287 lines
35 KiB
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287 lines
35 KiB
XML
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<mods:title id="3EC9E8C07167D2671050E11CF29718B0">Review of the genus Engyprosopon Günther, 1862 (Pleuronectiformes: Bothidae) from waters off Taiwan, with descriptions of two new species</mods:title>
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<mods:namePart id="64CEFAB17E5C73560E782B4A1F591119">Amaoka, Kunio</mods:namePart>
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<mods:namePart id="53A29A828388A883C4308688E52A79F6">Ho, Hsuan-Ching</mods:namePart>
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<mods:title id="1A0A34CC640EAD3AC2E1CBE2841D1902">Zootaxa</mods:title>
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<treatment id="B25B3C79FFB5FF9630E1FB07FE059DFB" ID-DOI="http://doi.org/10.5281/zenodo.5967712" ID-GBIF-Taxon="143666440" ID-Zenodo-Dep="5967712" LSID="urn:lsid:plazi:treatment:B25B3C79FFB5FF9630E1FB07FE059DFB" httpUri="http://treatment.plazi.org/id/B25B3C79FFB5FF9630E1FB07FE059DFB" lastPageId="10" lastPageNumber="459" pageId="7" pageNumber="456">
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<paragraph id="3A4D8D6FFFB5FF9B30E1FB07FD159B95" blockId="7.[151,679,1196,1255]" box="[151,679,1196,1222]" pageId="7" pageNumber="456">
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<taxonomicName id="FDF2F6ECFFB5FF9B30E1FB07FD159B95" ID-CoL="39TRN" authority="Amaoka, 1963" authorityName="Amaoka" authorityYear="1963" box="[151,679,1196,1222]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="7" pageNumber="456" phylum="Chordata" rank="species" species="multisquama">
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<emphasis id="0886517DFFB5FF9B30E1FB07FD159B95" bold="true" box="[151,679,1196,1222]" pageId="7" pageNumber="456">
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<emphasis id="0886517DFFB5FF9B30E1FB07FE5E9B95" bold="true" box="[151,492,1196,1222]" italics="true" pageId="7" pageNumber="456">Engyprosopon multisquama</emphasis>
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<bibRefCitation id="5E63F09EFFB5FF9B3185FB07FD159B95" author="Amaoka, K." box="[499,679,1196,1222]" journalOrPublisher="Bulletin of the Misaki Marine Biological Institute, Kyoto University" pageId="7" pageNumber="456" pagination="107 - 121" part="4" refId="ref14181" refString="Amaoka, K. (1963) A revision of the flatfishes referable to the genus Engyprosopon found in the waters around Japan. Bulletin of the Misaki Marine Biological Institute, Kyoto University, (4), 107 - 121." title="A revision of the flatfishes referable to the genus Engyprosopon found in the waters around Japan" type="journal article" year="1963">Amaoka, 1963</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="3A4D8D6FFFB5FF9B30E1FB65FECF9BB4" blockId="7.[151,679,1196,1255]" box="[151,381,1230,1255]" pageId="7" pageNumber="456">
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<figureCitation id="A2C991EAFFB5FF9B30E1FB65FEAD9BB4" box="[151,287,1230,1255]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="8.[151,250,1579,1601]" captionStartId-1="9.[151,250,1578,1600]" captionTargetBox-0="[151,1436,193,1557]" captionTargetBox-1="[151,1436,193,1556]" captionTargetId-0="figure@8.[151,1436,193,1557]" captionTargetId-1="figure@9.[151,1436,193,1556]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 4. Engyprosopon multisquama Amaoka, 1963. Ocular side, NMMB-P8852, male, 89.5 mm SL. B. NMMB-P14467, female, 75.3 mm SL." captionText-1="FIGURE 5. Engyprosopon multisquama Amaoka, 1963.Blind side, same specimens as those in Figure 4." httpUri-0="https://zenodo.org/record/1227799/files/figure.png" httpUri-1="https://zenodo.org/record/1227801/files/figure.png" pageId="7" pageNumber="456">Figures 4–5</figureCitation>
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;
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<tableCitation id="7770B8D4FFB5FF9B315DFB65FECF9BB4" box="[299,381,1230,1255]" captionStart="TABLE 1" captionStartId="15.[151,239,1661,1684]" captionText="TABLE 1. Morphometric data (% of SL) of five Engyprosopon species in Taiwan. M = male(s); F = female(s); O = ocular side; B = blind side." pageId="7" pageNumber="456">Table 1</tableCitation>
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<subSubSection id="72E8DEE4FFB5FF9B30E1FAB3FB4A9A18" pageId="7" pageNumber="456" type="reference_group">
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<paragraph id="3A4D8D6FFFB5FF9B30E1FAB3FB4A9A18" blockId="7.[151,1436,1302,1355]" pageId="7" pageNumber="456">
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<taxonomicName id="FDF2F6ECFFB5FF9B30E1FAB3FDC99A7F" ID-CoL="39TRN" authority="Amaoka, 1963: 111" authorityName="Amaoka" authorityPageNumber="111" authorityYear="1963" box="[151,635,1302,1325]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="7" pageNumber="456" phylum="Chordata" rank="species" species="multisquama">
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<emphasis id="0886517DFFB5FF9B30E1FAB3FE019A7F" box="[151,435,1302,1325]" italics="true" pageId="7" pageNumber="456">Engyprosopon multisquama</emphasis>
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<bibRefCitation id="5E63F09EFFB5FF9B31CFFABDFDE39A7F" author="Amaoka, K." box="[441,593,1302,1324]" journalOrPublisher="Bulletin of the Misaki Marine Biological Institute, Kyoto University" pageId="7" pageNumber="456" pagination="107 - 121" part="4" refId="ref14181" refString="Amaoka, K. (1963) A revision of the flatfishes referable to the genus Engyprosopon found in the waters around Japan. Bulletin of the Misaki Marine Biological Institute, Kyoto University, (4), 107 - 121." title="A revision of the flatfishes referable to the genus Engyprosopon found in the waters around Japan" type="journal article" year="1963">Amaoka, 1963</bibRefCitation>
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:111
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</taxonomicName>
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, fig. 3 (type locality: Susaki, Kochi Prefecture, Japan).
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<bibRefCitation id="5E63F09EFFB5FF9B34DDFABDFAF19A7F" author="Amaoka, K." box="[1195,1347,1302,1324]" journalOrPublisher="Shimonoseki University of Fisheries" pageId="7" pageNumber="456" pagination="65 - 340" part="18" refId="ref14224" refString="Amaoka, K. (1969) Studies on the sinistral flounders found in the waters around Japan. Taxonomy, anatomy and phylogeny. Shimonoseki University of Fisheries, 18 (2), 65 - 340." title="Studies on the sinistral flounders found in the waters around Japan. Taxonomy, anatomy and phylogeny" type="journal article" year="1969">Amaoka, 1969</bibRefCitation>
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||
:146, fig. 47;
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<bibRefCitation id="5E63F09EFFB5FF9B309BFA9EFED19A18" author="Shen, S. - C." box="[237,355,1333,1355]" journalOrPublisher="Journal Taiwan Museum" pageId="7" pageNumber="456" pagination="1 - 42" part="36" refId="ref16090" refString="Shen, S. - C. (1983) Study on the bothid fishes (Family Bothidae) from Taiwan. Journal Taiwan Museum, 36 (1), 1 - 42." title="Study on the bothid fishes (Family Bothidae) from Taiwan" type="journal article" year="1983">Shen, 1983</bibRefCitation>
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:20, fig. 26;
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<bibRefCitation id="5E63F09EFFB5FF9B31AAFA9EFDE29A18" author="Shen, S. - C." box="[476,592,1333,1355]" editor="Shen, S. - C." journalOrPublisher="National Taiwan University, Taipei" pageId="7" pageNumber="456" pagination="565 - 585" refId="ref16013" refString="Shen, S. - C. (1993) Pleuronectiformes. In: Shen, S. - C. (Ed.), Fishes of Taiwan. National Taiwan University, Taipei, pp. 565 - 585." title="Pleuronectiformes" type="book chapter" volumeTitle="Fishes of Taiwan" year="1993">Shen, 1993</bibRefCitation>
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||
:569, pl. 19-9;
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<bibRefCitation id="5E63F09EFFB5FF9B3295FA9EFC209A18" author="Shao, K. - T. & Ho, H. - C. & Lin, P. - L. & Lee, P. - F. & Lee, M. - Y. & Tsai, C. - Y. & Liao, Y. - C. & Lin, Y. - C." box="[739,914,1333,1355]" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="7" pageNumber="456" pagination="233 - 271" part="19" refId="ref15921" refString="Shao, K. - T., Ho, H. - C., Lin, P. - L., Lee, P. - F., Lee, M. - Y., Tsai, C. - Y., Liao, Y. - C. & Lin, Y. - C. (2008) A checklist of the fishes of southern Taiwan, Northern South China Sea. Raffles Bulletin of Zoology, Supplement 19, 233 - 271." title="A checklist of the fishes of southern Taiwan, Northern South China Sea" type="journal article" year="2008">
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Shao
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<emphasis id="0886517DFFB5FF9B336AFA9DFCFB9A18" box="[796,841,1333,1355]" italics="true" pageId="7" pageNumber="456">et al</emphasis>
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., 2008
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</bibRefCitation>
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:264;
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<bibRefCitation id="5E63F09EFFB5FF9B33BFFA9EFB369A18" author="Shen, S. - C. & Wu, K. - Y." bookContentInfo="896 pp." box="[969,1156,1333,1355]" journalOrPublisher="National Museum of Marine Biology and Aquarium, Pingtung" pageId="7" pageNumber="456" refId="ref16055" refString="Shen, S. - C. & Wu, K. - Y. (2012) Fishes of Taiwan. National Museum of Marine Biology and Aquarium, Pingtung, 896 pp." title="Fishes of Taiwan" type="book" year="2012">Shen & Wu, 2012</bibRefCitation>
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||
:751, 2 figs.
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</paragraph>
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<emphasis id="0886517DFFB5FF9B30E1FADCFE319AC3" bold="true" box="[151,387,1399,1424]" pageId="7" pageNumber="456">Material examined.</emphasis>
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,
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|
||
<specimenCode id="6A542514FFB5FF9B31FBFA14FDFA9A8B" box="[397,584,1471,1496]" collectionCode="NMMB-P" pageId="7" pageNumber="456">NMMB-P22258</specimenCode>
|
||
,
|
||
<specimenCount id="2CF446E6FFB5FF9B3224FA6BFD199A8B" box="[594,683,1471,1496]" pageId="7" pageNumber="456" type="male">2 males</specimenCount>
|
||
, 82.0–97.0 mm SL, Ke-tzu-liao,
|
||
<collectingRegion id="F836438DFFB5FF9B346CFA14FB249A8B" box="[1050,1174,1471,1496]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="456">Kaohsiung</collectingRegion>
|
||
,
|
||
<collectingCountry id="42E5CDFFFFB5FF9B34D7FA14FB449A8B" box="[1185,1270,1471,1496]" name="Taiwan" pageId="7" pageNumber="456">Taiwan</collectingCountry>
|
||
,
|
||
<date id="4E4CABAFFFB5FF9B3576FA6BFA259A8B" box="[1280,1431,1471,1496]" pageId="7" pageNumber="456" value="2015-03-11">
|
||
<collectingDate id="5E085247FFB5FF9B3576FA6BFA259A8B" box="[1280,1431,1471,1496]" pageId="7" pageNumber="456" value="2015-03-11">11 Mar. 2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="8A9A8732FFB5FF9B30E1FA4FFB019AAE" ID-GBIF-Occurrence="1838430526" box="[151,1203,1508,1533]" collectingDate="2015-02-11" collectionCode="NMMB-P" country="Taiwan" location="Kaohsiung" pageId="7" pageNumber="456" specimenCode="NMMB-P22245" specimenCount="1" specimenCount-female="1" stateProvince="Kaohsiung">
|
||
<specimenCode id="6A542514FFB5FF9B30E1FA4FFEE39AAE" box="[151,337,1508,1533]" collectionCode="NMMB-P" pageId="7" pageNumber="456">NMMB-P22245</specimenCode>
|
||
,
|
||
<specimenCount id="2CF446E6FFB5FF9B3114FA4FFE649AAE" box="[354,470,1508,1533]" pageId="7" pageNumber="456" type="female">1 females</specimenCount>
|
||
,
|
||
<quantity id="FD0A208AFFB5FF9B3190FA4FFDFD9AAF" box="[486,591,1508,1532]" metricMagnitude="-2" metricUnit="m" metricValue="9.17" pageId="7" pageNumber="456" unit="mm" value="91.7">91.7 mm</quantity>
|
||
SL, Ke-tzu-liao,
|
||
<collectingRegion id="F836438DFFB5FF9B3357FA4FFC2F9AAE" box="[801,925,1508,1533]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="456">Kaohsiung</collectingRegion>
|
||
,
|
||
<collectingCountry id="42E5CDFFFFB5FF9B33D8FA4FFBB19AAE" box="[942,1027,1508,1533]" name="Taiwan" pageId="7" pageNumber="456">Taiwan</collectingCountry>
|
||
,
|
||
<date id="4E4CABAFFFB5FF9B3465FA4FFB019AAE" box="[1043,1203,1508,1533]" pageId="7" pageNumber="456" value="2015-02-11">
|
||
<collectingDate id="5E085247FFB5FF9B3465FA4FFB019AAE" box="[1043,1203,1508,1533]" pageId="7" pageNumber="456" value="2015-02-11">11 Feb. 2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="8A9A8732FFB5FF9B34B5FA4FFC589973" ID-GBIF-Occurrence="1838430522" collectingDate="2015-11-01" collectionCode="NMMB-P" country="Taiwan" elevation="50" location="Chie-din" pageId="7" pageNumber="456" specimenCode="NMMB-P22770" specimenCount="3" specimenCount-male="3" stateProvince="Kaohsiung">
|
||
<specimenCode id="6A542514FFB5FF9B34B5FA4FFACC9AAF" box="[1219,1406,1508,1532]" collectionCode="NMMB-P" pageId="7" pageNumber="456">NMMB-P22770</specimenCode>
|
||
,
|
||
<specimenCount id="2CF446E6FFB5FF9B35F8FA4FFF699973" pageId="7" pageNumber="456" type="male">3 males</specimenCount>
|
||
,
|
||
<quantity id="FD0A208AFFB5FF9B3090F9A3FE3B994C" box="[230,393,1544,1568]" metricMagnitude="-2" metricUnit="m" metricValue="8.78" metricValueMax="9.23" metricValueMin="8.33" pageId="7" pageNumber="456" unit="mm" value="87.8" valueMax="92.3" valueMin="83.3">83.3–92.3 mm</quantity>
|
||
SL,
|
||
<location id="3F2DDBB4FFB5FF9B31C8F9ACFD919973" LSID="urn:lsid:plazi:treatment:B25B3C79FFB5FF9630E1FB07FE059DFB:3F2DDBB4FFB5FF9B31C8F9ACFD919973" box="[446,547,1543,1568]" country="Taiwan" name="Chie-din" pageId="7" pageNumber="456" stateProvince="Kaohsiung">Chie-din</location>
|
||
,
|
||
<collectingRegion id="F836438DFFB5FF9B3259F9ACFD199973" box="[559,683,1543,1568]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="456">Kaohsiung</collectingRegion>
|
||
,
|
||
<collectingCountry id="42E5CDFFFFB5FF9B32C0F9ACFCB99973" box="[694,779,1543,1568]" name="Taiwan" pageId="7" pageNumber="456">Taiwan</collectingCountry>
|
||
,
|
||
<quantity id="FD0A208AFFB5FF9B3360F9ACFCE3994C" box="[790,849,1543,1568]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="456" unit="m" value="50.0">
|
||
<elevation id="B1DF6A5CFFB5FF9B3360F9ACFCE3994C" box="[790,849,1543,1568]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="456" unit="m" value="50.0">50 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="4E4CABAFFFB5FF9B332BF9A3FC589973" box="[861,1002,1543,1568]" pageId="7" pageNumber="456" value="2015-11-01">
|
||
<collectingDate id="5E085247FFB5FF9B332BF9A3FC589973" box="[861,1002,1543,1568]" pageId="7" pageNumber="456" value="2015-11-01">1 Nov. 2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="72E8DEE4FFB5FF9B30B1F987FDC399DF" pageId="7" pageNumber="456" type="diagnosis">
|
||
<paragraph id="3A4D8D6FFFB5FF9B30B1F987FDC399DF" blockId="7.[151,1436,1399,2037]" pageId="7" pageNumber="456">
|
||
<emphasis id="0886517DFFB5FF9B30B1F987FEF39916" bold="true" box="[199,321,1580,1605]" pageId="7" pageNumber="456">Diagnosis.</emphasis>
|
||
Caudal fin with a pair of black blotches, arranged between the second and fourth fin rays from the uppermost and lowermost fin rays in this fin; ocular-side pectoral fin with greatly elongate filament, longer in males than females; gill rakers not serrate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="72E8DEE4FFB5FF9630B1F93CFE089E83" lastPageId="10" lastPageNumber="459" pageId="7" pageNumber="456" type="description">
|
||
<paragraph id="3A4D8D6FFFB5FF9B30B1F93CFC8E99AB" blockId="7.[151,1436,1399,2037]" pageId="7" pageNumber="456">
|
||
<emphasis id="0886517DFFB5FF9B30B1F93CFEEB99E3" bold="true" box="[199,345,1687,1712]" pageId="7" pageNumber="456">Description.</emphasis>
|
||
Dorsal-fin rays 89–98, anal-fin rays 68–75, ocular-side pectoral-fin rays10–12, blind-side pectoral-fin rays 9–10, caudal-fin rays 3 + 11 + 3, ocular-side pelvic-fin rays 6, blind-side pelvic-fin rays 6, scales
|
||
<quantity id="FD0A208AFFB5FF9B30E1F974FF4899AB" box="[151,250,1759,1784]" metricMagnitude="0" metricUnit="m" metricValue="1.2065" metricValueMax="1.27" metricValueMin="1.143" pageId="7" pageNumber="456" unit="in" value="47.5" valueMax="50.0" valueMin="45.0">45–50 in</quantity>
|
||
lateral line, gill rakers 0 + 6, vertebrae 10 + 25–26.
|
||
</paragraph>
|
||
<paragraph id="3A4D8D6FFFB5FF9B30B1F8AEFE1D98A6" blockId="7.[151,1436,1399,2037]" pageId="7" pageNumber="456">
|
||
In SL: HL 3.75–4.36, body depth 1.83–2.03. In HL: snout 4.67–5.25, upper eye diameter 3.27–3.92, lower eye diameter 3.26–3.87, interorbital width
|
||
<quantity id="FD0A208AFFB5FF9B323CF88CFD6A9813" box="[586,728,1831,1856]" metricMagnitude="-1" metricUnit="m" metricValue="1.19507" metricValueMax="1.43764" metricValueMin="0.9525" pageId="7" pageNumber="456" unit="in" value="4.705" valueMax="5.66" valueMin="3.75">3.75–5.66 in</quantity>
|
||
males,
|
||
<quantity id="FD0A208AFFB5FF9B3346F88CFC0F9813" box="[816,957,1831,1856]" metricMagnitude="-1" metricUnit="m" metricValue="2.00025" metricValueMax="2.22504" metricValueMin="1.77546" pageId="7" pageNumber="456" unit="in" value="7.875" valueMax="8.76" valueMin="6.99">6.99–8.76 in</quantity>
|
||
females, ocular-side upper jaw 2.94–3.18, blind-side upper jaw 2.90–3.18, ocular-side lower jaw 2.11–2.38, blind-side lower jaw 1.96–2.13, caudal peduncle depth 1.82–2.22, ocular-side pectoral fin
|
||
<quantity id="FD0A208AFFB5FF9B3219F8DBFD4D98DB" box="[623,767,1903,1928]" metricMagnitude="-2" metricUnit="m" metricValue="1.524" metricValueMax="1.8288" metricValueMin="1.2192" pageId="7" pageNumber="456" unit="in" value="0.6" valueMax="0.72" valueMin="0.48">0.48–0.72 in</quantity>
|
||
males,
|
||
<quantity id="FD0A208AFFB5FF9B332FF8C4FC5A98DB" box="[857,1000,1903,1928]" metricMagnitude="-2" metricUnit="m" metricValue="2.6162" metricValueMax="2.8194" metricValueMin="2.413" pageId="7" pageNumber="456" unit="in" value="1.03" valueMax="1.11" valueMin="0.95">0.95–1.11 in</quantity>
|
||
females, blind-side pectoral fin 1.70– 1.94, ocular-side pelvic fin 1.75–2.33, blind-side pelvic fin 2.01–2.85, base of ocular-side pelvic fin 2.16–2.59, base of blind-side pelvic fin 5.08–7.86, longest dorsal fin ray 1.73–2.28, longest anal fin ray 1.71–2.07, middle caudal fin ray 1.01–1.16.
|
||
</paragraph>
|
||
<caption id="6E8DDDE7FFBAFF9430E1F980FEDF9933" httpUri="https://zenodo.org/record/1227799/files/figure.png" pageId="8" pageNumber="457" startId="8.[151,250,1579,1601]" targetBox="[151,1436,193,1557]" targetPageId="8">
|
||
<paragraph id="3A4D8D6FFFBAFF9430E1F980FEDF9933" blockId="8.[151,1436,1579,1632]" pageId="8" pageNumber="457">
|
||
<emphasis id="0886517DFFBAFF9430E1F980FEBF9912" bold="true" box="[151,269,1579,1601]" pageId="8" pageNumber="457">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="FDF2F6ECFFBAFF94316CF987FD649912" authority="Amaoka, 1963" authorityName="Amaoka" authorityYear="1963" box="[282,726,1579,1601]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="8" pageNumber="457" phylum="Chordata" rank="species" species="multisquama">
|
||
<emphasis id="0886517DFFBAFF94316CF987FD849912" box="[282,566,1579,1601]" italics="true" pageId="8" pageNumber="457">Engyprosopon multisquama</emphasis>
|
||
Amaoka, 1963
|
||
</taxonomicName>
|
||
. Ocular side, NMMB-P8852, male, 89.5 mm SL. B. NMMB-P14467, female, 75.3 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3A4D8D6FFFBAFF9430B1F92CFEC398B7" blockId="8.[151,1436,1671,2020]" pageId="8" pageNumber="457">
|
||
Body rather narrowly ovate, deepest about at middle of body, body depth usually less than 1/2 of SL; dorsal and ventral contours of body almost symmetrical. Caudal peduncle about 1/4 of body depth. Head small, its length about 1/4 of SL; upper profile with distinct concavity anterodorsal to upper margin of lower eye, profile steep in mature males, not so in females and juveniles. Snout short, much shorter than eye diameter. Ocular-side rostral spine on snout tip in males, absent or feeble in females and juveniles. Eyes small, eye diameter distinctly less than upper jaw length; lower eye slightly in advance of upper eye.
|
||
<collectingRegion id="F836438DFFBAFF943306F897FC209800" box="[880,914,1852,1875]" country="Maldives" name="Uthuru" pageId="8" pageNumber="457">No</collectingRegion>
|
||
orbital spine in either sex. Interorbital space moderately concave, distinct sexual and ontogenetic differences in its width; interorbital width greater in males than in females. Ocular-side nostrils anterior to upper margin of lower eye; anterior nostril tubular with wide flap; posterior nostril a short tube; blind-side nostrils very small and similar to those on ocular side, closely set below origin of dorsal fin.
|
||
</paragraph>
|
||
<caption id="6E8DDDE7FFBBFF9530E1F981FB759913" box="[151,1223,1578,1600]" httpUri="https://zenodo.org/record/1227801/files/figure.png" pageId="9" pageNumber="458" startId="9.[151,250,1578,1600]" targetBox="[151,1436,193,1556]" targetPageId="9">
|
||
<paragraph id="3A4D8D6FFFBBFF9530E1F981FB759913" blockId="9.[151,1223,1578,1600]" box="[151,1223,1578,1600]" pageId="9" pageNumber="458">
|
||
<emphasis id="0886517DFFBBFF9530E1F981FEBF9913" bold="true" box="[151,269,1578,1600]" pageId="9" pageNumber="458">FIGURE 5</emphasis>
|
||
.
|
||
<taxonomicName id="FDF2F6ECFFBBFF95316FF980FD619913" authority="Amaoka, 1963" authorityName="Amaoka" authorityYear="1963" box="[281,723,1578,1600]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="9" pageNumber="458" phylum="Chordata" rank="species" species="multisquama">
|
||
<emphasis id="0886517DFFBBFF95316FF980FD879913" box="[281,565,1578,1600]" italics="true" pageId="9" pageNumber="458">Engyprosopon multisquama</emphasis>
|
||
Amaoka, 1963
|
||
</taxonomicName>
|
||
. Blind side, same specimens as those in Figure 4.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3A4D8D6FFFBBFF9530B1F9C7FD32980E" blockId="9.[151,1437,1644,2029]" pageId="9" pageNumber="458">Mouth small, oblique; maxilla extending ventroposteriorly slightly beyond vertical through anterior margin of lower eye; anterior tips of both jaws about on same vertical line, when mouth closed; small knob on mandibular symphysis. Teeth in upper jaw biserial, those in outer series larger and more widely spaced than those in inner series; some teeth anteriorly canine-like; lower jaw teeth uniserial, conical, almost same size as upper inner teeth. Gill rakers on first arch moderately slender, sharp at tip, inside edge not serrate; no gill rakers on upper limb. Ocular-side scales ctenoid with short ctenii, large and deciduous; tips of both jaws and tip of ocular side of snout naked; scales cycloid on blind side of body.</paragraph>
|
||
<paragraph id="3A4D8D6FFFBBFF9630B1F8C3FEC29F86" blockId="9.[151,1437,1644,2029]" lastBlockId="10.[151,1437,151,680]" lastPageId="10" lastPageNumber="459" pageId="9" pageNumber="458">Dorsal and anal fins without elongate rays. Ocular-side pectoral fin with elongate filamentous rays in both sexes; rays in males longer than those in females; blind-side pectoral fin not elongate; same length in both sexes. Ocular-side pelvic-fin origin at tip of isthmus, fifth ray of ocular-side fin opposite to first ray of blind-side fin; anterior margin of first ray of ocular-side pelvic fin with skin flap. Tip of isthmus near vertical line through middle of lower eye. Caudal-fin rays branched except for three upper- and three lowermost rays. Tip of pelvic bone not distinctly extruded.</paragraph>
|
||
<paragraph id="3A4D8D6FFFB8FF9630B1FF4BFC6E9E36" blockId="10.[151,1437,151,680]" pageId="10" pageNumber="459">
|
||
<emphasis id="0886517DFFB8FF9630B1FF4BFE739FAB" box="[199,449,224,248]" italics="true" pageId="10" pageNumber="459">Coloration in alcohol</emphasis>
|
||
. Ocular side of body uniformly light yellowish green without any distinct markings; caudal fin with pair of large black blotches arranged between second to fourth fin rays from upper- and lowermost fin rays, respectively; all fins with irregularly-scattered, small black spots; blind side of body dark, except for part of head in males; blind side of body uniformly yellowish white in females.
|
||
</paragraph>
|
||
<paragraph id="3A4D8D6FFFB8FF9630B1FEC4FE089E83" blockId="10.[151,1437,151,680]" pageId="10" pageNumber="459">
|
||
<emphasis id="0886517DFFB8FF9630B1FEC4FE0B9EDB" bold="true" box="[199,441,367,392]" pageId="10" pageNumber="459">Sexual dimorphism.</emphasis>
|
||
This species shows sexual dimorphism in slope of the anterior margin of the head, in interorbital width, presence or absence of a rostral spine, length of the ocular-side pectoral fin, and in coloration on the blind side of the body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="72E8DEE4FFB8FF9630B1FE77FBE19D4B" pageId="10" pageNumber="459" type="distribution">
|
||
<paragraph id="3A4D8D6FFFB8FF9630B1FE77FBE19D4B" blockId="10.[151,1437,151,680]" pageId="10" pageNumber="459">
|
||
<emphasis id="0886517DFFB8FF9630B1FE77FED29EA6" bold="true" box="[199,352,476,501]" pageId="10" pageNumber="459">Distribution.</emphasis>
|
||
<collectingRegion id="F836438DFFB8FF96311FFE76FE769EA7" box="[361,452,477,500]" country="Sri Lanka" name="Western" pageId="10" pageNumber="459">Western</collectingRegion>
|
||
Pacific Ocean off
|
||
<collectingCountry id="42E5CDFFFFB8FF9632EAFE77FD5C9EA6" box="[668,750,476,501]" name="Taiwan" pageId="10" pageNumber="459">Taiwan</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="42E5CDFFFFB8FF9633E4FE76FC679EA7" box="[914,981,477,500]" name="Japan" pageId="10" pageNumber="459">Japan</collectingCountry>
|
||
; also found in mouth of Mekong River,
|
||
<collectingCountry id="42E5CDFFFFB8FF9630E1FE54FF449D4B" box="[151,246,511,536]" name="Vietnam" pageId="10" pageNumber="459">Vietnam</collectingCountry>
|
||
(KA, pers. data). Commonly collected at depths less than
|
||
<quantity id="FD0A208AFFB8FF9633F2FDABFC799D44" box="[900,971,512,536]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="459" unit="m" value="100.0">100 m</quantity>
|
||
off
|
||
<collectingCountry id="42E5CDFFFFB8FF96338FFE54FBFD9D4B" box="[1017,1103,511,536]" name="Taiwan" pageId="10" pageNumber="459">Taiwan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="72E8DEE4FFB8FF9630B1FD8FFE059DFB" pageId="10" pageNumber="459" type="discussion">
|
||
<paragraph id="3A4D8D6FFFB8FF9630B1FD8FFE059DFB" blockId="10.[151,1437,151,680]" pageId="10" pageNumber="459">
|
||
<emphasis id="0886517DFFB8FF9630B1FD8FFE899D6E" bold="true" box="[199,315,548,573]" pageId="10" pageNumber="459">Remarks.</emphasis>
|
||
This species has frequently been confused with
|
||
<taxonomicName id="FDF2F6ECFFB8FF96332AFD8EFBAC9D6F" baseAuthorityName="Temminck & Schlegel" baseAuthorityYear="1846" box="[860,1054,548,572]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="10" pageNumber="459" phylum="Chordata" rank="species" species="grandisquama">
|
||
<emphasis id="0886517DFFB8FF96332AFD8EFBAC9D6F" box="[860,1054,548,572]" italics="true" pageId="10" pageNumber="459">E. grandisquama</emphasis>
|
||
</taxonomicName>
|
||
in having a pair of black blotches on the caudal fin. However,
|
||
<taxonomicName id="FDF2F6ECFFB8FF9631A4FDE2FD339D33" authorityName="Amaoka" authorityYear="1963" box="[466,641,584,608]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="10" pageNumber="459" phylum="Chordata" rank="species" species="multisquama">
|
||
<emphasis id="0886517DFFB8FF9631A4FDE2FD339D33" box="[466,641,584,608]" italics="true" pageId="10" pageNumber="459">E. multisquama</emphasis>
|
||
</taxonomicName>
|
||
differs from the latter in having larger black blotches on the caudal fin, a more elongate body, a more elongate ocular-side pectoral fin, greater number of scales in the lateral line, and by its higher meristic values.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |