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<document id="7F03A731D311A34C3AC4ACF29EBD61B1" ID-DOI="10.11646/zootaxa.3619.5.6" ID-GBIF-Dataset="37929844-8332-4afb-8672-d0b5c18ddf8c" ID-ISSN="1175-5326" ID-Zenodo-Dep="216429" ID-ZooBank="0AB793D1-D40D-47FF-9835-C83BBD94CE27" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460578850316" checkinUser="plazi" docAuthor="Tuzovskij, Petr V." docDate="2013" docId="CD0C7A0F243CFFF4B79E3FC2FDFED5C8" docLanguage="en" docName="zt03619p580.pdf" docOrigin="Zootaxa 3619 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lebertia (Pilolebertia) longiseta Bader 1955" docType="treatment" docVersion="7" lastPageNumber="577" masterDocId="313502772438FFFCB7093E1EFF87D704" masterDocTitle="Larval morphology of Lebertia longiseta Bader, 1955, L. dubia Thor, 1899 and Oxus nodigerus Koenike, 1898 (Acari, Hydrachnidia: Lebertiidae, Oxidae)" masterLastPageNumber="580" masterPageNumber="569" pageNumber="573" updateTime="1698323779179" updateUser="plazi">
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<mods:title id="A3B9C2E886F9CEAF3DDBD8F72DD6B2EE">Larval morphology of Lebertia longiseta Bader, 1955, L. dubia Thor, 1899 and Oxus nodigerus Koenike, 1898 (Acari, Hydrachnidia: Lebertiidae, Oxidae)</mods:title>
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<mods:namePart id="5BF2E95DEC038D01EE5020A4D85EF478">Tuzovskij, Petr V.</mods:namePart>
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<mods:date id="C3DC65551642F48F05A78908C7FA28C1">2013</mods:date>
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<treatment id="CD0C7A0F243CFFF4B79E3FC2FDFED5C8" ID-DOI="http://doi.org/10.5281/zenodo.6160821" ID-GBIF-Taxon="119560875" ID-Zenodo-Dep="6160821" LSID="urn:lsid:plazi:treatment:CD0C7A0F243CFFF4B79E3FC2FDFED5C8" httpUri="http://treatment.plazi.org/id/CD0C7A0F243CFFF4B79E3FC2FDFED5C8" lastPageId="8" lastPageNumber="577" pageId="4" pageNumber="573">
<subSubSection id="0DBF9892243CFFF8B79E3FC2FEA0D512" pageId="4" pageNumber="573" type="nomenclature">
<paragraph id="451ACB19243CFFF8B79E3FC2FD36D6F1" blockId="4.[151,689,475,534]" box="[151,689,475,502]" pageId="4" pageNumber="573">
<heading id="1E527C75243CFFF8B79E3FC2FD36D6F1" bold="true" box="[151,689,475,502]" fontSize="11" level="1" pageId="4" pageNumber="573" reason="1">
<taxonomicName id="82A5B09A243CFFF8B79E3FC2FD36D6F1" authority="Bader, 1955" authorityName="Bader" authorityYear="1955" box="[151,689,475,502]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="longiseta" subGenus="Pilolebertia">
<emphasis id="77D1170B243CFFF8B79E3FC2FD36D6F1" bold="true" box="[151,689,475,502]" pageId="4" pageNumber="573">
<emphasis id="77D1170B243CFFF8B79E3FC2FD91D6F2" bold="true" box="[151,534,476,502]" italics="true" pageId="4" pageNumber="573">Lebertia (Pilolebertia) longiseta</emphasis>
Bader, 1955
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="451ACB19243CFFF8B79E3FE0FEA0D512" blockId="4.[151,689,475,534]" box="[151,295,510,534]" pageId="4" pageNumber="573">
(
<figureCitation id="DD9ED79C243CFFF8B7963FE0FF76D512" box="[159,241,510,534]" captionStart="FIGURES 15 16" captionStartId="4.[151,264,2015,2038]" captionTargetBox="[217,1374,1108,1966]" captionTargetId="figure@4.[197,1389,1086,1994]" captionTargetPageId="4" captionText="FIGURES 15 16. Lebertia longiseta Bader, 1955, larva: 15, dorsal view; 16, ventral view. Scale bar: 50 μm." httpUri="https://zenodo.org/record/216432/files/figure.png" pageId="4" pageNumber="573">Figs 15</figureCitation>
26)
</paragraph>
</subSubSection>
<subSubSection id="0DBF9892243CFFF8B79E3C58FB7FD5A2" pageId="4" pageNumber="573" type="materials_examined">
<paragraph id="451ACB19243CFFF8B79E3C58FB7FD5A2" blockId="4.[151,1436,582,1003]" pageId="4" pageNumber="573">
<emphasis id="77D1170B243CFFF8B79E3C58FEFBD55B" bold="true" box="[151,380,582,607]" pageId="4" pageNumber="573">Material examined</emphasis>
. Larvae (n = 16) were reared from four females collected in a cold brook near the village Postyltsevo, Nekouz District, Yaroslavl Province: two females were collected on
<date id="311BEDD9243CFFF8B3303C74FB52D586" box="[1081,1237,618,642]" pageId="4" pageNumber="573" value="2002-06-16">16 June 2002</date>
, one female on
<date id="311BEDD9243CFFF8B2873C74FE82D5A2" pageId="4" pageNumber="573" value="2002-07-02">2 July 2002</date>
and one female on
<date id="311BEDD9243CFFF8B6E93C90FD0DD5A2" box="[480,650,654,679]" pageId="4" pageNumber="573" value="2005-08-08">8 August 2005</date>
. The duration of the embryonic period was 1116 days.
</paragraph>
</subSubSection>
<subSubSection id="0DBF9892243CFFF8B7CE3CACFA16D416" pageId="4" pageNumber="573" type="diagnosis">
<paragraph id="451ACB19243CFFF8B7CE3CACFA16D416" blockId="4.[151,1436,582,1003]" pageId="4" pageNumber="573">
<emphasis id="77D1170B243CFFF8B7CE3CACFEBDD5CF" bold="true" box="[199,314,690,715]" pageId="4" pageNumber="573">Diagnosis</emphasis>
. Dorsal plate elongated (L/W ratio 1.752.15), setae
<emphasis id="77D1170B243CFFF8B4BE3CADFC5FD5CE" box="[951,984,691,714]" italics="true" pageId="4" pageNumber="573">Oe</emphasis>
,
<emphasis id="77D1170B243CFFF8B4E23CAAFBF7D5CE" box="[1003,1136,691,715]" italics="true" pageId="4" pageNumber="573">Hi, He, Sci</emphasis>
and
<emphasis id="77D1170B243CFFF8B3B83CADFB5FD5CE" box="[1201,1240,691,714]" italics="true" pageId="4" pageNumber="573">Sce</emphasis>
equal in length; excretory pore plate wider than long (L/W ratio 0.750.95);
<emphasis id="77D1170B243CFFF8B4623CC8FC0ED5EA" box="[875,905,726,750]" italics="true" pageId="4" pageNumber="573">Ae</emphasis>
longer and thicker than
<emphasis id="77D1170B243CFFF8B3BD3CC8FB4AD5EA" box="[1204,1229,726,750]" italics="true" pageId="4" pageNumber="573">Ai</emphasis>
; chelicerae bases completely fused to each other; IILeg-4 with a single thickened seta, IIILeg-4 with two greatly thickened setae.
</paragraph>
</subSubSection>
<subSubSection id="0DBF9892243CFFFBB7C03D00FEE2D0AD" lastPageId="7" lastPageNumber="576" pageId="4" pageNumber="573" type="description">
<paragraph id="451ACB19243CFFF8B7C03D00FB24D4EE" blockId="4.[151,1436,582,1003]" pageId="4" pageNumber="573">
<emphasis id="77D1170B243CFFF8B7C03D00FEDCD433" bold="true" box="[201,347,798,823]" pageId="4" pageNumber="573">Description.</emphasis>
Idiosoma flat, dorsal plate in unengorged larvae covering large part of dorsum (
<figureCitation id="DD9ED79C243CFFF8B3F43D00FAD5D432" box="[1277,1362,798,822]" captionStart="FIGURES 15 16" captionStartId="4.[151,264,2015,2038]" captionTargetBox="[217,1374,1108,1966]" captionTargetId="figure@4.[197,1389,1086,1994]" captionTargetPageId="4" captionText="FIGURES 15 16. Lebertia longiseta Bader, 1955, larva: 15, dorsal view; 16, ventral view. Scale bar: 50 μm." httpUri="https://zenodo.org/record/216432/files/figure.png" pageId="4" pageNumber="573">Fig. 15</figureCitation>
), with slightly convex lateral margins. Anterior margin of the dorsal plate slightly convex, posterior margin rounded, with short and punctuate scale-like patterns and bearing four pairs of setae: two pairs of simple setae (
<emphasis id="77D1170B243CFFF8B3E93D78FAB6D47A" box="[1248,1329,870,894]" italics="true" pageId="4" pageNumber="573">Fch, Vi</emphasis>
) and two pairs of trichobothria (
<emphasis id="77D1170B243CFFF8B69F3D92FE67D4A6" box="[406,480,907,931]" italics="true" pageId="4" pageNumber="573">Fp, Oi</emphasis>
). Simple setae long and thick, anterior setae (
<emphasis id="77D1170B243CFFF8B4EE3D95FB95D4A7" box="[999,1042,907,931]" italics="true" pageId="4" pageNumber="573">Fch</emphasis>
) shorter and thinner than posterior setae (
<emphasis id="77D1170B243CFFF8B7D63DB1FF73D4C2" box="[223,244,943,966]" italics="true" pageId="4" pageNumber="573">Vi</emphasis>
). Both pairs of trichobothria short and thin, equal in size and branched. Seven pairs of long setae (
<emphasis id="77D1170B243CFFF8B2433DB1FEEBD4EF" italics="true" pageId="4" pageNumber="573">Oe, Hi, He, Sci, Sce, Li, Le</emphasis>
) situated on soft wrinkled membrane:
<emphasis id="77D1170B243CFFF8B4173DCDFCB8D4EE" box="[798,831,979,1002]" italics="true" pageId="4" pageNumber="573">Oe</emphasis>
longest, and
<emphasis id="77D1170B243CFFF8B4DD3DCAFC6BD4EF" box="[980,1004,980,1003]" italics="true" pageId="4" pageNumber="573">Li</emphasis>
and
<emphasis id="77D1170B243CFFF8B3283DCAFBBAD4EF" box="[1057,1085,980,1003]" italics="true" pageId="4" pageNumber="573">Le</emphasis>
shortest.
</paragraph>
<caption id="11DA9B91243CFFF8B79E39C1FB6DD0F7" box="[151,1258,2014,2038]" httpUri="https://zenodo.org/record/216432/files/figure.png" pageId="4" pageNumber="573" targetBox="[217,1374,1108,1966]" targetPageId="4">
<paragraph id="451ACB19243CFFF8B79E39C1FB6DD0F7" blockId="4.[151,1258,2014,2038]" box="[151,1258,2014,2038]" pageId="4" pageNumber="573">
<emphasis id="77D1170B243CFFF8B79E39C1FED4D0F0" bold="true" box="[151,339,2015,2038]" pageId="4" pageNumber="573">FIGURES 1516.</emphasis>
<taxonomicName id="82A5B09A243CFFF8B65339C1FD11D0F1" authority="Bader, 1955" authorityName="Bader" authorityYear="1955" box="[346,662,2015,2037]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="longiseta">
<emphasis id="77D1170B243CFFF8B65339C1FD94D0F1" box="[346,531,2015,2037]" italics="true" pageId="4" pageNumber="573">Lebertia longiseta</emphasis>
Bader, 1955
</taxonomicName>
, larva: 15, dorsal view; 16, ventral view. Scale bar: 50 μm.
</paragraph>
</caption>
<caption id="11DA9B91243DFFF9B79E3B43FC01D295" httpUri="https://zenodo.org/record/216433/files/figure.png" pageId="5" pageNumber="574" targetBox="[220,1352,197,1373]" targetPageId="5">
<paragraph id="451ACB19243DFFF9B79E3B43FC01D295" blockId="5.[151,1436,1373,1426]" pageId="5" pageNumber="574">
<emphasis id="77D1170B243DFFF9B79E3B43FEDDD277" bold="true" box="[151,346,1373,1396]" pageId="5" pageNumber="574">FIGURES 1721.</emphasis>
<taxonomicName id="82A5B09A243DFFF9B66F3B40FD33D270" authority="Bader, 1955" authorityName="Bader" authorityYear="1955" box="[358,692,1374,1396]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="574" phylum="Arthropoda" rank="species" species="longiseta">
<emphasis id="77D1170B243DFFF9B66F3B40FDA2D270" box="[358,549,1374,1396]" italics="true" pageId="5" pageNumber="574">Lebertia longiseta</emphasis>
Bader, 1955
</taxonomicName>
, larva: 1718, excretory pore plate; 19, chelicerae, dorsal view; 20, cheliceral stylet, lateral view; 21, pedipalp, lateral view. Scale bar: 20 μm.
</paragraph>
</caption>
<paragraph id="451ACB19243DFFF9B7C03BA4FCB9D182" blockId="5.[151,1436,1466,2031]" pageId="5" pageNumber="574">
Coxal plates large, plates IIIII on each side completely fused to each other, suture line between them developed only in their lateral parts (
<figureCitation id="DD9ED79C243DFFF9B5303BC0FD0AD2F2" box="[569,653,1502,1526]" captionStart="FIGURES 15 16" captionStartId="4.[151,264,2015,2038]" captionTargetBox="[217,1374,1108,1966]" captionTargetId="figure@4.[197,1389,1086,1994]" captionTargetPageId="4" captionText="FIGURES 15 16. Lebertia longiseta Bader, 1955, larva: 15, dorsal view; 16, ventral view. Scale bar: 50 μm." httpUri="https://zenodo.org/record/216432/files/figure.png" pageId="5" pageNumber="574">Fig. 16</figureCitation>
). Anterior coxal plates completely separated from coxal plates II and III. Setae C1 slightly shorter than other coxal setae (C2C4) and approximately equal in length. Posterior margin of coxal plate III has two unequal setae,
<emphasis id="77D1170B243DFFF9B5373839FDDBD13A" box="[574,604,1575,1598]" italics="true" pageId="5" pageNumber="574">Pe</emphasis>
slightly longer than
<emphasis id="77D1170B243DFFF9B44D3839FCDAD13A" box="[836,861,1575,1598]" italics="true" pageId="5" pageNumber="574">Pi</emphasis>
. Setae
<emphasis id="77D1170B243DFFF9B4A73839FC4FD13A" box="[942,968,1575,1598]" italics="true" pageId="5" pageNumber="574">Ci</emphasis>
very long, located on small tubercles and much longer than
<emphasis id="77D1170B243DFFF9B66B3855FEFFD166" box="[354,376,1611,1634]" italics="true" pageId="5" pageNumber="574">Si</emphasis>
and
<emphasis id="77D1170B243DFFF9B6A63855FE4DD166" box="[431,458,1611,1634]" italics="true" pageId="5" pageNumber="574">Se</emphasis>
. All coxal plates with reticulated pattern consisting of elongated cells. Urstigma lateral to the border between coxal plates I and II, covered by flaps.
</paragraph>
<paragraph id="451ACB19243DFFF9B7C0388DFE5BD012" blockId="5.[151,1436,1466,2031]" pageId="5" pageNumber="574">
Excretory pore plate wider than long, with more or less developed anteromedial projection (
<figureCitation id="DD9ED79C243DFFF9B205388CFA09D1AE" box="[1292,1422,1682,1706]" captionStart="FIGURES 17 21" captionStartId="5.[151,264,1373,1396]" captionTargetBox="[220,1352,197,1373]" captionTargetId="figure@5.[220,1367,171,1373]" captionTargetPageId="5" captionText="FIGURES 17 21. Lebertia longiseta Bader, 1955, larva: 17 18, excretory pore plate; 19, chelicerae, dorsal view; 20, cheliceral stylet, lateral view; 21, pedipalp, lateral view. Scale bar: 20 μm." httpUri="https://zenodo.org/record/216433/files/figure.png" pageId="5" pageNumber="574">Fig. 1718</figureCitation>
), rounded posteriorly; setae
<emphasis id="77D1170B243DFFF9B6CF38A8FE64D1CA" box="[454,483,1718,1742]" italics="true" pageId="5" pageNumber="574">Ae</emphasis>
much longer than
<emphasis id="77D1170B243DFFF9B5B538A8FD52D1CA" box="[700,725,1718,1742]" italics="true" pageId="5" pageNumber="574">Ai</emphasis>
, located laterally in the posterior half of plate. Excretory pore placed posteromedial to
<emphasis id="77D1170B243DFFF9B6A538C4FE4ED1F6" box="[428,457,1754,1778]" italics="true" pageId="5" pageNumber="574">Ae</emphasis>
near posterior margin of plate; distance between setae
<emphasis id="77D1170B243DFFF9B33338C4FB04D1F6" box="[1082,1155,1754,1778]" italics="true" pageId="5" pageNumber="574">AeAe</emphasis>
almost two times longer than distance between
<emphasis id="77D1170B243DFFF9B69C38E0FE52D012" box="[405,469,1790,1814]" italics="true" pageId="5" pageNumber="574">AiAi</emphasis>
.
</paragraph>
<paragraph id="451ACB19243DFFF9B7C0393CFB18D05A" blockId="5.[151,1436,1466,2031]" pageId="5" pageNumber="574">
Chelicerae bases completely fused to each other (
<figureCitation id="DD9ED79C243DFFF9B5F8393CFCC4D03E" box="[753,835,1826,1850]" captionStart="FIGURES 17 21" captionStartId="5.[151,264,1373,1396]" captionTargetBox="[220,1352,197,1373]" captionTargetId="figure@5.[220,1367,171,1373]" captionTargetPageId="5" captionText="FIGURES 17 21. Lebertia longiseta Bader, 1955, larva: 17 18, excretory pore plate; 19, chelicerae, dorsal view; 20, cheliceral stylet, lateral view; 21, pedipalp, lateral view. Scale bar: 20 μm." httpUri="https://zenodo.org/record/216433/files/figure.png" pageId="5" pageNumber="574">Fig. 19</figureCitation>
), its dorsal margins convex, ventral margins concave; cheliceral stylets relatively small, crescentshaped, with two small subapical teeth (
<figureCitation id="DD9ED79C243DFFF9B3373958FB15D05A" box="[1086,1170,1862,1886]" captionStart="FIGURES 17 21" captionStartId="5.[151,264,1373,1396]" captionTargetBox="[220,1352,197,1373]" captionTargetId="figure@5.[220,1367,171,1373]" captionTargetPageId="5" captionText="FIGURES 17 21. Lebertia longiseta Bader, 1955, larva: 17 18, excretory pore plate; 19, chelicerae, dorsal view; 20, cheliceral stylet, lateral view; 21, pedipalp, lateral view. Scale bar: 20 μm." httpUri="https://zenodo.org/record/216433/files/figure.png" pageId="5" pageNumber="574">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="451ACB19243DFFF9B7C03974FAC1D0EA" blockId="5.[151,1436,1466,2031]" pageId="5" pageNumber="574">
Pedipalps moderately developed (
<figureCitation id="DD9ED79C243DFFF9B54D3974FD1FD086" box="[580,664,1898,1922]" captionStart="FIGURES 17 21" captionStartId="5.[151,264,1373,1396]" captionTargetBox="[220,1352,197,1373]" captionTargetId="figure@5.[220,1367,171,1373]" captionTargetPageId="5" captionText="FIGURES 17 21. Lebertia longiseta Bader, 1955, larva: 17 18, excretory pore plate; 19, chelicerae, dorsal view; 20, cheliceral stylet, lateral view; 21, pedipalp, lateral view. Scale bar: 20 μm." httpUri="https://zenodo.org/record/216433/files/figure.png" pageId="5" pageNumber="574">Fig. 21</figureCitation>
): P1 short and glabrous; P2 large with convex dorsal margin and a single dorsoproximal seta; P3 with very long, thick lateroproximal seta and a relatively short, fine dorsoproximal one; P4 with two rather long unequal lateroproximal setae, one relatively short dorsodistal seta and a massive dorsodistal claw; P5 small, with single solenidion and seven unequal simple setae, two of them very long.
</paragraph>
<paragraph id="451ACB19243EFFFAB79E3858FDD3D178" blockId="6.[151,1436,1606,1662]" pageId="6" pageNumber="575">
<emphasis id="77D1170B243EFFFAB79E3858FED3D158" bold="true" box="[151,340,1606,1629]" pageId="6" pageNumber="575">FIGURES 2226.</emphasis>
<taxonomicName id="82A5B09A243EFFFAB6503859FD13D159" authority="Bader, 1955" authorityName="Bader" authorityYear="1955" box="[345,660,1607,1629]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="575" phylum="Arthropoda" rank="species" species="longiseta">
<emphasis id="77D1170B243EFFFAB6503859FD95D159" box="[345,530,1607,1629]" italics="true" pageId="6" pageNumber="575">Lebertia longiseta</emphasis>
Bader, 1955
</taxonomicName>
, larva: 22, leg I; 23, leg II; 24, leg III; 25, claws of leg III, 26—claw of leg I. Scale bars: 2224 = 50 μm, 2526 = 20 μm.
</paragraph>
<paragraph id="451ACB19243EFFFBB7C038B0FF4CD7F3" blockId="6.[151,1436,1709,2022]" lastBlockId="7.[151,1436,151,535]" lastPageId="7" lastPageNumber="576" pageId="6" pageNumber="575">Legs 5segmented, basifemur and telofemur of all legs fused to each other, with indistinct suture line. Shape and arrangement of setae on legs segments as shown in Figs 2224. Total number of leg setae, excluding eupathidia, is as follows (specialized setae given in parentheses): ILeg-15: 1, 7, 5 (s), 10 (2s), 15 (s, ac); IILeg- 15: 1, 7, 5 (s), 11 (2s), 15 (s, ac); IIILeg-15: 1, 6, 5 (s), 10 (s), 12 (11); proximal ventral setae on tarsus III sometimes absent. Number of thickened setae from trochanter to tarsus: ILeg: 0, 1, 1, 1, 0; IILeg: 0, 2, 2, 1, 0; IIILeg: 0, 2, 2, 2, 0. ILeg-1 with relatively short seta, IILeg-1 and IIILeg-1 each with long seta. Solenidia on genu of legs III and tibia III short and approximately equal in length; solenidion on genu III shorter than other legs solenidia. Both solenidia on I-leg-4 located in distal half of segments; proximal solenidion on II-leg-4 and solenidion on III-leg-4 situated proximally to middle of these segments. Acanthoid setae are present only on tarsus I and II, each bent two times. Claws of legs III (Fig. 25) larger than claws of legs I and II (Fig. 26). Lateral claws longer and thinner than heavy empodial claw: all claws are sickle shaped and provided with a lateral spur on each side.</paragraph>
<paragraph id="451ACB19243FFFFBB7C03F1BFC63D513" blockId="7.[151,1436,151,535]" pageId="7" pageNumber="576">
Measurements, n=10. L of dorsal plate 255275, W 125155; L of setae
<emphasis id="77D1170B243FFFFBB3123F1AFBC0D618" box="[1051,1095,260,284]" italics="true" pageId="7" pageNumber="576">Fch</emphasis>
6470, L of setae
<emphasis id="77D1170B243FFFFBB22D3F1BFAC4D618" box="[1316,1347,261,284]" italics="true" pageId="7" pageNumber="576">Fp</emphasis>
and
<emphasis id="77D1170B243FFFFBB2893F1BFA1BD618" box="[1408,1436,261,284]" italics="true" pageId="7" pageNumber="576">Oi</emphasis>
1923, L of setae Vi 140150, L of setae
<emphasis id="77D1170B243FFFFBB5783F36FD6AD644" box="[625,749,296,320]" italics="true" pageId="7" pageNumber="576">Oe, Hi, He</emphasis>
110125; L of setae,
<emphasis id="77D1170B243FFFFBB4EF3F36FB8FD63B" box="[998,1032,296,319]" italics="true" pageId="7" pageNumber="576">Sci</emphasis>
and
<emphasis id="77D1170B243FFFFBB3483F36FBEFD63B" box="[1089,1128,296,319]" italics="true" pageId="7" pageNumber="576">Sce</emphasis>
110125; L of setae
<emphasis id="77D1170B243FFFFBB2503F37FA1BD644" box="[1369,1436,297,320]" italics="true" pageId="7" pageNumber="576">Li, Le</emphasis>
95115; L of setae
<emphasis id="77D1170B243FFFFBB6703F53FE09D660" box="[377,398,333,356]" italics="true" pageId="7" pageNumber="576">Si</emphasis>
8085, L of setae
<emphasis id="77D1170B243FFFFBB5653F53FD00D660" box="[620,647,333,356]" italics="true" pageId="7" pageNumber="576">Se</emphasis>
90105, L of setae
<emphasis id="77D1170B243FFFFBB47A3F53FC09D660" box="[883,910,333,356]" italics="true" pageId="7" pageNumber="576">Ci</emphasis>
140150, L of setae
<emphasis id="77D1170B243FFFFBB38E3F53FB27D660" box="[1159,1184,333,356]" italics="true" pageId="7" pageNumber="576">Pi</emphasis>
6065, L of setae
<emphasis id="77D1170B243FFFFBB2773F53FA1CD660" box="[1406,1435,333,356]" italics="true" pageId="7" pageNumber="576">Pe</emphasis>
7075, L of setae C1 6575, L of setae C2C4 100115; L of medial margins of coxae
<date id="311BEDD9243FFFFBB3923F6EFB4BD683" box="[1179,1228,367,391]" pageId="7" pageNumber="576" value="1980-01">I 80</date>
100, L of medial margins of coxae IIIII 110125; L of excretory pore plate 2030, W 2535; L of capitulum 8996; L of basal segments of chelicerae 8390, L of cheliceral stylet 1216; L of pedipalpal segments (P15): 913, 3542, 2938, 2530, 1216; L of legs segments: ILeg-15: 2835, 4952, 5055, 6570, 6480; IILeg-15: 4048, 4852, 5055, 8083, 89100; IIILeg-15: 4555, 5560, 5765, 8993, 95115.
</paragraph>
<paragraph id="451ACB19243FFFFBB79E396DFEE2D0AD" blockId="7.[151,1436,1907,1962]" pageId="7" pageNumber="576">
<emphasis id="77D1170B243FFFFBB79E396DFED1D08C" bold="true" box="[151,342,1907,1930]" pageId="7" pageNumber="576">FIGURES 2729.</emphasis>
<taxonomicName id="82A5B09A243FFFFBB654396DFD20D08D" authority="Koch, 1837" authorityName="Koch" authorityYear="1837" box="[349,679,1907,1929]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="576" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="77D1170B243FFFFBB654396DFDA3D08D" box="[349,548,1907,1929]" italics="true" pageId="7" pageNumber="576">Lebertia inaequalis</emphasis>
Koch, 1837
</taxonomicName>
, larva: 27, ventral view; 2829, excretory pore plate. Scale bars: 27 = 50 μm, 2829 = 20 μm.
</paragraph>
</subSubSection>
<subSubSection id="0DBF98922430FFF4B7C03E86FD6FD55B" pageId="8" pageNumber="577" type="discussion">
<paragraph id="451ACB192430FFF4B7C03E86FBB7D683" blockId="8.[151,1436,151,716]" pageId="8" pageNumber="577">
<emphasis id="77D1170B2430FFF4B7C03E86FEBAD7B4" bold="true" box="[201,317,152,176]" pageId="8" pageNumber="577">Remarks.</emphasis>
The larva of
<taxonomicName id="82A5B09A2430FFF4B6DB3E87FDD1D7B4" box="[466,598,152,176]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="577" phylum="Arthropoda" rank="species" species="longiseta">
<emphasis id="77D1170B2430FFF4B6DB3E87FDD1D7B4" box="[466,598,152,176]" italics="true" pageId="8" pageNumber="577">L. longiseta</emphasis>
</taxonomicName>
is similar to the larva of
<taxonomicName id="82A5B09A2430FFF4B4623E87FB1DD7AB" authority="Koch, 1837" authorityName="Koch" authorityYear="1837" box="[875,1178,151,176]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="577" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="77D1170B2430FFF4B4623E87FC7AD7B4" box="[875,1021,152,176]" italics="true" pageId="8" pageNumber="577">L. inaequalis</emphasis>
(Koch, 1837)
</taxonomicName>
, the latter described by Wainstein (1980) and Martin (2000), but the description were very different, most probably refer two different species. A comparison is made with the Yaroslavl larvae of this species, which is similar to Wainsteins description (Figs 2729). The idiosoma in unengorged of larva
<taxonomicName id="82A5B09A2430FFF4B5D03F1BFCEBD618" box="[729,876,260,284]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="577" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="77D1170B2430FFF4B5D03F1BFCEBD618" box="[729,876,260,284]" italics="true" pageId="8" pageNumber="577">L. inaequalis</emphasis>
</taxonomicName>
has a distinct posteromedian protrusion (Fig. 27), the excretory pore plate has short setae, which are thick and nearly equal in length (Fig. 2829). In contrast, the idiosoma in unengorged of larval
<taxonomicName id="82A5B09A2430FFF4B5103F53FD18D660" box="[537,671,332,356]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="577" phylum="Arthropoda" rank="species" species="longiseta">
<emphasis id="77D1170B2430FFF4B5103F53FD18D660" box="[537,671,332,356]" italics="true" pageId="8" pageNumber="577">L. longiseta</emphasis>
</taxonomicName>
are without posteromedian protrusion (
<figureCitation id="DD9ED79C2430FFF4B36B3F52FB31D660" box="[1122,1206,332,356]" captionStart="FIGURES 15 16" captionStartId="4.[151,264,2015,2038]" captionTargetBox="[217,1374,1108,1966]" captionTargetId="figure@4.[197,1389,1086,1994]" captionTargetPageId="4" captionText="FIGURES 15 16. Lebertia longiseta Bader, 1955, larva: 15, dorsal view; 16, ventral view. Scale bar: 50 μm." httpUri="https://zenodo.org/record/216432/files/figure.png" pageId="8" pageNumber="577">Fig. 16</figureCitation>
), the excretory pore plate has relatively long, thin unequal setae,
<emphasis id="77D1170B2430FFF4B5803F6EFD21D68C" box="[649,678,368,392]" italics="true" pageId="8" pageNumber="577">Ae</emphasis>
much longer than
<emphasis id="77D1170B2430FFF4B4703F6EFC15D68C" box="[889,914,368,392]" italics="true" pageId="8" pageNumber="577">Ai</emphasis>
(
<figureCitation id="DD9ED79C2430FFF4B4A93F71FBA4D683" box="[928,1059,367,391]" captionStart="FIGURES 17 21" captionStartId="5.[151,264,1373,1396]" captionTargetBox="[220,1352,197,1373]" captionTargetId="figure@5.[220,1367,171,1373]" captionTargetPageId="5" captionText="FIGURES 17 21. Lebertia longiseta Bader, 1955, larva: 17 18, excretory pore plate; 19, chelicerae, dorsal view; 20, cheliceral stylet, lateral view; 21, pedipalp, lateral view. Scale bar: 20 μm." httpUri="https://zenodo.org/record/216433/files/figure.png" pageId="8" pageNumber="577">Figs 1718</figureCitation>
).
</paragraph>
<paragraph id="451ACB192430FFF4B7C03F8AFE63D538" blockId="8.[151,1436,151,716]" pageId="8" pageNumber="577">
The morphology of the larva of
<taxonomicName id="82A5B09A2430FFF4B5263F8BFD34D6A8" box="[559,691,404,428]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="577" phylum="Arthropoda" rank="species" species="longiseta">
<emphasis id="77D1170B2430FFF4B5263F8BFD34D6A8" box="[559,691,404,428]" italics="true" pageId="8" pageNumber="577">L. longiseta</emphasis>
</taxonomicName>
is very similar to Martins larva
<taxonomicName id="82A5B09A2430FFF4B3163F8BFB36D6A8" box="[1055,1201,404,428]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="577" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="77D1170B2430FFF4B3163F8BFB36D6A8" box="[1055,1201,404,428]" italics="true" pageId="8" pageNumber="577">L. inaequalis</emphasis>
</taxonomicName>
but these species are not identical. In Martins larva
<taxonomicName id="82A5B09A2430FFF4B6F83FA7FD04D6D4" box="[497,643,440,464]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="577" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="77D1170B2430FFF4B6F83FA7FD04D6D4" box="[497,643,440,464]" italics="true" pageId="8" pageNumber="577">L. inaequalis</emphasis>
</taxonomicName>
III-Leg-5 with 10 setae + 1 solenidion, all segments of III-Leg without thickened setae (Martin 2000). In contrast, in the larva of
<taxonomicName id="82A5B09A2430FFF4B4153FC3FC26D6F0" box="[796,929,476,500]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="577" phylum="Arthropoda" rank="species" species="longiseta">
<emphasis id="77D1170B2430FFF4B4153FC3FC26D6F0" box="[796,929,476,500]" italics="true" pageId="8" pageNumber="577">L. longiseta</emphasis>
</taxonomicName>
III-Leg-5 bearing only simple setae, III-Leg- 24 with two thickened setae each. I believe, that is possible Martins larva
<taxonomicName id="82A5B09A2430FFF4B3063C1FFB22D51C" box="[1039,1189,512,536]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="577" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="77D1170B2430FFF4B3063C1FFB22D51C" box="[1039,1189,512,536]" italics="true" pageId="8" pageNumber="577">L. inaequalis</emphasis>
</taxonomicName>
belongs to any other species of the genus
<taxonomicName id="82A5B09A2430FFF4B6773C3AFE5AD538" box="[382,477,548,572]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="77D1170B2430FFF4B6773C3AFE5AD538" box="[382,477,548,572]" italics="true" pageId="8" pageNumber="577">Lebertia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="451ACB192430FFF4B7C03C59FD6FD55B" blockId="8.[151,1436,151,716]" box="[201,744,583,607]" pageId="8" pageNumber="577">
This species is found in
<collectingCountry id="3DB28B892430FFF4B6D13C59FDA4D55B" box="[472,547,583,607]" name="Russia" pageId="8" pageNumber="577">Russia</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
<subSubSection id="0DBF98922430FFF4B7CE3C75FDFED5C8" pageId="8" pageNumber="577" type="distribution">
<paragraph id="451ACB192430FFF4B7CE3C75FE85D5A3" blockId="8.[151,1436,151,716]" pageId="8" pageNumber="577">
<emphasis id="77D1170B2430FFF4B7CE3C75FEE7D580" bold="true" box="[199,352,619,644]" pageId="8" pageNumber="577">Distribution.</emphasis>
Europe: Northern Alps and Pre-Alps (K.
<collectingCountry id="3DB28B892430FFF4B45C3C72FCEBD580" box="[853,876,620,644]" name="Iceland" pageId="8" pageNumber="577">O</collectingCountry>
. Viets 1978, Gerecke 2009);
<collectingCountry id="3DB28B892430FFF4B3DA3C72FAA7D580" box="[1235,1312,620,644]" name="Russia" pageId="8" pageNumber="577">Russia</collectingCountry>
, Yaroslavl Province.
</paragraph>
<paragraph id="451ACB192430FFF4B7CE3CADFDFED5C8" blockId="8.[151,1436,151,716]" box="[199,633,691,716]" pageId="8" pageNumber="577">
<emphasis id="77D1170B2430FFF4B7CE3CADFEADD5C8" bold="true" box="[199,298,691,716]" pageId="8" pageNumber="577">Habitat.</emphasis>
Standing and running waters.
</paragraph>
</subSubSection>
</treatment>
</document>