 3" captionStartId="5.[151,250,1674,1697]" captionTargetBox="[279,1308,201,1641]" captionTargetId="figure@5.[272,1314,193,1645]" captionTargetPageId="5" captionText="FIGURE 3. Neocallichirus brocchii (Lőrenthey, 1897) new combination, major chelae (propodi and dactyli): A, left chela (robust morphotype), KGP-MH DH 001; B, right chela (slender morphotype), PCMH- 001; C D, right chela from inner and outer view (robust morphotype), PCRB-DH 008; E, left dactylus, PCRB-DH 001; F, left dactylus, PCRB-DH 002; G, right dactylus, KGP-MH DH 012; H, right dactylus, PCRB-DH 004; I, right dactylus, PCRB-DH 003; J, left chela (robust morphotype), KGP-MH DH 064, note serration on the lower margin (K); L M, fragmentary right chela (robust morphotype), PCRB-DH 005; N, left chela (robust morphotype), SNM Z- 37534, note serration on the lower margin (O); P, right chela (slender morphotype), PCRB-DH 010, note short carpus; Q, fragmented right chela (slender morphotype), PCMH- 003, note occlusal keel on the fixed finger positioned medially; R, right chela (slender morphotype), PCRB-DH 007, note occlusal keel on the fixed finger positioned medially; S, fragmented right chela (robust morphotype), PCMH- 002, note occlusal keel on the fixed finger positioned laterally; T, right chela, KGP-MH DH- 66, note the unusual length / height ratio. All specimens come from the Dúbravská hlavica locality (Vienna Basin, Slovakia), Middle Miocene. All specimens are to scale and were covered with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/209487/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
T) which shows a propodus length/height ratio (L/H = 0.6) rather different from all other studied specimens. The nature of the fixed finger (position of serrated keel) resembles that of the slender morphotype. The specimen itself is, however, much bigger than the usual size of the slender morphotype propodus.
</paragraph>
</subSubSection>
<subSubSection id="1089656DFFBCFFDDFF0E82641B2B6ED3" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="582C36E6FFBCFFDEFF0E82641C4F6A84" blockId="9.[151,1436,151,463]" pageId="9" pageNumber="10">
<emphasis id="6AE7EAF4FFBCFFDEFF0E826418B36A2F" bold="true" box="[199,435,331,356]" pageId="9" pageNumber="10">Material examined.</emphasis>
FI M.2329 (
<typeStatus id="87288844FFBCFFDEFD8082631BAD6A2F" box="[585,685,332,356]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
left propodus of robust morphotype articulated with dactylus); KGP-MH DH-001KGP-MH DH-015; M.86.249 (1 left and 1 right fragmented propodi;
<collectionCode id="3E82AE23FFBCFFDEFB4182401DC56ACC" box="[1160,1221,367,391]" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="9" pageNumber="10">SNM</collectionCode>
<collectionCode id="3E82AE23FFBCFFDEFB0582401DE66ACC" box="[1228,1254,367,391]" pageId="9" pageNumber="10">Z-</collectionCode>
7943 (collective number),
<collectionCode id="3E82AE23FFBCFFDEFECC82BB18426AE7" box="[261,322,404,428]" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="9" pageNumber="10">SNM</collectionCode>
<collectionCode id="3E82AE23FFBCFFDEFE8382BB18646AE7" box="[330,356,404,428]" pageId="9" pageNumber="10">Z-</collectionCode>
16419 (collective number),
<collectionCode id="3E82AE23FFBCFFDEFD6982BB1BDD6AE7" box="[672,733,404,428]" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="9" pageNumber="10">SNM</collectionCode>
<collectionCode id="3E82AE23FFBCFFDEFD2C82BB1A006AE7" box="[741,768,404,428]" pageId="9" pageNumber="10">Z-</collectionCode>
16433 (collective number),
<collectionCode id="3E82AE23FFBCFFDEFBF282BB1D786AE7" box="[1083,1144,404,428]" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="9" pageNumber="10">SNM</collectionCode>
<collectionCode id="3E82AE23FFBCFFDEFB4982BB1D9A6AE7" box="[1152,1178,404,428]" pageId="9" pageNumber="10">Z-</collectionCode>
37532
<collectionCode id="3E82AE23FFBCFFDEFB2682BB1C2C6AE7" box="[1263,1324,404,428]" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="9" pageNumber="10">SNM</collectionCode>
<collectionCode id="3E82AE23FFBCFFDEFAFD82BB1C4F6AE7" box="[1332,1359,404,428]" pageId="9" pageNumber="10">Z-</collectionCode>
37553; PCMH-001PCMH-003;
<collectionCode id="3E82AE23FFBCFFDEFE7082981B006A84" box="[441,512,439,463]" pageId="9" pageNumber="10">PCRB</collectionCode>
DH-001
<collectionCode id="3E82AE23FFBCFFDEFDB982981BB96A84" box="[624,697,439,463]" pageId="9" pageNumber="10">PCRB</collectionCode>
DH-016. For measurements and all the details see
<tableCitation id="1511035DFFBCFFDEFB3C82981C496A84" box="[1269,1353,439,463]" captionStart="TABLE 1" captionStartId="9.[151,235,506,527]" captionTargetBox="[151,1436,657,1332]" captionTargetPageId="9" captionText="TABLE 1. Measurements of selected well preserved specimens of Neocallichirus brocchii (Lőrenthey, 1897) comb. nov. Handedness: L = left; R = right; measurements: L = max. length; H = max. height; ~ = approximate value;&gt; = &quot; more than &quot; (in case the approximation is not possible due to the state of preservation); values are in mm." httpUri="http://table.plazi.org/id/0CEC666EFFBCFFDEFF5E82D51AEF6905" pageId="9" pageNumber="10" tableUuid="0CEC666EFFBCFFDEFF5E82D51AEF6905">Table 1</tableCitation>
.
</paragraph>
<caption id="0CEC666EFFBCFFDEFF5E82D51AEF6905" ID-Table-UUID="0CEC666EFFBCFFDEFF5E82D51AEF6905" httpUri="http://table.plazi.org/id/0CEC666EFFBCFFDEFF5E82D51AEF6905" pageId="9" pageNumber="10" targetBox="[151,1436,657,1332]" targetIsTable="true" targetPageId="9">
<paragraph id="582C36E6FFBCFFDEFF5E82D51AEF6905" blockId="9.[151,1436,506,638]" pageId="9" pageNumber="10">
<emphasis id="6AE7EAF4FFBCFFDEFF5E82D51804695B" bold="true" box="[151,260,506,528]" pageId="9" pageNumber="10">TABLE 1.</emphasis>
Measurements of selected well preserved specimens of
<taxonomicName id="9F934D65FFBCFFDEFCF182D51DE2695A" authority="Lorenthey, 1897" authorityName="Lorenthey" authorityYear="1897" box="[824,1250,506,529]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brocchii" status="comb. nov.">
<emphasis id="6AE7EAF4FFBCFFDEFCF182D51D27695B" box="[824,1063,506,528]" italics="true" pageId="9" pageNumber="10">Neocallichirus brocchii</emphasis>
(Lőrenthey, 1897)
</taxonomicName>
<taxonomicNameLabel id="71D4578FFFBCFFDEFB2182D51C58695B" box="[1256,1368,506,528]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
Handedness: L = left; R = right; measurements: L = max. length; H = max. height; ~ = approximate value;&gt; = &quot;more than&quot; (in case the approximation is not possible due to the state of preservation); values are in mm.
</paragraph>
</caption>
<paragraph id="582C36E6FFBCFFDDFCC581461A526E3F" blockId="9.[151,1436,506,638]" lastBlockId="10.[151,1437,1264,2009]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
merus carpus propodus dactylus
<collectionCode id="3E82AE23FFBCFFDEFF56866819D66E17" box="[159,214,1351,1372]" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="9" pageNumber="10">SNM</collectionCode>
<collectionCode id="3E82AE23FFBCFFDEFF15866819F46E16" box="[220,244,1351,1373]" pageId="9" pageNumber="10">Z-</collectionCode>
37548 unknown R 8.7 5.0 - - - - - continued next page
<emphasis id="6AE7EAF4FFBFFFDDFF0E87DF18596E42" bold="true" box="[199,345,1264,1289]" pageId="10" pageNumber="11">Occurrence.</emphasis>
The species has been reported only from the Serravallian (Middle Miocene), specifically from the upper 'Badenian' of the Rákos Formation localities Rákos, Budafok and Gyakorló út in the Budapest City area (
<collectingCountry id="20847676FFBFFFDDFF57861718036E1B" box="[158,259,1336,1360]" name="Hungary" pageId="10" pageNumber="11">Hungary</collectingCountry>
) (
<bibRefCitation id="3C024B17FFBFFFDDFED486181B5C6E1B" author="Lorenthey" box="[285,604,1335,1360]" pageId="10" pageNumber="11" refString="Lorenthey, E. &amp; Beurlen, K. (1929) Die fossilen Dekapoden der Lander der Ungarischen Krone. Geologica Hungarica, Series Palaeontologica, 3, 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen 1929</bibRefCitation>
;
<bibRefCitation id="3C024B17FFBFFFDDFDA286171BFE6E1B" author="Muller" box="[619,766,1336,1360]" pageId="10" pageNumber="11" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller 1984</bibRefCitation>
); and the lower 'Sarmatian' of the Studienka Formation locality Dúbravská hlavica in the Bratislava district (
<collectingCountry id="20847676FFBFFFDDFD2C86731A4A6E3F" box="[741,842,1372,1396]" name="Slovakia" pageId="10" pageNumber="11">Slovakia</collectingCountry>
)
</paragraph>
<paragraph id="582C36E6FFBCFFDEFF5681BE1C266E7F" pageId="9" pageNumber="10">
<table id="2A93C446FFBC0028FF5E81BE1C9C6E7F" box="[151,1436,657,1332]" gridcols="8" gridrows="12" pageId="9" pageNumber="10">
<tr id="E6A334A4FFBC0028FF5E81BE1C9C69EC" box="[151,1436,657,679]" gridrow="0" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E81BE183369EC" box="[151,307,657,679]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">Specimen</th>
<th id="A5725DD8FFBC0028FE6C81BE1B2669EC" box="[421,550,657,679]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">morphotype</th>
<th id="A5725DD8FFBC0028FD9481BE1BD269EC" box="[605,722,657,679]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">handedness</th>
<th id="A5725DD8FFBC0028FCC581BE1A1B69EC" box="[780,795,657,679]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">L</th>
<th id="A5725DD8FFBC0028FCA781BE1A8069EC" box="[878,896,657,679]" gridcol="4" gridrow="0" pageId="9" pageNumber="10">H</th>
<th id="A5725DD8FFBC0028FC7B81BE1ADE69EC" box="[946,990,657,679]" gridcol="5" gridrow="0" pageId="9" pageNumber="10">L</th>
<th id="A5725DD8FFBC0028FBDD81BE1D4169EC" box="[1044,1089,657,679]" gridcol="6" gridrow="0" pageId="9" pageNumber="10">H</th>
<th id="A5725DD8FFBC0028FBBE81BE1C9C69EC" box="[1143,1436,657,679]" gridcol="7" gridrow="0" pageId="9" pageNumber="10">L H L</th>
</tr>
<tr id="E6A334A4FFBC0028FF5E81951C9C699B" box="[151,1436,698,720]" gridrow="1" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E81951833699B" box="[151,307,698,720]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">SNM Z-37532</th>
<td id="A5725DD8FFBC0028FE6C81951B26699B" box="[421,550,698,720]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9481951BD2699B" box="[605,722,698,720]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">L</td>
<td id="A5725DD8FFBC0028FCC581951A1B699B" box="[780,795,698,720]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA781951A80699B" box="[878,896,698,720]" gridcol="4" gridrow="1" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B81951ADE699B" box="[946,990,698,720]" gridcol="5" gridrow="1" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD81951D41699B" box="[1044,1089,698,720]" gridcol="6" gridrow="1" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE81951C9C699B" box="[1143,1436,698,720]" gridcol="7" gridrow="1" pageId="9" pageNumber="10">14.0 14.4&gt;9.4</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E81CC1C9C6801" box="[151,1436,739,842]" gridrow="2" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E81CC18336801" box="[151,307,739,842]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">SNM Z-37533 SNM Z-37534 SNM Z-37535</th>
<td id="A5725DD8FFBC0028FE6C81CC1B266801" box="[421,550,739,842]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">major robust major robust major robust</td>
<td id="A5725DD8FFBC0028FD9481CC1BD26801" box="[605,722,739,842]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">L L L</td>
<td id="A5725DD8FFBC0028FCC581CC1A1B6801" box="[780,795,739,842]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FCA781CC1A806801" box="[878,896,739,842]" gridcol="4" gridrow="2" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FC7B81CC1ADE6801" box="[946,990,739,842]" gridcol="5" gridrow="2" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FBDD81CC1D416801" box="[1044,1089,739,842]" gridcol="6" gridrow="2" pageId="9" pageNumber="10">- - 15.7</td>
<td id="A5725DD8FFBC0028FBBE81CC1C9C6801" box="[1143,1436,739,842]" gridcol="7" gridrow="2" pageId="9" pageNumber="10">10.8 18.3 - ~13&gt;12 9.0 18.5 17.0&gt;8.0</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E80721C9C6838" box="[151,1436,861,883]" gridrow="3" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E807218336838" box="[151,307,861,883]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">SNM Z-37536</th>
<td id="A5725DD8FFBC0028FE6C80721B266838" box="[421,550,861,883]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9480721BD26838" box="[605,722,861,883]" gridcol="2" gridrow="3" pageId="9" pageNumber="10">L</td>
<td id="A5725DD8FFBC0028FCC580721A1B6838" box="[780,795,861,883]" gridcol="3" gridrow="3" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA780721A806838" box="[878,896,861,883]" gridcol="4" gridrow="3" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B80721ADE6838" box="[946,990,861,883]" gridcol="5" gridrow="3" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD80721D416838" box="[1044,1089,861,883]" gridcol="6" gridrow="3" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE80721C9C6838" box="[1143,1436,861,883]" gridcol="7" gridrow="3" pageId="9" pageNumber="10">18.3 17.4 -</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E80A91C9C68D7" box="[151,1436,902,924]" gridrow="4" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E80A9183368D7" box="[151,307,902,924]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">SNM Z-37537</th>
<td id="A5725DD8FFBC0028FE6C80A91B2668D7" box="[421,550,902,924]" gridcol="1" gridrow="4" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9480A91BD268D7" box="[605,722,902,924]" gridcol="2" gridrow="4" pageId="9" pageNumber="10">L</td>
<td id="A5725DD8FFBC0028FCC580A91A1B68D7" box="[780,795,902,924]" gridcol="3" gridrow="4" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA780A91A8068D7" box="[878,896,902,924]" gridcol="4" gridrow="4" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B80A91ADE68D7" box="[946,990,902,924]" gridcol="5" gridrow="4" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD80A91D4168D7" box="[1044,1089,902,924]" gridcol="6" gridrow="4" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE80A91C9C68D7" box="[1143,1436,902,924]" gridcol="7" gridrow="4" pageId="9" pageNumber="10">13.0 13.2 -</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E80801C9C688E" box="[151,1436,943,965]" gridrow="5" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E80801833688E" box="[151,307,943,965]" gridcol="0" gridrow="5" pageId="9" pageNumber="10">SNM Z-37538</th>
<td id="A5725DD8FFBC0028FE6C80801B26688E" box="[421,550,943,965]" gridcol="1" gridrow="5" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9480801BD2688E" box="[605,722,943,965]" gridcol="2" gridrow="5" pageId="9" pageNumber="10">L</td>
<td id="A5725DD8FFBC0028FCC580801A1B688E" box="[780,795,943,965]" gridcol="3" gridrow="5" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA780801A80688E" box="[878,896,943,965]" gridcol="4" gridrow="5" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B80801ADE688E" box="[946,990,943,965]" gridcol="5" gridrow="5" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD80801D41688E" box="[1044,1089,943,965]" gridcol="6" gridrow="5" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE80801C9C688E" box="[1143,1436,943,965]" gridcol="7" gridrow="5" pageId="9" pageNumber="10">- 15.0 -</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E80F71C9C6F5D" box="[151,1436,984,1046]" gridrow="6" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E80F718336F5D" box="[151,307,984,1046]" gridcol="0" gridrow="6" pageId="9" pageNumber="10">SNM Z-37539 SNM Z-37540</th>
<td id="A5725DD8FFBC0028FE6C80F71B266F5D" box="[421,550,984,1046]" gridcol="1" gridrow="6" pageId="9" pageNumber="10">major robust major robust</td>
<td id="A5725DD8FFBC0028FD9480F71BD26F5D" box="[605,722,984,1046]" gridcol="2" gridrow="6" pageId="9" pageNumber="10">L R</td>
<td id="A5725DD8FFBC0028FCC580F71A1B6F5D" box="[780,795,984,1046]" gridcol="3" gridrow="6" pageId="9" pageNumber="10">- -</td>
<td id="A5725DD8FFBC0028FCA780F71A806F5D" box="[878,896,984,1046]" gridcol="4" gridrow="6" pageId="9" pageNumber="10">- -</td>
<td id="A5725DD8FFBC0028FC7B80F71ADE6F5D" box="[946,990,984,1046]" gridcol="5" gridrow="6" pageId="9" pageNumber="10">- -</td>
<td id="A5725DD8FFBC0028FBDD80F71D416F5D" box="[1044,1089,984,1046]" gridcol="6" gridrow="6" pageId="9" pageNumber="10">- -</td>
<td id="A5725DD8FFBC0028FBBE80F71C9C6F5D" box="[1143,1436,984,1046]" gridcol="7" gridrow="6" pageId="9" pageNumber="10">11.4 11.4 - 14.7 15.0&gt;8.0</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E87061C9C6F74" box="[151,1436,1065,1087]" gridrow="7" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E870618336F74" box="[151,307,1065,1087]" gridcol="0" gridrow="7" pageId="9" pageNumber="10">SNM Z-37541</th>
<td id="A5725DD8FFBC0028FE6C87061B266F74" box="[421,550,1065,1087]" gridcol="1" gridrow="7" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9487061BD26F74" box="[605,722,1065,1087]" gridcol="2" gridrow="7" pageId="9" pageNumber="10">R</td>
<td id="A5725DD8FFBC0028FCC587061A1B6F74" box="[780,795,1065,1087]" gridcol="3" gridrow="7" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA787061A806F74" box="[878,896,1065,1087]" gridcol="4" gridrow="7" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B87061ADE6F74" box="[946,990,1065,1087]" gridcol="5" gridrow="7" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD87061D416F74" box="[1044,1089,1065,1087]" gridcol="6" gridrow="7" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE87061C9C6F74" box="[1143,1436,1065,1087]" gridcol="7" gridrow="7" pageId="9" pageNumber="10">14.4 14.0 -</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E877D1C9C6F23" box="[151,1436,1106,1128]" gridrow="8" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E877D18336F23" box="[151,307,1106,1128]" gridcol="0" gridrow="8" pageId="9" pageNumber="10">SNM Z-37542</th>
<td id="A5725DD8FFBC0028FE6C877D1B266F23" box="[421,550,1106,1128]" gridcol="1" gridrow="8" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD94877D1BD26F23" box="[605,722,1106,1128]" gridcol="2" gridrow="8" pageId="9" pageNumber="10">R</td>
<td id="A5725DD8FFBC0028FCC5877D1A1B6F23" box="[780,795,1106,1128]" gridcol="3" gridrow="8" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA7877D1A806F23" box="[878,896,1106,1128]" gridcol="4" gridrow="8" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B877D1ADE6F23" box="[946,990,1106,1128]" gridcol="5" gridrow="8" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD877D1D416F23" box="[1044,1089,1106,1128]" gridcol="6" gridrow="8" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE877D1C9C6F23" box="[1143,1436,1106,1128]" gridcol="7" gridrow="8" pageId="9" pageNumber="10">17.6 14.2 -</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E87541C9C6FA9" box="[151,1436,1147,1250]" gridrow="9" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E875418336FA9" box="[151,307,1147,1250]" gridcol="0" gridrow="9" pageId="9" pageNumber="10">SNM Z-37543 SNM Z-37544 SNM Z-37545</th>
<td id="A5725DD8FFBC0028FE6C87541B266FA9" box="[421,550,1147,1250]" gridcol="1" gridrow="9" pageId="9" pageNumber="10">major robust major robust major robust</td>
<td id="A5725DD8FFBC0028FD9487541BD26FA9" box="[605,722,1147,1250]" gridcol="2" gridrow="9" pageId="9" pageNumber="10">R R R</td>
<td id="A5725DD8FFBC0028FCC587541A1B6FA9" box="[780,795,1147,1250]" gridcol="3" gridrow="9" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FCA787541A806FA9" box="[878,896,1147,1250]" gridcol="4" gridrow="9" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FC7B87541ADE6FA9" box="[946,990,1147,1250]" gridcol="5" gridrow="9" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FBDD87541D416FA9" box="[1044,1089,1147,1250]" gridcol="6" gridrow="9" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FBBE87541C9C6FA9" box="[1143,1436,1147,1250]" gridcol="7" gridrow="9" pageId="9" pageNumber="10">13.2 12.2 - 17.0 17.2 - 18.0 17.0 -</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E87DA1C9C6E40" box="[151,1436,1269,1291]" gridrow="10" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E87DA18336E40" box="[151,307,1269,1291]" gridcol="0" gridrow="10" pageId="9" pageNumber="10">SNM Z-37546</th>
<td id="A5725DD8FFBC0028FE6C87DA1B266E40" box="[421,550,1269,1291]" gridcol="1" gridrow="10" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9487DA1BD26E40" box="[605,722,1269,1291]" gridcol="2" gridrow="10" pageId="9" pageNumber="10">L</td>
<td id="A5725DD8FFBC0028FCC587DA1A1B6E40" box="[780,795,1269,1291]" gridcol="3" gridrow="10" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA787DA1A806E40" box="[878,896,1269,1291]" gridcol="4" gridrow="10" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B87DA1ADE6E40" box="[946,990,1269,1291]" gridcol="5" gridrow="10" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD87DA1D416E40" box="[1044,1089,1269,1291]" gridcol="6" gridrow="10" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE87DA1C9C6E40" box="[1143,1436,1269,1291]" gridcol="7" gridrow="10" pageId="9" pageNumber="10">- - 12.6</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E86311C9C6E7F" box="[151,1436,1310,1332]" gridrow="11" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E863118336E7F" box="[151,307,1310,1332]" gridcol="0" gridrow="11" pageId="9" pageNumber="10">SNM Z-37547</th>
<td id="A5725DD8FFBC0028FE6C86311B266E7F" box="[421,550,1310,1332]" gridcol="1" gridrow="11" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9486311BD26E7F" box="[605,722,1310,1332]" gridcol="2" gridrow="11" pageId="9" pageNumber="10">L</td>
<td id="A5725DD8FFBC0028FCC586311A1B6E7F" box="[780,795,1310,1332]" gridcol="3" gridrow="11" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA786311A806E7F" box="[878,896,1310,1332]" gridcol="4" gridrow="11" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B86311ADE6E7F" box="[946,990,1310,1332]" gridcol="5" gridrow="11" pageId="9" pageNumber="10">11.3</td>
<td id="A5725DD8FFBC0028FBDD86311D416E7F" box="[1044,1089,1310,1332]" gridcol="6" gridrow="11" pageId="9" pageNumber="10">14.1</td>
<td id="A5725DD8FFBC0028FBBE86311C9C6E7F" box="[1143,1436,1310,1332]" gridcol="7" gridrow="11" pageId="9" pageNumber="10">- - -</td>
</tr>
</table>
</paragraph>
<paragraph id="582C36E6FFBCFFDEFF56865F1C256C8A" pageId="9" pageNumber="10">
<table id="2A93C446FFBC0028FF5E865F1C9C6C8A" box="[151,1436,1392,1985]" gridcols="10" gridrows="11" pageId="9" pageNumber="10">
<tr id="E6A334A4FFBC0028FF5E865F1C9C6EE5" box="[151,1436,1392,1454]" gridrow="0" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E865F185C6EE5" box="[151,348,1392,1454]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">SNM Z-37549 SNM Z-37550</th>
<th id="A5725DD8FFBC0028FE6C865F1B306EE5" box="[421,560,1392,1454]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">major robust major slender</th>
<th id="A5725DD8FFBC0028FD94865F1B6E6EE5" box="[605,622,1392,1454]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">L R</th>
<th id="A5725DD8FFBC0028FCC5865F1A466EE5" box="[780,838,1392,1454]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">14 -</th>
<th id="A5725DD8FFBC0028FCA7865F1A8E6EE5" box="[878,910,1392,1454]" gridcol="4" gridrow="0" pageId="9" pageNumber="10">9.6 -</th>
<th id="A5725DD8FFBC0028FC7B865F1AED6EE5" box="[946,1005,1392,1454]" gridcol="5" gridrow="0" pageId="9" pageNumber="10">17.4 6.8</th>
<th id="A5725DD8FFBC0028FBDD865F1D416EE5" box="[1044,1089,1392,1454]" gridcol="6" gridrow="0" pageId="9" pageNumber="10">15.7 8.4</th>
<th id="A5725DD8FFBC0028FBBE865F1DA56EE5" box="[1143,1189,1392,1454]" gridcol="7" gridrow="0" pageId="9" pageNumber="10">~20 -</th>
<th id="A5725DD8FFBC0028FB02865F1DF76EE5" box="[1227,1271,1392,1454]" gridcol="8" gridrow="0" pageId="9" pageNumber="10">14.0 -</th>
<th id="A5725DD8FFBC0028FAD4865F1C9C6EE5" box="[1309,1436,1392,1454]" gridcol="9" gridrow="0" pageId="9" pageNumber="10">&gt;10.0 -</th>
</tr>
<tr id="E6A334A4FFBC0028FF5E86EE1C9C6E9C" box="[151,1436,1473,1495]" gridrow="1" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E86EE185C6E9C" box="[151,348,1473,1495]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">SNM Z-37551</th>
<td id="A5725DD8FFBC0028FE6C86EE1B306E9C" box="[421,560,1473,1495]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">major slender</td>
<td id="A5725DD8FFBC0028FD9486EE1B6E6E9C" box="[605,622,1473,1495]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">L</td>
<td id="A5725DD8FFBC0028FCC586EE1A466E9C" box="[780,838,1473,1495]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA786EE1A8E6E9C" box="[878,910,1473,1495]" gridcol="4" gridrow="1" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B86EE1AED6E9C" box="[946,1005,1473,1495]" gridcol="5" gridrow="1" pageId="9" pageNumber="10">5.5</td>
<td id="A5725DD8FFBC0028FBDD86EE1D416E9C" box="[1044,1089,1473,1495]" gridcol="6" gridrow="1" pageId="9" pageNumber="10">8.2</td>
<td id="A5725DD8FFBC0028FBBE86EE1DA56E9C" box="[1143,1189,1473,1495]" gridcol="7" gridrow="1" pageId="9" pageNumber="10">&gt;8.0</td>
<td id="A5725DD8FFBC0028FB0286EE1DF76E9C" box="[1227,1271,1473,1495]" gridcol="8" gridrow="1" pageId="9" pageNumber="10">8.5</td>
<td id="A5725DD8FFBC0028FAD486EE1C9C6E9C" box="[1309,1436,1473,1495]" gridcol="9" gridrow="1" pageId="9" pageNumber="10">-</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E86C51C9C6D4B" box="[151,1436,1514,1536]" gridrow="2" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E86C5185C6D4B" box="[151,348,1514,1536]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">SNM Z-37552</th>
<td id="A5725DD8FFBC0028FE6C86C51B306D4B" box="[421,560,1514,1536]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9486C51B6E6D4B" box="[605,622,1514,1536]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">L</td>
<td id="A5725DD8FFBC0028FCC586C51A466D4B" box="[780,838,1514,1536]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA786C51A8E6D4B" box="[878,910,1514,1536]" gridcol="4" gridrow="2" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B86C51AED6D4B" box="[946,1005,1514,1536]" gridcol="5" gridrow="2" pageId="9" pageNumber="10">17.5</td>
<td id="A5725DD8FFBC0028FBDD86C51D416D4B" box="[1044,1089,1514,1536]" gridcol="6" gridrow="2" pageId="9" pageNumber="10">17.1</td>
<td id="A5725DD8FFBC0028FBBE86C51DA56D4B" box="[1143,1189,1514,1536]" gridcol="7" gridrow="2" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FB0286C51DF76D4B" box="[1227,1271,1514,1536]" gridcol="8" gridrow="2" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FAD486C51C9C6D4B" box="[1309,1436,1514,1536]" gridcol="9" gridrow="2" pageId="9" pageNumber="10">-</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E853C1C9C6D31" box="[151,1436,1555,1658]" gridrow="3" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E853C185C6D31" box="[151,348,1555,1658]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">SNM Z-37553 KGP-MH DH-001 KGP-MH DH-002</th>
<td id="A5725DD8FFBC0028FE6C853C1B306D31" box="[421,560,1555,1658]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">minor major robust major robust</td>
<td id="A5725DD8FFBC0028FD94853C1B6E6D31" box="[605,622,1555,1658]" gridcol="2" gridrow="3" pageId="9" pageNumber="10">R L R</td>
<td id="A5725DD8FFBC0028FCC5853C1A466D31" box="[780,838,1555,1658]" gridcol="3" gridrow="3" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FCA7853C1A8E6D31" box="[878,910,1555,1658]" gridcol="4" gridrow="3" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FC7B853C1AED6D31" box="[946,1005,1555,1658]" gridcol="5" gridrow="3" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FBDD853C1D416D31" box="[1044,1089,1555,1658]" gridcol="6" gridrow="3" pageId="9" pageNumber="10">- - -</td>
<td id="A5725DD8FFBC0028FBBE853C1DA56D31" box="[1143,1189,1555,1658]" gridcol="7" gridrow="3" pageId="9" pageNumber="10">8.6 15.0 14.3</td>
<td id="A5725DD8FFBC0028FB02853C1DF76D31" box="[1227,1271,1555,1658]" gridcol="8" gridrow="3" pageId="9" pageNumber="10">8.5 14.2 13.0</td>
<td id="A5725DD8FFBC0028FAD4853C1C9C6D31" box="[1309,1436,1555,1658]" gridcol="9" gridrow="3" pageId="9" pageNumber="10">- 10.5 -</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E85A21C9C6DE8" box="[151,1436,1677,1699]" gridrow="4" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E85A2185C6DE8" box="[151,348,1677,1699]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">KGP-MH DH-003</th>
<td id="A5725DD8FFBC0028FE6C85A21B306DE8" box="[421,560,1677,1699]" gridcol="1" gridrow="4" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9485A21B6E6DE8" box="[605,622,1677,1699]" gridcol="2" gridrow="4" pageId="9" pageNumber="10">L</td>
<td id="A5725DD8FFBC0028FCC585A21A466DE8" box="[780,838,1677,1699]" gridcol="3" gridrow="4" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA785A21A8E6DE8" box="[878,910,1677,1699]" gridcol="4" gridrow="4" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B85A21AED6DE8" box="[946,1005,1677,1699]" gridcol="5" gridrow="4" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD85A21D416DE8" box="[1044,1089,1677,1699]" gridcol="6" gridrow="4" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE85A21DA56DE8" box="[1143,1189,1677,1699]" gridcol="7" gridrow="4" pageId="9" pageNumber="10">&gt;15</td>
<td id="A5725DD8FFBC0028FB0285A21DF76DE8" box="[1227,1271,1677,1699]" gridcol="8" gridrow="4" pageId="9" pageNumber="10">15.3</td>
<td id="A5725DD8FFBC0028FAD485A21C9C6DE8" box="[1309,1436,1677,1699]" gridcol="9" gridrow="4" pageId="9" pageNumber="10">-</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E85991C9C6D87" box="[151,1436,1718,1740]" gridrow="5" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E8599185C6D87" box="[151,348,1718,1740]" gridcol="0" gridrow="5" pageId="9" pageNumber="10">KGP-MH DH-004</th>
<td id="A5725DD8FFBC0028FE6C85991B306D87" box="[421,560,1718,1740]" gridcol="1" gridrow="5" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9485991B6E6D87" box="[605,622,1718,1740]" gridcol="2" gridrow="5" pageId="9" pageNumber="10">R</td>
<td id="A5725DD8FFBC0028FCC585991A466D87" box="[780,838,1718,1740]" gridcol="3" gridrow="5" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA785991A8E6D87" box="[878,910,1718,1740]" gridcol="4" gridrow="5" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B85991AED6D87" box="[946,1005,1718,1740]" gridcol="5" gridrow="5" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD85991D416D87" box="[1044,1089,1718,1740]" gridcol="6" gridrow="5" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE85991DA56D87" box="[1143,1189,1718,1740]" gridcol="7" gridrow="5" pageId="9" pageNumber="10">11.3</td>
<td id="A5725DD8FFBC0028FB0285991DF76D87" box="[1227,1271,1718,1740]" gridcol="8" gridrow="5" pageId="9" pageNumber="10">11.0</td>
<td id="A5725DD8FFBC0028FAD485991C9C6D87" box="[1309,1436,1718,1740]" gridcol="9" gridrow="5" pageId="9" pageNumber="10">-</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E85F01C9C6DBE" box="[151,1436,1759,1781]" gridrow="6" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E85F0185C6DBE" box="[151,348,1759,1781]" gridcol="0" gridrow="6" pageId="9" pageNumber="10">KGP-MH DH-005</th>
<td id="A5725DD8FFBC0028FE6C85F01B306DBE" box="[421,560,1759,1781]" gridcol="1" gridrow="6" pageId="9" pageNumber="10">major slender</td>
<td id="A5725DD8FFBC0028FD9485F01B6E6DBE" box="[605,622,1759,1781]" gridcol="2" gridrow="6" pageId="9" pageNumber="10">R</td>
<td id="A5725DD8FFBC0028FCC585F01A466DBE" box="[780,838,1759,1781]" gridcol="3" gridrow="6" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA785F01A8E6DBE" box="[878,910,1759,1781]" gridcol="4" gridrow="6" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B85F01AED6DBE" box="[946,1005,1759,1781]" gridcol="5" gridrow="6" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD85F01D416DBE" box="[1044,1089,1759,1781]" gridcol="6" gridrow="6" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE85F01DA56DBE" box="[1143,1189,1759,1781]" gridcol="7" gridrow="6" pageId="9" pageNumber="10">8.6</td>
<td id="A5725DD8FFBC0028FB0285F01DF76DBE" box="[1227,1271,1759,1781]" gridcol="8" gridrow="6" pageId="9" pageNumber="10">8.6</td>
<td id="A5725DD8FFBC0028FAD485F01C9C6DBE" box="[1309,1436,1759,1781]" gridcol="9" gridrow="6" pageId="9" pageNumber="10">-</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E84271C9C6C0D" box="[151,1436,1800,1862]" gridrow="7" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E8427185C6C0D" box="[151,348,1800,1862]" gridcol="0" gridrow="7" pageId="9" pageNumber="10">KGP-MH DH-007 KGP-MH DH-008</th>
<td id="A5725DD8FFBC0028FE6C84271B306C0D" box="[421,560,1800,1862]" gridcol="1" gridrow="7" pageId="9" pageNumber="10">major robust major robust</td>
<td id="A5725DD8FFBC0028FD9484271B6E6C0D" box="[605,622,1800,1862]" gridcol="2" gridrow="7" pageId="9" pageNumber="10">R R</td>
<td id="A5725DD8FFBC0028FCC584271A466C0D" box="[780,838,1800,1862]" gridcol="3" gridrow="7" pageId="9" pageNumber="10">- -</td>
<td id="A5725DD8FFBC0028FCA784271A8E6C0D" box="[878,910,1800,1862]" gridcol="4" gridrow="7" pageId="9" pageNumber="10">- -</td>
<td id="A5725DD8FFBC0028FC7B84271AED6C0D" box="[946,1005,1800,1862]" gridcol="5" gridrow="7" pageId="9" pageNumber="10">&gt;20.3&gt;19.4</td>
<td id="A5725DD8FFBC0028FBDD84271D416C0D" box="[1044,1089,1800,1862]" gridcol="6" gridrow="7" pageId="9" pageNumber="10">15.4 14.5</td>
<td id="A5725DD8FFBC0028FBBE84271DA56C0D" box="[1143,1189,1800,1862]" gridcol="7" gridrow="7" pageId="9" pageNumber="10">- -</td>
<td id="A5725DD8FFBC0028FB0284271DF76C0D" box="[1227,1271,1800,1862]" gridcol="8" gridrow="7" pageId="9" pageNumber="10">- -</td>
<td id="A5725DD8FFBC0028FAD484271C9C6C0D" box="[1309,1436,1800,1862]" gridcol="9" gridrow="7" pageId="9" pageNumber="10">- -</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E84761C9C6C24" box="[151,1436,1881,1903]" gridrow="8" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E8476185C6C24" box="[151,348,1881,1903]" gridcol="0" gridrow="8" pageId="9" pageNumber="10">KGP-MH DH-009</th>
<td id="A5725DD8FFBC0028FE6C84761B306C24" box="[421,560,1881,1903]" gridcol="1" gridrow="8" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9484761B6E6C24" box="[605,622,1881,1903]" gridcol="2" gridrow="8" pageId="9" pageNumber="10">R</td>
<td id="A5725DD8FFBC0028FCC584761A466C24" box="[780,838,1881,1903]" gridcol="3" gridrow="8" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FCA784761A8E6C24" box="[878,910,1881,1903]" gridcol="4" gridrow="8" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FC7B84761AED6C24" box="[946,1005,1881,1903]" gridcol="5" gridrow="8" pageId="9" pageNumber="10">~17</td>
<td id="A5725DD8FFBC0028FBDD84761D416C24" box="[1044,1089,1881,1903]" gridcol="6" gridrow="8" pageId="9" pageNumber="10">~14</td>
<td id="A5725DD8FFBC0028FBBE84761DA56C24" box="[1143,1189,1881,1903]" gridcol="7" gridrow="8" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FB0284761DF76C24" box="[1227,1271,1881,1903]" gridcol="8" gridrow="8" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FAD484761C9C6C24" box="[1309,1436,1881,1903]" gridcol="9" gridrow="8" pageId="9" pageNumber="10">-</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E84AD1C9C6CD3" box="[151,1436,1922,1944]" gridrow="9" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E84AD185C6CD3" box="[151,348,1922,1944]" gridcol="0" gridrow="9" pageId="9" pageNumber="10">KGP-MH DH-010</th>
<td id="A5725DD8FFBC0028FE6C84AD1B306CD3" box="[421,560,1922,1944]" gridcol="1" gridrow="9" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9484AD1B6E6CD3" box="[605,622,1922,1944]" gridcol="2" gridrow="9" pageId="9" pageNumber="10">L</td>
<td id="A5725DD8FFBC0028FCC584AD1A466CD3" box="[780,838,1922,1944]" gridcol="3" gridrow="9" pageId="9" pageNumber="10">13.2</td>
<td id="A5725DD8FFBC0028FCA784AD1A8E6CD3" box="[878,910,1922,1944]" gridcol="4" gridrow="9" pageId="9" pageNumber="10">8.5</td>
<td id="A5725DD8FFBC0028FC7B84AD1AED6CD3" box="[946,1005,1922,1944]" gridcol="5" gridrow="9" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD84AD1D416CD3" box="[1044,1089,1922,1944]" gridcol="6" gridrow="9" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE84AD1DA56CD3" box="[1143,1189,1922,1944]" gridcol="7" gridrow="9" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FB0284AD1DF76CD3" box="[1227,1271,1922,1944]" gridcol="8" gridrow="9" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FAD484AD1C9C6CD3" box="[1309,1436,1922,1944]" gridcol="9" gridrow="9" pageId="9" pageNumber="10">-</td>
</tr>
<tr id="E6A334A4FFBC0028FF5E84841C9C6C8A" box="[151,1436,1963,1985]" gridrow="10" pageId="9" pageNumber="10">
<th id="A5725DD8FFBC0028FF5E8484185C6C8A" box="[151,348,1963,1985]" gridcol="0" gridrow="10" pageId="9" pageNumber="10">KGP-MH DH-011</th>
<td id="A5725DD8FFBC0028FE6C84841B306C8A" box="[421,560,1963,1985]" gridcol="1" gridrow="10" pageId="9" pageNumber="10">major robust</td>
<td id="A5725DD8FFBC0028FD9484841B6E6C8A" box="[605,622,1963,1985]" gridcol="2" gridrow="10" pageId="9" pageNumber="10">R</td>
<td id="A5725DD8FFBC0028FCC584841A466C8A" box="[780,838,1963,1985]" gridcol="3" gridrow="10" pageId="9" pageNumber="10">&gt;15.0</td>
<td id="A5725DD8FFBC0028FCA784841A8E6C8A" box="[878,910,1963,1985]" gridcol="4" gridrow="10" pageId="9" pageNumber="10">~9</td>
<td id="A5725DD8FFBC0028FC7B84841AED6C8A" box="[946,1005,1963,1985]" gridcol="5" gridrow="10" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBDD84841D416C8A" box="[1044,1089,1963,1985]" gridcol="6" gridrow="10" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FBBE84841DA56C8A" box="[1143,1189,1963,1985]" gridcol="7" gridrow="10" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FB0284841DF76C8A" box="[1227,1271,1963,1985]" gridcol="8" gridrow="10" pageId="9" pageNumber="10">-</td>
<td id="A5725DD8FFBC0028FAD484841C9C6C8A" box="[1309,1436,1963,1985]" gridcol="9" gridrow="10" pageId="9" pageNumber="10">-</td>
</tr>
</table>
</paragraph>
<paragraph id="582C36E6FFBFFFDDFF5683B01C266FFA" pageId="10" pageNumber="11">
<table id="2A93C446FFBF0028FF5683B01C716FFA" box="[159,1393,159,1201]" gridcols="9" gridrows="26" pageId="10" pageNumber="11">
<tr id="E6A334A4FFBF0028FF5683B01C716BFE" box="[159,1393,159,181]" gridrow="0" pageId="10" pageNumber="11" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="A5725DD8FFBF0028FF5683B018876BFE" box="[159,391,159,181]" gridcol="0" gridrow="0" pageId="10" pageNumber="11">
<emphasis id="6AE7EAF4FFBFFFDDFF5683B0180C6BFF" bold="true" box="[159,268,159,180]" pageId="10" pageNumber="11">TABLE 1.</emphasis>
(continued)
</th>
</tr>
<tr id="E6A334A4FFBF0028FF5683E71C716B95" box="[159,1393,200,222]" gridrow="1" pageId="10" pageNumber="11" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-6="1">
<td id="A5725DD8FFBF0028FCC583E71A4A6B95" box="[780,842,200,222]" gridcol="3" gridrow="1" pageId="10" pageNumber="11">merus</td>
<td id="A5725DD8FFBF0028FC7B83E71AF46B95" box="[946,1012,200,222]" gridcol="5" gridrow="1" pageId="10" pageNumber="11">carpus</td>
<td id="A5725DD8FFBF0028FBBE83E71DF76B95" box="[1143,1271,200,222]" gridcol="7" gridrow="1" pageId="10" pageNumber="11">propodus</td>
<td id="A5725DD8FFBF0028FAD483E71C716B95" box="[1309,1393,200,222]" gridcol="8" gridrow="1" pageId="10" pageNumber="11">dactylus</td>
</tr>
<tr id="E6A334A4FFBF0028FF5683DE1C716A4C" box="[159,1393,241,263]" gridrow="2" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF5683DE18876A4C" box="[159,391,241,263]" gridcol="0" gridrow="2" pageId="10" pageNumber="11">Specimen</th>
<td id="A5725DD8FFBF0028FE6C83DE1B306A4C" box="[421,560,241,263]" gridcol="1" gridrow="2" pageId="10" pageNumber="11">morphotype</td>
<td id="A5725DD8FFBF0028FD9483DE1BD26A4C" box="[605,722,241,263]" gridcol="2" gridrow="2" pageId="10" pageNumber="11">handedness</td>
<td id="A5725DD8FFBF0028FCC583DE1A4A6A4C" box="[780,842,241,263]" gridcol="3" gridrow="2" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCA783DE1A8E6A4C" box="[878,910,241,263]" gridcol="4" gridrow="2" pageId="10" pageNumber="11">H</td>
<td id="A5725DD8FFBF0028FC7B83DE1AF46A4C" box="[946,1012,241,263]" gridcol="5" gridrow="2" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FBDD83DE1D4F6A4C" box="[1044,1103,241,263]" gridcol="6" gridrow="2" pageId="10" pageNumber="11">H</td>
<td id="A5725DD8FFBF0028FBBE83DE1DF76A4C" box="[1143,1271,241,263]" gridcol="7" gridrow="2" pageId="10" pageNumber="11">L H</td>
<td id="A5725DD8FFBF0028FAD483DE1C716A4C" box="[1309,1393,241,263]" gridcol="8" gridrow="2" pageId="10" pageNumber="11">L</td>
</tr>
<tr id="E6A334A4FFBF0028FF5682351C716A7B" box="[159,1393,282,304]" gridrow="3" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56823518876A7B" box="[159,391,282,304]" gridcol="0" gridrow="3" pageId="10" pageNumber="11">KGP-MH DH-012</th>
<td id="A5725DD8FFBF0028FE6C82351B306A7B" box="[421,560,282,304]" gridcol="1" gridrow="3" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD9482351BD26A7B" box="[605,722,282,304]" gridcol="2" gridrow="3" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC582351A4A6A7B" box="[780,842,282,304]" gridcol="3" gridrow="3" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA782351A8E6A7B" box="[878,910,282,304]" gridcol="4" gridrow="3" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B82351AF46A7B" box="[946,1012,282,304]" gridcol="5" gridrow="3" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD82351D4F6A7B" box="[1044,1103,282,304]" gridcol="6" gridrow="3" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE82351DF76A7B" box="[1143,1271,282,304]" gridcol="7" gridrow="3" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD482351C716A7B" box="[1309,1393,282,304]" gridcol="8" gridrow="3" pageId="10" pageNumber="11">12.0</td>
</tr>
<tr id="E6A334A4FFBF0028FF56826D1C716A13" box="[159,1393,322,344]" gridrow="4" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56826D18876A13" box="[159,391,322,344]" gridcol="0" gridrow="4" pageId="10" pageNumber="11">KGP-MH DH-058</th>
<td id="A5725DD8FFBF0028FE6C826D1B306A13" box="[421,560,322,344]" gridcol="1" gridrow="4" pageId="10" pageNumber="11">minor</td>
<td id="A5725DD8FFBF0028FD94826D1BD26A13" box="[605,722,322,344]" gridcol="2" gridrow="4" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC5826D1A4A6A13" box="[780,842,322,344]" gridcol="3" gridrow="4" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA7826D1A8E6A13" box="[878,910,322,344]" gridcol="4" gridrow="4" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B826D1AF46A13" box="[946,1012,322,344]" gridcol="5" gridrow="4" pageId="10" pageNumber="11">7.7</td>
<td id="A5725DD8FFBF0028FBDD826D1D4F6A13" box="[1044,1103,322,344]" gridcol="6" gridrow="4" pageId="10" pageNumber="11">6.5</td>
<td id="A5725DD8FFBF0028FBBE826D1DF76A13" box="[1143,1271,322,344]" gridcol="7" gridrow="4" pageId="10" pageNumber="11">5.4 5.4</td>
<td id="A5725DD8FFBF0028FAD4826D1C716A13" box="[1309,1393,322,344]" gridcol="8" gridrow="4" pageId="10" pageNumber="11">&gt;5.0</td>
</tr>
<tr id="E6A334A4FFBF0028FF5682441C716ACA" box="[159,1393,363,385]" gridrow="5" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56824418876ACA" box="[159,391,363,385]" gridcol="0" gridrow="5" pageId="10" pageNumber="11">KGP-MH DH-058</th>
<td id="A5725DD8FFBF0028FE6C82441B306ACA" box="[421,560,363,385]" gridcol="1" gridrow="5" pageId="10" pageNumber="11">major slender</td>
<td id="A5725DD8FFBF0028FD9482441BD26ACA" box="[605,722,363,385]" gridcol="2" gridrow="5" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC582441A4A6ACA" box="[780,842,363,385]" gridcol="3" gridrow="5" pageId="10" pageNumber="11">8.7</td>
<td id="A5725DD8FFBF0028FCA782441A8E6ACA" box="[878,910,363,385]" gridcol="4" gridrow="5" pageId="10" pageNumber="11">5.7</td>
<td id="A5725DD8FFBF0028FC7B82441AF46ACA" box="[946,1012,363,385]" gridcol="5" gridrow="5" pageId="10" pageNumber="11">7.6</td>
<td id="A5725DD8FFBF0028FBDD82441D4F6ACA" box="[1044,1103,363,385]" gridcol="6" gridrow="5" pageId="10" pageNumber="11">9.6</td>
<td id="A5725DD8FFBF0028FBBE82441DF76ACA" box="[1143,1271,363,385]" gridcol="7" gridrow="5" pageId="10" pageNumber="11">11.4 10.5</td>
<td id="A5725DD8FFBF0028FAD482441C716ACA" box="[1309,1393,363,385]" gridcol="8" gridrow="5" pageId="10" pageNumber="11">7.8</td>
</tr>
<tr id="E6A334A4FFBF0028FF5682BB1C716AE1" box="[159,1393,404,426]" gridrow="6" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF5682BB18876AE1" box="[159,391,404,426]" gridcol="0" gridrow="6" pageId="10" pageNumber="11">KGP-MH DH-064</th>
<td id="A5725DD8FFBF0028FE6C82BB1B306AE1" box="[421,560,404,426]" gridcol="1" gridrow="6" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD9482BB1BD26AE1" box="[605,722,404,426]" gridcol="2" gridrow="6" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC582BB1A4A6AE1" box="[780,842,404,426]" gridcol="3" gridrow="6" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA782BB1A8E6AE1" box="[878,910,404,426]" gridcol="4" gridrow="6" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B82BB1AF46AE1" box="[946,1012,404,426]" gridcol="5" gridrow="6" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD82BB1D4F6AE1" box="[1044,1103,404,426]" gridcol="6" gridrow="6" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE82BB1DF76AE1" box="[1143,1271,404,426]" gridcol="7" gridrow="6" pageId="10" pageNumber="11">18.0 16.6</td>
<td id="A5725DD8FFBF0028FAD482BB1C716AE1" box="[1309,1393,404,426]" gridcol="8" gridrow="6" pageId="10" pageNumber="11">-</td>
</tr>
<tr id="E6A334A4FFBF0028FF5682921C716A98" box="[159,1393,445,467]" gridrow="7" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56829218876A98" box="[159,391,445,467]" gridcol="0" gridrow="7" pageId="10" pageNumber="11">KGP-MH DH-065</th>
<td id="A5725DD8FFBF0028FE6C82921B306A98" box="[421,560,445,467]" gridcol="1" gridrow="7" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD9482921BD26A98" box="[605,722,445,467]" gridcol="2" gridrow="7" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC582921A4A6A98" box="[780,842,445,467]" gridcol="3" gridrow="7" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA782921A8E6A98" box="[878,910,445,467]" gridcol="4" gridrow="7" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B82921AF46A98" box="[946,1012,445,467]" gridcol="5" gridrow="7" pageId="10" pageNumber="11">17.0</td>
<td id="A5725DD8FFBF0028FBDD82921D4F6A98" box="[1044,1103,445,467]" gridcol="6" gridrow="7" pageId="10" pageNumber="11">&gt;14.3</td>
<td id="A5725DD8FFBF0028FBBE82921DF76A98" box="[1143,1271,445,467]" gridcol="7" gridrow="7" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD482921C716A98" box="[1309,1393,445,467]" gridcol="8" gridrow="7" pageId="10" pageNumber="11">-</td>
</tr>
<tr id="E6A334A4FFBF0028FF5682CA1C716AB0" box="[159,1393,485,507]" gridrow="8" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF5682CA18876AB0" box="[159,391,485,507]" gridcol="0" gridrow="8" pageId="10" pageNumber="11">KGP-MH DH-066</th>
<td id="A5725DD8FFBF0028FE6C82CA1B306AB0" box="[421,560,485,507]" gridcol="1" gridrow="8" pageId="10" pageNumber="11">unknown</td>
<td id="A5725DD8FFBF0028FD9482CA1BD26AB0" box="[605,722,485,507]" gridcol="2" gridrow="8" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC582CA1A4A6AB0" box="[780,842,485,507]" gridcol="3" gridrow="8" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA782CA1A8E6AB0" box="[878,910,485,507]" gridcol="4" gridrow="8" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B82CA1AF46AB0" box="[946,1012,485,507]" gridcol="5" gridrow="8" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD82CA1D4F6AB0" box="[1044,1103,485,507]" gridcol="6" gridrow="8" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE82CA1DF76AB0" box="[1143,1271,485,507]" gridcol="7" gridrow="8" pageId="10" pageNumber="11">8.7 14.7</td>
<td id="A5725DD8FFBF0028FAD482CA1C716AB0" box="[1309,1393,485,507]" gridcol="8" gridrow="8" pageId="10" pageNumber="11">-</td>
</tr>
<tr id="E6A334A4FFBF0028FF5681211C71696F" box="[159,1393,526,548]" gridrow="9" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF5681211887696F" box="[159,391,526,548]" gridcol="0" gridrow="9" pageId="10" pageNumber="11">KGP-MH DH-067</th>
<td id="A5725DD8FFBF0028FE6C81211B30696F" box="[421,560,526,548]" gridcol="1" gridrow="9" pageId="10" pageNumber="11">minor</td>
<td id="A5725DD8FFBF0028FD9481211BD2696F" box="[605,722,526,548]" gridcol="2" gridrow="9" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC581211A4A696F" box="[780,842,526,548]" gridcol="3" gridrow="9" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA781211A8E696F" box="[878,910,526,548]" gridcol="4" gridrow="9" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B81211AF4696F" box="[946,1012,526,548]" gridcol="5" gridrow="9" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD81211D4F696F" box="[1044,1103,526,548]" gridcol="6" gridrow="9" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE81211DF7696F" box="[1143,1271,526,548]" gridcol="7" gridrow="9" pageId="10" pageNumber="11">&gt;6.0 6.7</td>
<td id="A5725DD8FFBF0028FAD481211C71696F" box="[1309,1393,526,548]" gridcol="8" gridrow="9" pageId="10" pageNumber="11">&gt;5.5</td>
</tr>
<tr id="E6A334A4FFBF0028FF5681181C716906" box="[159,1393,567,589]" gridrow="10" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56811818876906" box="[159,391,567,589]" gridcol="0" gridrow="10" pageId="10" pageNumber="11">PCMH-001</th>
<td id="A5725DD8FFBF0028FE6C81181B306906" box="[421,560,567,589]" gridcol="1" gridrow="10" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD9481181BD26906" box="[605,722,567,589]" gridcol="2" gridrow="10" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC581181A4A6906" box="[780,842,567,589]" gridcol="3" gridrow="10" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA781181A8E6906" box="[878,910,567,589]" gridcol="4" gridrow="10" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B81181AF46906" box="[946,1012,567,589]" gridcol="5" gridrow="10" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD81181D4F6906" box="[1044,1103,567,589]" gridcol="6" gridrow="10" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE81181DF76906" box="[1143,1271,567,589]" gridcol="7" gridrow="10" pageId="10" pageNumber="11">12.0 11.8</td>
<td id="A5725DD8FFBF0028FAD481181C716906" box="[1309,1393,567,589]" gridcol="8" gridrow="10" pageId="10" pageNumber="11">9.2</td>
</tr>
<tr id="E6A334A4FFBF0028FF56814F1C71693D" box="[159,1393,608,630]" gridrow="11" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56814F1887693D" box="[159,391,608,630]" gridcol="0" gridrow="11" pageId="10" pageNumber="11">PCRB DH-001</th>
<td id="A5725DD8FFBF0028FE6C814F1B30693D" box="[421,560,608,630]" gridcol="1" gridrow="11" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD94814F1BD2693D" box="[605,722,608,630]" gridcol="2" gridrow="11" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC5814F1A4A693D" box="[780,842,608,630]" gridcol="3" gridrow="11" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA7814F1A8E693D" box="[878,910,608,630]" gridcol="4" gridrow="11" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B814F1AF4693D" box="[946,1012,608,630]" gridcol="5" gridrow="11" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD814F1D4F693D" box="[1044,1103,608,630]" gridcol="6" gridrow="11" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE814F1DF7693D" box="[1143,1271,608,630]" gridcol="7" gridrow="11" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD4814F1C71693D" box="[1309,1393,608,630]" gridcol="8" gridrow="11" pageId="10" pageNumber="11">12.2</td>
</tr>
<tr id="E6A334A4FFBF0028FF5681A61C7169D4" box="[159,1393,649,671]" gridrow="12" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF5681A6188769D4" box="[159,391,649,671]" gridcol="0" gridrow="12" pageId="10" pageNumber="11">PCRB DH-002</th>
<td id="A5725DD8FFBF0028FE6C81A61B3069D4" box="[421,560,649,671]" gridcol="1" gridrow="12" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD9481A61BD269D4" box="[605,722,649,671]" gridcol="2" gridrow="12" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC581A61A4A69D4" box="[780,842,649,671]" gridcol="3" gridrow="12" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA781A61A8E69D4" box="[878,910,649,671]" gridcol="4" gridrow="12" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B81A61AF469D4" box="[946,1012,649,671]" gridcol="5" gridrow="12" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD81A61D4F69D4" box="[1044,1103,649,671]" gridcol="6" gridrow="12" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE81A61DF769D4" box="[1143,1271,649,671]" gridcol="7" gridrow="12" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD481A61C7169D4" box="[1309,1393,649,671]" gridcol="8" gridrow="12" pageId="10" pageNumber="11">11.1</td>
</tr>
<tr id="E6A334A4FFBF0028FF56819D1C716983" box="[159,1393,690,712]" gridrow="13" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56819D18876983" box="[159,391,690,712]" gridcol="0" gridrow="13" pageId="10" pageNumber="11">PCRB DH-003</th>
<td id="A5725DD8FFBF0028FE6C819D1B306983" box="[421,560,690,712]" gridcol="1" gridrow="13" pageId="10" pageNumber="11">unknown</td>
<td id="A5725DD8FFBF0028FD94819D1BD26983" box="[605,722,690,712]" gridcol="2" gridrow="13" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC5819D1A4A6983" box="[780,842,690,712]" gridcol="3" gridrow="13" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA7819D1A8E6983" box="[878,910,690,712]" gridcol="4" gridrow="13" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B819D1AF46983" box="[946,1012,690,712]" gridcol="5" gridrow="13" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD819D1D4F6983" box="[1044,1103,690,712]" gridcol="6" gridrow="13" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE819D1DF76983" box="[1143,1271,690,712]" gridcol="7" gridrow="13" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD4819D1C716983" box="[1309,1393,690,712]" gridcol="8" gridrow="13" pageId="10" pageNumber="11">7.0</td>
</tr>
<tr id="E6A334A4FFBF0028FF5681F51C7169BB" box="[159,1393,730,752]" gridrow="14" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF5681F5188769BB" box="[159,391,730,752]" gridcol="0" gridrow="14" pageId="10" pageNumber="11">PCRB DH-004</th>
<td id="A5725DD8FFBF0028FE6C81F51B3069BB" box="[421,560,730,752]" gridcol="1" gridrow="14" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD9481F51BD269BB" box="[605,722,730,752]" gridcol="2" gridrow="14" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC581F51A4A69BB" box="[780,842,730,752]" gridcol="3" gridrow="14" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA781F51A8E69BB" box="[878,910,730,752]" gridcol="4" gridrow="14" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B81F51AF469BB" box="[946,1012,730,752]" gridcol="5" gridrow="14" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD81F51D4F69BB" box="[1044,1103,730,752]" gridcol="6" gridrow="14" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE81F51DF769BB" box="[1143,1271,730,752]" gridcol="7" gridrow="14" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD481F51C7169BB" box="[1309,1393,730,752]" gridcol="8" gridrow="14" pageId="10" pageNumber="11">&gt;11.0</td>
</tr>
<tr id="E6A334A4FFBF0028FF56802C1C716852" box="[159,1393,771,793]" gridrow="15" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56802C18876852" box="[159,391,771,793]" gridcol="0" gridrow="15" pageId="10" pageNumber="11">PCRB DH-005</th>
<td id="A5725DD8FFBF0028FE6C802C1B306852" box="[421,560,771,793]" gridcol="1" gridrow="15" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD94802C1BD26852" box="[605,722,771,793]" gridcol="2" gridrow="15" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC5802C1A4A6852" box="[780,842,771,793]" gridcol="3" gridrow="15" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA7802C1A8E6852" box="[878,910,771,793]" gridcol="4" gridrow="15" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B802C1AF46852" box="[946,1012,771,793]" gridcol="5" gridrow="15" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD802C1D4F6852" box="[1044,1103,771,793]" gridcol="6" gridrow="15" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE802C1DF76852" box="[1143,1271,771,793]" gridcol="7" gridrow="15" pageId="10" pageNumber="11">- 14.0</td>
<td id="A5725DD8FFBF0028FAD4802C1C716852" box="[1309,1393,771,793]" gridcol="8" gridrow="15" pageId="10" pageNumber="11">11.2</td>
</tr>
<tr id="E6A334A4FFBF0028FF5680031C716809" box="[159,1393,812,834]" gridrow="16" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56800318876809" box="[159,391,812,834]" gridcol="0" gridrow="16" pageId="10" pageNumber="11">PCRB DH-007</th>
<td id="A5725DD8FFBF0028FE6C80031B306809" box="[421,560,812,834]" gridcol="1" gridrow="16" pageId="10" pageNumber="11">major slender</td>
<td id="A5725DD8FFBF0028FD9480031BD26809" box="[605,722,812,834]" gridcol="2" gridrow="16" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC580031A4A6809" box="[780,842,812,834]" gridcol="3" gridrow="16" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA780031A8E6809" box="[878,910,812,834]" gridcol="4" gridrow="16" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B80031AF46809" box="[946,1012,812,834]" gridcol="5" gridrow="16" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD80031D4F6809" box="[1044,1103,812,834]" gridcol="6" gridrow="16" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE80031DF76809" box="[1143,1271,812,834]" gridcol="7" gridrow="16" pageId="10" pageNumber="11">9.0 8.4</td>
<td id="A5725DD8FFBF0028FAD480031C716809" box="[1309,1393,812,834]" gridcol="8" gridrow="16" pageId="10" pageNumber="11">-</td>
</tr>
<tr id="E6A334A4FFBF0028FF56807A1C716820" box="[159,1393,853,875]" gridrow="17" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56807A18876820" box="[159,391,853,875]" gridcol="0" gridrow="17" pageId="10" pageNumber="11">PCRB DH-008</th>
<td id="A5725DD8FFBF0028FE6C807A1B306820" box="[421,560,853,875]" gridcol="1" gridrow="17" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD94807A1BD26820" box="[605,722,853,875]" gridcol="2" gridrow="17" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC5807A1A4A6820" box="[780,842,853,875]" gridcol="3" gridrow="17" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA7807A1A8E6820" box="[878,910,853,875]" gridcol="4" gridrow="17" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B807A1AF46820" box="[946,1012,853,875]" gridcol="5" gridrow="17" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD807A1D4F6820" box="[1044,1103,853,875]" gridcol="6" gridrow="17" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE807A1DF76820" box="[1143,1271,853,875]" gridcol="7" gridrow="17" pageId="10" pageNumber="11">16.1 14.8</td>
<td id="A5725DD8FFBF0028FAD4807A1C716820" box="[1309,1393,853,875]" gridcol="8" gridrow="17" pageId="10" pageNumber="11">&gt;10.0</td>
</tr>
<tr id="E6A334A4FFBF0028FF5680521C7168D8" box="[159,1393,893,915]" gridrow="18" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF568052188768D8" box="[159,391,893,915]" gridcol="0" gridrow="18" pageId="10" pageNumber="11">PCRB DH-009</th>
<td id="A5725DD8FFBF0028FE6C80521B3068D8" box="[421,560,893,915]" gridcol="1" gridrow="18" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD9480521BD268D8" box="[605,722,893,915]" gridcol="2" gridrow="18" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC580521A4A68D8" box="[780,842,893,915]" gridcol="3" gridrow="18" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA780521A8E68D8" box="[878,910,893,915]" gridcol="4" gridrow="18" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B80521AF468D8" box="[946,1012,893,915]" gridcol="5" gridrow="18" pageId="10" pageNumber="11">&gt;12.0</td>
<td id="A5725DD8FFBF0028FBDD80521D4F68D8" box="[1044,1103,893,915]" gridcol="6" gridrow="18" pageId="10" pageNumber="11">14.4</td>
<td id="A5725DD8FFBF0028FBBE80521DF768D8" box="[1143,1271,893,915]" gridcol="7" gridrow="18" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD480521C7168D8" box="[1309,1393,893,915]" gridcol="8" gridrow="18" pageId="10" pageNumber="11">-</td>
</tr>
<tr id="E6A334A4FFBF0028FF5680891C7168F7" box="[159,1393,934,956]" gridrow="19" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF568089188768F7" box="[159,391,934,956]" gridcol="0" gridrow="19" pageId="10" pageNumber="11">PCRB DH-010</th>
<td id="A5725DD8FFBF0028FE6C80891B3068F7" box="[421,560,934,956]" gridcol="1" gridrow="19" pageId="10" pageNumber="11">major slender</td>
<td id="A5725DD8FFBF0028FD9480891BD268F7" box="[605,722,934,956]" gridcol="2" gridrow="19" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC580891A4A68F7" box="[780,842,934,956]" gridcol="3" gridrow="19" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA780891A8E68F7" box="[878,910,934,956]" gridcol="4" gridrow="19" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B80891AF468F7" box="[946,1012,934,956]" gridcol="5" gridrow="19" pageId="10" pageNumber="11">6.2</td>
<td id="A5725DD8FFBF0028FBDD80891D4F68F7" box="[1044,1103,934,956]" gridcol="6" gridrow="19" pageId="10" pageNumber="11">7.3</td>
<td id="A5725DD8FFBF0028FBBE80891DF768F7" box="[1143,1271,934,956]" gridcol="7" gridrow="19" pageId="10" pageNumber="11">8.1 7.6</td>
<td id="A5725DD8FFBF0028FAD480891C7168F7" box="[1309,1393,934,956]" gridcol="8" gridrow="19" pageId="10" pageNumber="11">-</td>
</tr>
<tr id="E6A334A4FFBF0028FF5680E01C7168AE" box="[159,1393,975,997]" gridrow="20" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF5680E0188768AE" box="[159,391,975,997]" gridcol="0" gridrow="20" pageId="10" pageNumber="11">PCRB DH-011</th>
<td id="A5725DD8FFBF0028FE6C80E01B3068AE" box="[421,560,975,997]" gridcol="1" gridrow="20" pageId="10" pageNumber="11">minor</td>
<td id="A5725DD8FFBF0028FD9480E01BD268AE" box="[605,722,975,997]" gridcol="2" gridrow="20" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC580E01A4A68AE" box="[780,842,975,997]" gridcol="3" gridrow="20" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA780E01A8E68AE" box="[878,910,975,997]" gridcol="4" gridrow="20" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B80E01AF468AE" box="[946,1012,975,997]" gridcol="5" gridrow="20" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD80E01D4F68AE" box="[1044,1103,975,997]" gridcol="6" gridrow="20" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE80E01DF768AE" box="[1143,1271,975,997]" gridcol="7" gridrow="20" pageId="10" pageNumber="11">7.0 7.6</td>
<td id="A5725DD8FFBF0028FAD480E01C7168AE" box="[1309,1393,975,997]" gridcol="8" gridrow="20" pageId="10" pageNumber="11">-</td>
</tr>
<tr id="E6A334A4FFBF0028FF5680D71C716F45" box="[159,1393,1016,1038]" gridrow="21" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF5680D718876F45" box="[159,391,1016,1038]" gridcol="0" gridrow="21" pageId="10" pageNumber="11">PCRB DH-012</th>
<td id="A5725DD8FFBF0028FE6C80D71B306F45" box="[421,560,1016,1038]" gridcol="1" gridrow="21" pageId="10" pageNumber="11">minor</td>
<td id="A5725DD8FFBF0028FD9480D71BD26F45" box="[605,722,1016,1038]" gridcol="2" gridrow="21" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC580D71A4A6F45" box="[780,842,1016,1038]" gridcol="3" gridrow="21" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA780D71A8E6F45" box="[878,910,1016,1038]" gridcol="4" gridrow="21" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B80D71AF46F45" box="[946,1012,1016,1038]" gridcol="5" gridrow="21" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD80D71D4F6F45" box="[1044,1103,1016,1038]" gridcol="6" gridrow="21" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE80D71DF76F45" box="[1143,1271,1016,1038]" gridcol="7" gridrow="21" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD480D71C716F45" box="[1309,1393,1016,1038]" gridcol="8" gridrow="21" pageId="10" pageNumber="11">11.3</td>
</tr>
<tr id="E6A334A4FFBF0028FF56870E1C716F7C" box="[159,1393,1057,1079]" gridrow="22" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56870E18876F7C" box="[159,391,1057,1079]" gridcol="0" gridrow="22" pageId="10" pageNumber="11">PCRB DH-013</th>
<td id="A5725DD8FFBF0028FE6C870E1B306F7C" box="[421,560,1057,1079]" gridcol="1" gridrow="22" pageId="10" pageNumber="11">minor</td>
<td id="A5725DD8FFBF0028FD94870E1BD26F7C" box="[605,722,1057,1079]" gridcol="2" gridrow="22" pageId="10" pageNumber="11">R</td>
<td id="A5725DD8FFBF0028FCC5870E1A4A6F7C" box="[780,842,1057,1079]" gridcol="3" gridrow="22" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FCA7870E1A8E6F7C" box="[878,910,1057,1079]" gridcol="4" gridrow="22" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FC7B870E1AF46F7C" box="[946,1012,1057,1079]" gridcol="5" gridrow="22" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD870E1D4F6F7C" box="[1044,1103,1057,1079]" gridcol="6" gridrow="22" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE870E1DF76F7C" box="[1143,1271,1057,1079]" gridcol="7" gridrow="22" pageId="10" pageNumber="11">8.0 8.0</td>
<td id="A5725DD8FFBF0028FAD4870E1C716F7C" box="[1309,1393,1057,1079]" gridcol="8" gridrow="22" pageId="10" pageNumber="11">-</td>
</tr>
<tr id="E6A334A4FFBF0028FF5687651C716F2B" box="[159,1393,1098,1120]" gridrow="23" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56876518876F2B" box="[159,391,1098,1120]" gridcol="0" gridrow="23" pageId="10" pageNumber="11">PCRB DH-014</th>
<td id="A5725DD8FFBF0028FE6C87651B306F2B" box="[421,560,1098,1120]" gridcol="1" gridrow="23" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD9487651BD26F2B" box="[605,722,1098,1120]" gridcol="2" gridrow="23" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC587651A4A6F2B" box="[780,842,1098,1120]" gridcol="3" gridrow="23" pageId="10" pageNumber="11">11.7</td>
<td id="A5725DD8FFBF0028FCA787651A8E6F2B" box="[878,910,1098,1120]" gridcol="4" gridrow="23" pageId="10" pageNumber="11">7.3</td>
<td id="A5725DD8FFBF0028FC7B87651AF46F2B" box="[946,1012,1098,1120]" gridcol="5" gridrow="23" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD87651D4F6F2B" box="[1044,1103,1098,1120]" gridcol="6" gridrow="23" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE87651DF76F2B" box="[1143,1271,1098,1120]" gridcol="7" gridrow="23" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD487651C716F2B" box="[1309,1393,1098,1120]" gridcol="8" gridrow="23" pageId="10" pageNumber="11">-</td>
</tr>
<tr id="E6A334A4FFBF0028FF56875D1C716FC3" box="[159,1393,1138,1160]" gridrow="24" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF56875D18876FC3" box="[159,391,1138,1160]" gridcol="0" gridrow="24" pageId="10" pageNumber="11">PCRB DH-015</th>
<td id="A5725DD8FFBF0028FE6C875D1B306FC3" box="[421,560,1138,1160]" gridcol="1" gridrow="24" pageId="10" pageNumber="11">major robust</td>
<td id="A5725DD8FFBF0028FD94875D1BD26FC3" box="[605,722,1138,1160]" gridcol="2" gridrow="24" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC5875D1A4A6FC3" box="[780,842,1138,1160]" gridcol="3" gridrow="24" pageId="10" pageNumber="11">&gt;11.0</td>
<td id="A5725DD8FFBF0028FCA7875D1A8E6FC3" box="[878,910,1138,1160]" gridcol="4" gridrow="24" pageId="10" pageNumber="11">8.0</td>
<td id="A5725DD8FFBF0028FC7B875D1AF46FC3" box="[946,1012,1138,1160]" gridcol="5" gridrow="24" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD875D1D4F6FC3" box="[1044,1103,1138,1160]" gridcol="6" gridrow="24" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE875D1DF76FC3" box="[1143,1271,1138,1160]" gridcol="7" gridrow="24" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD4875D1C716FC3" box="[1309,1393,1138,1160]" gridcol="8" gridrow="24" pageId="10" pageNumber="11">-</td>
</tr>
<tr id="E6A334A4FFBF0028FF5687B41C716FFA" box="[159,1393,1179,1201]" gridrow="25" pageId="10" pageNumber="11">
<th id="A5725DD8FFBF0028FF5687B418876FFA" box="[159,391,1179,1201]" gridcol="0" gridrow="25" pageId="10" pageNumber="11">PCRB DH-016</th>
<td id="A5725DD8FFBF0028FE6C87B41B306FFA" box="[421,560,1179,1201]" gridcol="1" gridrow="25" pageId="10" pageNumber="11">major slender</td>
<td id="A5725DD8FFBF0028FD9487B41BD26FFA" box="[605,722,1179,1201]" gridcol="2" gridrow="25" pageId="10" pageNumber="11">L</td>
<td id="A5725DD8FFBF0028FCC587B41A4A6FFA" box="[780,842,1179,1201]" gridcol="3" gridrow="25" pageId="10" pageNumber="11">11.8</td>
<td id="A5725DD8FFBF0028FCA787B41A8E6FFA" box="[878,910,1179,1201]" gridcol="4" gridrow="25" pageId="10" pageNumber="11">5.5</td>
<td id="A5725DD8FFBF0028FC7B87B41AF46FFA" box="[946,1012,1179,1201]" gridcol="5" gridrow="25" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBDD87B41D4F6FFA" box="[1044,1103,1179,1201]" gridcol="6" gridrow="25" pageId="10" pageNumber="11">-</td>
<td id="A5725DD8FFBF0028FBBE87B41DF76FFA" box="[1143,1271,1179,1201]" gridcol="7" gridrow="25" pageId="10" pageNumber="11">- -</td>
<td id="A5725DD8FFBF0028FAD487B41C716FFA" box="[1309,1393,1179,1201]" gridcol="8" gridrow="25" pageId="10" pageNumber="11">-</td>
</tr>
</table>
</paragraph>
<paragraph id="582C36E6FFBFFFDDFF5E86AF1B2B6ED3" blockId="10.[151,1437,1264,2009]" box="[151,555,1408,1433]" pageId="10" pageNumber="11">
(
<bibRefCitation id="3C024B17FFBFFFDDFF5786AF18376ED2" author="Hyzny" box="[158,311,1408,1433]" pageId="10" pageNumber="11" refString="Hyzny, M. (2011 a) In situ mud shrimps (Decapoda: Axiidea: Callianassidae) preserved within their burrows from the middle Miocene of the Central Paratethys. Bulletin of the Mizunami Fossil Museum, 37, 37 - 46." type="journal article" year="2011" yearSuffix="a">
Hyžný 2011
<emphasis id="6AE7EAF4FFBFFFDDFEE086AD18376ED2" box="[297,311,1410,1433]" italics="true" pageId="10" pageNumber="11">a</emphasis>
</bibRefCitation>
,
<bibRefCitation id="3C024B17FFBFFFDDFE8C86AE18536ED2" author="Hyzny" box="[325,339,1409,1433]" pageId="10" pageNumber="11" refString="Hyzny M. (2011 b) Synopsis of fossil decapod crustaceans from Slovakia (Western Carpathians). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 260 (2), 165 - 171." type="journal article" year="2011" yearSuffix="b">
<emphasis id="6AE7EAF4FFBFFFDDFE8C86AE18536ED2" box="[325,339,1409,1433]" italics="true" pageId="10" pageNumber="11">b</emphasis>
</bibRefCitation>
; this contribution).
</paragraph>
</subSubSection>
<subSubSection id="1089656DFFBFFFDDFF0E868A18DC6C03" pageId="10" pageNumber="11" type="discussion">
<paragraph id="582C36E6FFBFFFDDFF0E868A184E6D63" blockId="10.[151,1437,1264,2009]" pageId="10" pageNumber="11">
<emphasis id="6AE7EAF4FFBFFFDDFF0E868A183B6EF6" bold="true" box="[199,315,1445,1469]" pageId="10" pageNumber="11">Remarks.</emphasis>
The species possesses a combination of characters which is quite typical for
<taxonomicName id="9F934D65FFBFFFDDFB75868A1C616EF6" box="[1212,1377,1445,1469]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFBFFFDDFB75868A1C616EF6" box="[1212,1377,1445,1469]" italics="true" pageId="10" pageNumber="11">Neocallichirus</emphasis>
</taxonomicName>
: serrated lower margins of the merus, carpus and propodus; rounded lower margin of the carpus; upper margin of the propodus converging distally; an edentulous fixed finger (which is only finely denticulated), and an armed dactylus with hooked tip.
</paragraph>
<paragraph id="582C36E6FFBFFFDDFF0E851A186F6DF3" blockId="10.[151,1437,1264,2009]" pageId="10" pageNumber="11">
In his major work,
<bibRefCitation id="3C024B17FFBFFFDDFE57851B1B686D07" author="Muller" box="[414,616,1588,1612]" pageId="10" pageNumber="11" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984: 51)</bibRefCitation>
already noted the morphological similarity of
<taxonomicName id="9F934D65FFBFFFDDFBBD851A1C5D6D06" box="[1140,1373,1589,1613]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBFFFDDFBBD851A1C5D6D06" box="[1140,1373,1589,1613]" italics="true" pageId="10" pageNumber="11">Callianassa brocchii</emphasis>
</taxonomicName>
to
<taxonomicName id="9F934D65FFBFFFDDFA4B851A18B96D3B" authority="Barnard, 1947" authorityName="Barnard" authorityYear="1947" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="6AE7EAF4FFBFFFDDFA4B851A18096D3A" italics="true" pageId="10" pageNumber="11">C. natalensis</emphasis>
<bibRefCitation id="3C024B17FFBFFFDDFEDB857718B96D3B" author="Barnard" box="[274,441,1624,1648]" pageId="10" pageNumber="11" refString="Barnard, K. H. (1947) Descriptions of new species of South African decapod Crustacea, with notes on synonymy and new records. Annals and Magazine of Natural History, series 11, 13, 361 - 392. [Dated 1946, published 1947]" type="journal article" year="1947">Barnard, 1947</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="3C024B17FFBFFFDDFE0F85771B666D3B" author="Muller" box="[454,614,1624,1648]" pageId="10" pageNumber="11" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984)</bibRefCitation>
examined only the
<typeStatus id="87288844FFBFFFDDFC8785771AB06D3B" box="[846,944,1624,1648]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
of
<taxonomicName id="9F934D65FFBFFFDDFC1385761D5A6D3A" box="[986,1114,1625,1649]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBFFFDDFC1385761D5A6D3A" box="[986,1114,1625,1649]" italics="true" pageId="10" pageNumber="11">C. brocchii</emphasis>
</taxonomicName>
and five additional propodi from the Budapest City localities (see above). At that time no carpi or meri were known, or at least were not attributed to
<taxonomicName id="9F934D65FFBFFFDDFF22858E18686DF2" box="[235,360,1697,1721]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBFFFDDFF22858E18686DF2" box="[235,360,1697,1721]" italics="true" pageId="10" pageNumber="11">C. brocchii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="582C36E6FFBFFFDDFF0E85EB18DC6C03" blockId="10.[151,1437,1264,2009]" pageId="10" pageNumber="11">
<bibRefCitation id="3C024B17FFBFFFDDFF0E85EB18936D97" author="Muller" box="[199,403,1732,1756]" pageId="10" pageNumber="11" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984: 51)</bibRefCitation>
stated that
<taxonomicName id="9F934D65FFBFFFDDFDDE85EA1A016D96" box="[535,769,1733,1757]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBFFFDDFDDE85EA1A016D96" box="[535,769,1733,1757]" italics="true" pageId="10" pageNumber="11">Callianassa brocchii</emphasis>
</taxonomicName>
can be differentiated from similar extant species (
<taxonomicName id="9F934D65FFBFFFDDFAF785EA18DD6C4B" authority="Barnard, 1947" authorityName="Barnard" authorityYear="1947" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gilchristi">
<emphasis id="6AE7EAF4FFBFFFDDFAF785EA18346C4A" italics="true" pageId="10" pageNumber="11">Callianassa gilchristi</emphasis>
<bibRefCitation id="3C024B17FFBFFFDDFEF285C718DD6C4B" author="Barnard" box="[315,477,1768,1792]" pageId="10" pageNumber="11" refString="Barnard, K. H. (1947) Descriptions of new species of South African decapod Crustacea, with notes on synonymy and new records. Annals and Magazine of Natural History, series 11, 13, 361 - 392. [Dated 1946, published 1947]" type="journal article" year="1947">Barnard, 1947</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="9F934D65FFBFFFDDFE2185C61B7A6C4A" box="[488,634,1769,1793]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="6AE7EAF4FFBFFFDDFE2185C61B7A6C4A" box="[488,634,1769,1793]" italics="true" pageId="10" pageNumber="11">C. natalensis</emphasis>
</taxonomicName>
) on the basis of &quot;the saw-like lower edge&quot;. This is not correct, however, as
<taxonomicName id="9F934D65FFBFFFDDFF7C842218476C6E" box="[181,327,1805,1829]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="6AE7EAF4FFBFFFDDFF7C842218476C6E" box="[181,327,1805,1829]" italics="true" pageId="10" pageNumber="11">C. natalensis</emphasis>
</taxonomicName>
does possess serration on the lower margin of the propodus (see e.g.
<bibRefCitation id="3C024B17FFBFFFDDFB8284231C166C6E" author="Dworschak" box="[1099,1302,1804,1829]" pageId="10" pageNumber="11" refString="Dworschak, P. C. (2011 a) Redescription of Callianassa jousseaumei Nobili, 1904, a junior (sic!) subjective synonym of Callianassa indica de Man, 1905 with description of a new species of Neocallichirus (Decapoda: Axiidea: Callianassidae). Zootaxa, 2746, 1 - 19." type="journal article" year="2011" yearSuffix="a">
Dworschak 2011
<emphasis id="6AE7EAF4FFBFFFDDFAC184211C166C6E" box="[1288,1302,1806,1829]" italics="true" pageId="10" pageNumber="11">a</emphasis>
</bibRefCitation>
: fig. 5D, E) as do
<taxonomicName id="9F934D65FFBFFFDDFF11841E187D6C02" box="[216,381,1841,1865]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFBFFFDDFF11841E187D6C02" box="[216,381,1841,1865]" italics="true" pageId="10" pageNumber="11">Neocallichirus</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="1089656DFFBFFFDCFF00847A1B9D6F1C" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="582C36E6FFBFFFDCFF00847A18DA6A74" blockId="10.[151,1437,1264,2009]" lastBlockId="11.[151,1437,151,1329]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<taxonomicName id="9F934D65FFBFFFDDFF00847A18D66C26" box="[201,470,1877,1901]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brocchii" status="comb. nov.">
<emphasis id="6AE7EAF4FFBFFFDDFF00847A18D66C26" box="[201,470,1877,1901]" italics="true" pageId="10" pageNumber="11">Neocallichirus brocchii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71D4578FFFBFFFDDFE28847B1B626C27" box="[481,610,1876,1900]" pageId="10" pageNumber="11">comb. nov.</taxonomicNameLabel>
is morphologically very similar to the extant
<taxonomicName id="9F934D65FFBFFFDDFB4E847A18486CDB" authority="Manning, 1993" authorityName="Manning" authorityYear="1993" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lemaitrei">
<emphasis id="6AE7EAF4FFBFFFDDFB4E847A1C9B6C26" box="[1159,1435,1877,1901]" italics="true" pageId="10" pageNumber="11">Neocallichirus lemaitrei</emphasis>
<bibRefCitation id="3C024B17FFBFFFDDFF5E845718486CDB" author="Manning" box="[151,328,1912,1936]" pageId="10" pageNumber="11" refString="Manning, R. B. (1993) Two new species of Neocallichirus from the Caribbean Sea (Crustacea: Decapoda: Callianassidae). Proceedings of the Biological Society of Washington, 106, 106 - 114." type="journal article" year="1993">Manning, 1993</bibRefCitation>
</taxonomicName>
, and allied species (see also comparison with
<taxonomicName id="9F934D65FFBFFFDDFCAC84551D1D6CDA" box="[869,1053,1913,1937]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="grandimanus">
<emphasis id="6AE7EAF4FFBFFFDDFCAC84551D1D6CDA" box="[869,1053,1913,1937]" italics="true" pageId="10" pageNumber="11">N. grandimanus</emphasis>
</taxonomicName>
and
<taxonomicName id="9F934D65FFBFFFDDFB9384551C086CDA" box="[1114,1288,1914,1937]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="jousseaumei">
<emphasis id="6AE7EAF4FFBFFFDDFB9384551C086CDA" box="[1114,1288,1914,1937]" italics="true" pageId="10" pageNumber="11">N. jousseaumei</emphasis>
</taxonomicName>
above). The armature of the dactylus is similar to extant
<taxonomicName id="9F934D65FFBFFFDDFDB784B11D2F6CFF" authority="Felder &amp; Manning, 1995" authorityName="Felder &amp; Manning" authorityYear="1995" box="[638,1071,1948,1973]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cacahuate">
<emphasis id="6AE7EAF4FFBFFFDDFDB784B11A116CFE" box="[638,785,1949,1973]" italics="true" pageId="10" pageNumber="11">N. cacahuate</emphasis>
<bibRefCitation id="3C024B17FFBFFFDDFCD184B31D2F6CFF" author="Felder" box="[792,1071,1948,1972]" pageId="10" pageNumber="11" refString="Felder, D. L. &amp; Manning, R. B. (1995) Neocallichirus cacahuate, a new species of ghost shrimp from the Atlantic coast of Florida, with reexamination of N. grandimana and N. lemaitrei (Crustacea: Decapoda: Callianassidae). Proceedings of the Biological Society of Washington, 108 (3), 477 - 490." type="journal article" year="1995">Felder &amp; Manning, 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="9F934D65FFBFFFDDFBF084B11DE36CFE" box="[1081,1251,1950,1973]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="jousseaumei">
<emphasis id="6AE7EAF4FFBFFFDDFBF084B11DE36CFE" box="[1081,1251,1950,1973]" italics="true" pageId="10" pageNumber="11">N. jousseaumei</emphasis>
</taxonomicName>
,
<taxonomicName id="9F934D65FFBFFFDDFB3884B11C756CFE" box="[1265,1397,1949,1973]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lemaitrei">
<emphasis id="6AE7EAF4FFBFFFDDFB3884B11C756CFE" box="[1265,1397,1949,1973]" italics="true" pageId="10" pageNumber="11">N. lemaitrei</emphasis>
</taxonomicName>
,
<taxonomicName id="9F934D65FFBFFFDDFA4B84B119EB6C92" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="monodi">
<emphasis id="6AE7EAF4FFBFFFDDFA4B84B119EB6C92" italics="true" pageId="10" pageNumber="11">N. monodi</emphasis>
</taxonomicName>
(de
<bibRefCitation id="3C024B17FFBFFFDDFED484EF1B8E6C93" author="Saint" box="[285,654,1984,2008]" pageId="10" pageNumber="11" refString="Saint Laurent, M. de &amp; Le Loeuff, P. (1979) Upogebiidae et Callianassidae. Crustaces Decapodes Thalassinidea, 1. Campagnes de la Calypso au large des cotes Atlantiques africaines (1956 et 1959) (suite), 22. Resultats scientifiques des campagnes de la Calypso, 11, 29 - 101." type="journal article" year="1979">Saint Laurent &amp; Le Loeuff, 1979</bibRefCitation>
) and
<taxonomicName id="9F934D65FFBFFFDDFD0484ED1A5F6C92" box="[717,863,1985,2009]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="6AE7EAF4FFBFFFDDFD0484ED1A5F6C92" box="[717,863,1985,2009]" italics="true" pageId="10" pageNumber="11">N. natalensis</emphasis>
</taxonomicName>
. In fossil taxa the variation in the shape of the dactylus is not very well known; however, none of those described can be fully compared to
<taxonomicName id="9F934D65FFBEFFDCFB5A83B71C9C6BFB" box="[1171,1436,152,176]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii" status="comb. nov.">
<emphasis id="6AE7EAF4FFBEFFDCFB5A83B71C9C6BFB" box="[1171,1436,152,176]" italics="true" pageId="11" pageNumber="12">Neocallichirus brocchii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71D4578FFFBEFFDCFF5E839318146B9F" box="[151,276,188,212]" pageId="11" pageNumber="12">comb. nov.</taxonomicNameLabel>
The combination of characters as presented in the diagnosis and description is unique for this species, although some similarities can be found with
<taxonomicName id="9F934D65FFBEFFDCFD5F83CF1C306BBC" authority="Schweitzer, Iturralde-Vinent, Hetler" authorityName="Schweitzer, Iturralde-Vinent, Hetler" box="[662,1328,223,248]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aetodes">
<emphasis id="6AE7EAF4FFBEFFDCFD5F83CF1A976BB3" box="[662,919,224,248]" italics="true" pageId="11" pageNumber="12">Neocallichirus aetodes</emphasis>
Schweitzer, Iturralde-Vinent, Hetler
</taxonomicName>
&amp; Velez- Juarbe, 2006 from the Lower Oligocene of
<collectingCountry id="20847676FFBEFFDCFDB1822A1BFD6A57" box="[632,765,260,284]" name="Puerto Rico" pageId="11" pageNumber="12">Puerto Rico</collectingCountry>
and
<taxonomicName id="9F934D65FFBEFFDCFCFB822A1DE76A57" authority="Karasawa &amp; Nakagawa, 2010" authorityName="Karasawa &amp; Nakagawa" authorityYear="2010" box="[818,1255,260,284]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="hattai">
<emphasis id="6AE7EAF4FFBEFFDCFCFB822A1A936A57" box="[818,915,260,284]" italics="true" pageId="11" pageNumber="12">N. hattai</emphasis>
<bibRefCitation id="3C024B17FFBEFFDCFC53822A1DE76A57" author="Karasawa" box="[922,1255,260,284]" pageId="11" pageNumber="12" refString="Karasawa, H. &amp; Nakagawa, T. (2010) A new ghost shrimp (Decapoda: Thalassinidea) from the Miocene Kunimi Formation, Fukui Prefecture, Japan. Bulletin of the Mizunami Fossil Museum, 36, 31 - 36." type="journal article" year="2010">Karasawa &amp; Nakagawa, 2010</bibRefCitation>
</taxonomicName>
from the Lower to Middle Miocene of
<collectingCountry id="20847676FFBEFFDCFE5A820718D76A74" box="[403,471,296,319]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
.
</paragraph>
<paragraph id="582C36E6FFBEFFDCFF0082631BA2685F" blockId="11.[151,1437,151,1329]" pageId="11" pageNumber="12">
The original description of
<taxonomicName id="9F934D65FFBEFFDCFDC982631BEB6A2F" box="[512,747,332,356]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBEFFDCFDC982631BEB6A2F" box="[512,747,332,356]" italics="true" pageId="11" pageNumber="12">Callianassa brocchii</emphasis>
</taxonomicName>
is based on a single specimen of left major propodus articulated with dactylus;
<bibRefCitation id="3C024B17FFBEFFDCFE4F82411B5C6ACC" author="Lorenthey" box="[390,604,366,392]" pageId="11" pageNumber="12" refString="Lorenthey, E. (1898 a) Beitrage zur Decapodenfauna des ungarischen Tertiars. Mathematischen und Naturwissenschaftlichen Berichte aus Ungarn, 14, 92 - 115." type="journal article" year="1898" yearSuffix="a">
Lőrenthey (1898
<emphasis id="6AE7EAF4FFBEFFDCFD8C825E1B536AC3" box="[581,595,369,392]" italics="true" pageId="11" pageNumber="12">a</emphasis>
)
</bibRefCitation>
and
<bibRefCitation id="3C024B17FFBEFFDCFD5282411AE96ACC" author="Lorenthey" box="[667,1001,366,391]" pageId="11" pageNumber="12" refString="Lorenthey, E. &amp; Beurlen, K. (1929) Die fossilen Dekapoden der Lander der Ungarischen Krone. Geologica Hungarica, Series Palaeontologica, 3, 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen (1929)</bibRefCitation>
erroneously treated it as a right one, although the accompanying figures clearly show it as being left, as re-examination of the
<typeStatus id="87288844FFBEFFDCFB5182BA1DC86AE6" box="[1176,1224,405,429]" pageId="11" pageNumber="12">type</typeStatus>
material also confirmed. Lőrenthey in
<bibRefCitation id="3C024B17FFBEFFDCFE4082991BF96A84" author="Lorenthey" box="[393,761,438,463]" pageId="11" pageNumber="12" refString="Lorenthey, E. &amp; Beurlen, K. (1929) Die fossilen Dekapoden der Lander der Ungarischen Krone. Geologica Hungarica, Series Palaeontologica, 3, 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen (1929: 65)</bibRefCitation>
mentioned another two specimens, which may probably be assigned to this species, although no figures of them were added. Another specimen was reported by Beurlen in
<bibRefCitation id="3C024B17FFBEFFDCFF5E82D11B0B695C" author="Lorenthey" box="[151,523,510,535]" pageId="11" pageNumber="12" refString="Lorenthey, E. &amp; Beurlen, K. (1929) Die fossilen Dekapoden der Lander der Ungarischen Krone. Geologica Hungarica, Series Palaeontologica, 3, 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen (1929: 69)</bibRefCitation>
. He classified the material as
<taxonomicName id="9F934D65FFBEFFDCFCA9812F1D706953" box="[864,1136,511,536]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBEFFDCFCA9812F1AE66953" box="[864,998,512,536]" italics="true" pageId="11" pageNumber="12">Callianassa</emphasis>
cf.
<emphasis id="6AE7EAF4FFBEFFDCFBDD812F1D706953" box="[1044,1136,512,536]" italics="true" pageId="11" pageNumber="12">brocchii</emphasis>
</taxonomicName>
, however, he stated that it morphologically conforms with the description of
<taxonomicName id="9F934D65FFBEFFDCFD02810A1A476977" box="[715,839,548,572]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBEFFDCFD02810A1A476977" box="[715,839,548,572]" italics="true" pageId="11" pageNumber="12">C. brocchii</emphasis>
</taxonomicName>
.
<collectingCountry id="20847676FFBEFFDCFC9C810B1A746977" box="[853,884,548,572]" name="American Samoa" pageId="11" pageNumber="12">As</collectingCountry>
no figure of the material was added, we are hesitant to assign the material to the species with certainty. Interestingly in the description it is mentioned that &quot;Form des Carpus, die Grösse, der Index, soweit es das abgebrochene Stück erkennen lässt, stimmen volkommen mit dieser Art überein&quot; (
<collectingCountry id="20847676FFBEFFDCFEB581A0189C69EC" box="[380,412,655,679]" name="American Samoa" pageId="11" pageNumber="12">As</collectingCountry>
far as the broken specimen can be identified, the form of carpus, its size, and the fixed finger correspond precisely to this species).
<collectingCountry id="20847676FFBEFFDCFDAF819B1B856987" box="[614,645,692,716]" name="American Samoa" pageId="11" pageNumber="12">As</collectingCountry>
<taxonomicName id="9F934D65FFBEFFDCFD45819A1A096987" box="[652,777,692,716]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBEFFDCFD45819A1A096987" box="[652,777,692,716]" italics="true" pageId="11" pageNumber="12">C. brocchii</emphasis>
</taxonomicName>
has been described on the basis of an articulated propodus and dactylus only, and no carpus was described in the original description, we assume that Beurlen made a mistake and meant, in fact, propodus instead of carpus.
</paragraph>
<paragraph id="582C36E6FFBEFFDCFF0080301B9D6F1C" blockId="11.[151,1437,151,1329]" pageId="11" pageNumber="12">
There is much confusion in citing Lőrenthey's works.
<collectingCountry id="20847676FFBEFFDCFCEA800F1A426873" box="[803,834,800,824]" name="American Samoa" pageId="11" pageNumber="12">As</collectingCountry>
stated by
<bibRefCitation id="3C024B17FFBEFFDCFC7D80301D80687C" author="Muller" box="[948,1152,799,823]" pageId="11" pageNumber="12" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984: 32)</bibRefCitation>
: &quot;Lőrenthey's first papers were published in parallel Hungarian and
<collectingCountry id="20847676FFBEFFDCFDBB806B1BCC6817" box="[626,716,836,860]" name="Germany" pageId="11" pageNumber="12">German</collectingCountry>
versions with identical contents. He presented the same data in preliminary publications without figures (
<bibRefCitation id="3C024B17FFBEFFDCFDAE80491A1E6834" author="Lorenthey" box="[615,798,870,895]" pageId="11" pageNumber="12" refString="Lorenthey, E. (1897) Adatok Magyarorszag harmadkoru rak faunajahoz. Mathematikai es Termeszettudomanyi Ertesito, 15, 149 - 169." type="journal article" year="1897">Lőrenthey 1897</bibRefCitation>
,
<bibRefCitation id="3C024B17FFBEFFDCFCE080481A716834" author="Lorenthey" box="[809,881,871,895]" pageId="11" pageNumber="12" refString="Lorenthey, E. (1898 a) Beitrage zur Decapodenfauna des ungarischen Tertiars. Mathematischen und Naturwissenschaftlichen Berichte aus Ungarn, 14, 92 - 115." type="journal article" year="1898" yearSuffix="a">1898a</bibRefCitation>
), and in definite papers with excellent figures and detailed descriptions (
<bibRefCitation id="3C024B17FFBEFFDCFE5D80A41B5D68EF" author="Lorenthey" box="[404,605,907,932]" pageId="11" pageNumber="12" refString="Lorenthey, E. (1898 b) Adatok Magyarorszag harmadkoru rak-faunajahoz. Mathematikai es Termeszettudomanyi Kozllemenyek, 27 (2), 103 - 271." type="journal article" year="1898" yearSuffix="b">Lőrenthey 1898b</bibRefCitation>
,
<bibRefCitation id="3C024B17FFBEFFDCFDA580A31B7B68EF" author="Lorenthey" box="[620,635,908,932]" pageId="11" pageNumber="12" refString="Lorenthey, E. (1898 c) Beitrage zur Decapodenfauna des Ungarischen Tertiars. Termeszetrajzi Fuzetek, 21, 1 - 133." type="journal article" year="1898" yearSuffix="c">c</bibRefCitation>
)&quot;.
<taxonomicName id="9F934D65FFBEFFDCFD5680A31A8C68EF" box="[671,908,908,932]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBEFFDCFD5680A31A8C68EF" box="[671,908,908,932]" italics="true" pageId="11" pageNumber="12">Callianassa brocchii</emphasis>
</taxonomicName>
was first described as a new species in such a preliminary publication in 1897. In this paper a short description with comparison with similar taxa can be found which satisfies the ICZN rules for a new species designation. Later the description of
<taxonomicName id="9F934D65FFBEFFDCFBAA80FA1DE068A7" box="[1123,1248,980,1004]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBEFFDCFBAA80FA1DE068A7" box="[1123,1248,980,1004]" italics="true" pageId="11" pageNumber="12">C. brocchii</emphasis>
</taxonomicName>
was refined and figures were added (
<bibRefCitation id="3C024B17FFBEFFDCFEB480D91B436F44" author="Lorenthey" box="[381,579,1014,1039]" pageId="11" pageNumber="12" refString="Lorenthey, E. (1898 b) Adatok Magyarorszag harmadkoru rak-faunajahoz. Mathematikai es Termeszettudomanyi Kozllemenyek, 27 (2), 103 - 271." type="journal article" year="1898" yearSuffix="b">Lőrenthey 1898b</bibRefCitation>
,
<bibRefCitation id="3C024B17FFBEFFDCFD8680D81B5E6F44" author="Lorenthey" box="[591,606,1015,1039]" pageId="11" pageNumber="12" refString="Lorenthey, E. (1898 c) Beitrage zur Decapodenfauna des Ungarischen Tertiars. Termeszetrajzi Fuzetek, 21, 1 - 133." type="journal article" year="1898" yearSuffix="c">c</bibRefCitation>
). Thus, the year 1897 is considered as the year of description of
<taxonomicName id="9F934D65FFBEFFDCFAF780D7182B6F7F" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBEFFDCFAF780D7182B6F7F" italics="true" pageId="11" pageNumber="12">Callianassa brocchii</emphasis>
</taxonomicName>
contrary to
<bibRefCitation id="3C024B17FFBEFFDCFE7A87331B736F7F" author="Glaessner" box="[435,627,1052,1076]" pageId="11" pageNumber="12" refString="Glaessner, M. F. (1929) Crustacea Decapoda. In. Pompeckj, F. J. (Ed.), Fossilium catalogus, 1: Animalium, 41, 1 - 464. W. Junk, Berlin." type="journal article" year="1929">Glaessner (1929)</bibRefCitation>
and
<bibRefCitation id="3C024B17FFBEFFDCFD6387331AB76F7F" box="[682,951,1052,1076]" pageId="11" pageNumber="12" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, 1 - 222." type="journal article">
Schweitzer
<emphasis id="6AE7EAF4FFBEFFDCFCE487321A5E6F7F" box="[813,862,1052,1076]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. (2010)
</bibRefCitation>
where the species appeared as described in 1898 (see also discussion in
<bibRefCitation id="3C024B17FFBEFFDCFE1E87101B666F1C" author="Muller" box="[471,614,1087,1111]" pageId="11" pageNumber="12" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller 1984</bibRefCitation>
: 32).
</paragraph>
</subSubSection>
<subSubSection id="1089656DFFBEFFDCFF00874B1C256E64" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="582C36E6FFBEFFDCFF00874B1C256E64" blockId="11.[151,1437,151,1329]" pageId="11" pageNumber="12">
<bibRefCitation id="3C024B17FFBEFFDCFF00874B18986F37" author="Muller" box="[201,408,1124,1148]" pageId="11" pageNumber="12" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984: 50)</bibRefCitation>
erroneously assumed that the handedness is quite typical for every callianassid species: &quot;Most of recent
<emphasis id="6AE7EAF4FFBEFFDCFE8687A718E76FEB" box="[335,487,1160,1184]" italics="true" pageId="11" pageNumber="12">Callianassids</emphasis>
(sic!) are strongly heterochelous. Surprisingly, in the diverse 'Badenian' material no evidence of heterochely was observed, as right and left chelae are found in same proportion and in same size as well, with the exception of
<taxonomicName id="9F934D65FFBEFFDCFE0E87FF1B436FA3" box="[455,579,1232,1256]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBEFFDCFE0E87FF1B436FA3" box="[455,579,1232,1256]" italics="true" pageId="11" pageNumber="12">C. brocchii</emphasis>
</taxonomicName>
&quot;. It should be noted here that
<bibRefCitation id="3C024B17FFBEFFDCFC5A87E01D306FAC" author="Muller" box="[915,1072,1231,1255]" pageId="11" pageNumber="12" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984)</bibRefCitation>
examined only six specimens of
<taxonomicName id="9F934D65FFBEFFDCFF5E87DA18196E47" box="[151,281,1268,1292]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBEFFDCFF5E87DA18196E47" box="[151,281,1268,1292]" italics="true" pageId="11" pageNumber="12">C. brocchii</emphasis>
</taxonomicName>
comparing to often tens and hundreds of specimens in other &quot;
<taxonomicName id="9F934D65FFBEFFDCFBC087DB1D8F6E47" box="[1033,1167,1268,1292]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="6AE7EAF4FFBEFFDCFBC087DB1D8F6E47" box="[1033,1167,1268,1292]" italics="true" pageId="11" pageNumber="12">Callianassa</emphasis>
</taxonomicName>
&quot; species. Therefore, it seemed to him that
<taxonomicName id="9F934D65FFBEFFDCFEBB863718EF6E7B" box="[370,495,1304,1328]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brocchii">
<emphasis id="6AE7EAF4FFBEFFDCFEBB863718EF6E7B" box="[370,495,1304,1328]" italics="true" pageId="11" pageNumber="12">C. brocchii</emphasis>
</taxonomicName>
was the only heterochelous species among 'Badenian' callianassid forms.
</paragraph>
</subSubSection>
</treatment>
</document>