160 lines
21 KiB
XML
160 lines
21 KiB
XML
<document id="7605723E480C0441581EF24EE0FF800F" ID-CLB-Dataset="30201" ID-DOI="10.11646/zootaxa.4413.3.2" ID-GBIF-Dataset="786fe950-08b0-4072-bf66-7f64702748df" ID-ISSN="1175-5326" ID-Zenodo-Dep="1227791" ID-ZooBank="4CBD6FA5-1E36-4857-8992-2DCBA902EE13" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524558778976" checkinUser="plazi" docAuthor="Amaoka, Kunio & Ho, Hsuan-Ching" docDate="2018" docId="B25B3C79FFB3FF9D30E1FCEBFDFE984B" docLanguage="en" docName="zootaxa.4413.3.2.pdf.imf" docOrigin="Zootaxa 4413 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Engyprosopon Gunther 1862" docType="treatment" docVersion="7" lastPageNumber="450" masterDocId="4E624401FFB2FF9C3076FFABFFB29F53" masterDocTitle="Review of the genus Engyprosopon Günther, 1862 (Pleuronectiformes: Bothidae) from waters off Taiwan, with descriptions of two new species" masterLastPageNumber="481" masterPageNumber="449" pageNumber="450" updateTime="1698540829403" updateUser="ExternalLinkService">
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<mods:title id="B360EF13CD4C16F6ABB8D9E3AB63B83B">Review of the genus Engyprosopon Günther, 1862 (Pleuronectiformes: Bothidae) from waters off Taiwan, with descriptions of two new species</mods:title>
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<mods:namePart id="F18BE5CA6F36FC6CC92FD0ADCC669DE4">Amaoka, Kunio</mods:namePart>
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<mods:namePart id="D50F83E2EBADA0609E2B49F6F521A1F9">Ho, Hsuan-Ching</mods:namePart>
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<mods:title id="3C17D4AD3DB8DE2B909AF6FA6BABFBF0">Zootaxa</mods:title>
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<mods:date id="97D90FEBCCCC29B283A951A7A87A85B4">2018</mods:date>
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<heading id="61053A03FFB3FF9D30E1FCEBFDEA9C09" bold="true" box="[151,600,831,858]" fontSize="11" level="1" pageId="1" pageNumber="450" reason="1">
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<emphasis id="0886517DFFB3FF9D30E1FCEBFDEA9C09" bold="true" box="[151,600,831,858]" pageId="1" pageNumber="450">
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Genus
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<taxonomicName id="FDF2F6ECFFB3FF9D3098FC94FDEA9C09" ID-CoL="62RVT" ID-ENA="195606" authority="Gunther, 1862" authorityName="Gunther" authorityYear="1862" box="[238,600,831,858]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="genus">
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<emphasis id="0886517DFFB3FF9D3098FC94FE2C9C0A" bold="true" box="[238,414,831,857]" italics="true" pageId="1" pageNumber="450">Engyprosopon</emphasis>
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<bibRefCitation id="5E63F09EFFB3FF9D31D3FCEBFDEA9C09" author="Gunther, A." bookContentInfo="534 pp." box="[421,600,832,858]" journalOrPublisher="British Museum of Natural History London" pageId="1" pageNumber="450" refId="ref14800" refString="Gunther, A. (1862) Catalogue of the fishes in the British Museum. vol. 4. British Museum of Natural History London, 534 pp." title="Catalogue of the fishes in the British Museum. vol. 4" type="book" year="1862">Günther, 1862</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="72E8DEE4FFB3FF9D30E1FC23FB779CE8" pageId="1" pageNumber="450" type="reference_group">
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<paragraph id="3A4D8D6FFFB3FF9D30E1FC23FC639CCF" blockId="1.[151,1221,902,955]" box="[151,977,902,925]" pageId="1" pageNumber="450">
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<treatmentCitationGroup id="1AE2AA41FFB3FF9D30E1FC23FC639CCF" box="[151,977,902,925]" pageId="1" pageNumber="450">
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<taxonomicName id="FDF2F6ECFFB3FF9D30E1FC23FE5F9CCF" ID-CoL="62RVT" authority="Gunther, 1862: 431" authorityName="Gunther" authorityPageNumber="431" authorityYear="1862" box="[151,493,902,925]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="genus">
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<emphasis id="0886517DFFB3FF9D30E1FC23FE989CCE" box="[151,298,904,925]" italics="true" pageId="1" pageNumber="450">Engyprosopon</emphasis>
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<treatmentCitation id="BB53AB7EFFB3FF9D3159FC2DFE5F9CCF" author="" box="[303,493,902,924]" page="431" pageId="1" pageNumber="450">
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<bibRefCitation id="5E63F09EFFB3FF9D3159FC2DFE719CCF" author="Gunther, A." bookContentInfo="534 pp." box="[303,451,902,924]" journalOrPublisher="British Museum of Natural History London" pageId="1" pageNumber="450" refId="ref14800" refString="Gunther, A. (1862) Catalogue of the fishes in the British Museum. vol. 4. British Museum of Natural History London, 534 pp." title="Catalogue of the fishes in the British Museum. vol. 4" type="book" year="1862">Günther, 1862</bibRefCitation>
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:431
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</treatmentCitation>
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</taxonomicName>
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(type species:
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<taxonomicName id="FDF2F6ECFFB3FF9D32F3FC2DFC749CCF" authority="Bleeker, 1854" authorityName="Bleeker" authorityYear="1854" box="[645,966,902,924]" class="Actinopterygii" family="Scophthalmidae" genus="Rhombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="species" species="mogkii">
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<emphasis id="0886517DFFB3FF9D32F3FC2DFC9D9CCF" box="[645,815,902,924]" italics="true" pageId="1" pageNumber="450">Rhombus mogkii</emphasis>
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<bibRefCitation id="5E63F09EFFB3FF9D3342FC2DFC749CCF" author="Bleeker, P." box="[820,966,902,924]" journalOrPublisher="Natuurkundig Tijdschrift voor Nederlandsch Indie" pageId="1" pageNumber="450" pagination="225 - 260" part="7" refId="ref14432" refString="Bleeker, P. (1854) Vijfde bijdrage tot de kennis der ichthyologische fauna van Celebes. Natuurkundig Tijdschrift voor Nederlandsch Indie, 7 (2), 225 - 260." title="Vijfde bijdrage tot de kennis der ichthyologische fauna van Celebes" type="journal article" year="1854">Bleeker, 1854</bibRefCitation>
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</taxonomicName>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="3A4D8D6FFFB3FF9D30E1FC0DFB779CE8" blockId="1.[151,1221,902,955]" box="[151,1221,933,955]" pageId="1" pageNumber="450">
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<treatmentCitationGroup id="1AE2AA41FFB3FF9D30E1FC0DFB779CE8" box="[151,1221,933,955]" pageId="1" pageNumber="450">
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<taxonomicName id="FDF2F6ECFFB3FF9D30E1FC0DFF589CE8" authorityName="Jordan & Starks" authorityYear="1904" box="[151,234,934,955]" class="Actinopterygii" family="Bothidae" genus="Scaeops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="genus">
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<emphasis id="0886517DFFB3FF9D30E1FC0DFF589CE8" box="[151,234,934,955]" italics="true" pageId="1" pageNumber="450">Scaeops</emphasis>
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</taxonomicName>
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<treatmentCitation id="BB53AB7EFFB3FF9D3086FC0EFE4D9CE8" author="" box="[240,511,933,955]" page="627" pageId="1" pageNumber="450">
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<bibRefCitation id="5E63F09EFFB3FF9D3086FC0EFE669CE8" author="Jordan D. S. & Starks, E. C." box="[240,468,933,955]" journalOrPublisher="Bulletin of the United States Fish Commission" pageId="1" pageNumber="450" pagination="577 - 628" part="22" refId="ref15318" refString="Jordan D. S. & Starks, E. C. (1904) List of fishes dredged by the steamer " Albatross " off the coast of Japan in the summer of 1900, with descriptions of new species and a review of the Japanese Macrouridae. Bulletin of the United States Fish Commission, 22, 577 - 628." title="List of fishes dredged by the steamer " Albatross " off the coast of Japan in the summer of 1900, with descriptions of new species and a review of the Japanese Macrouridae" type="journal article" year="1904">Jordan & Starks, 1904</bibRefCitation>
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:627
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</treatmentCitation>
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(type species:
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<taxonomicName id="FDF2F6ECFFB3FF9D32E0FC0EFB089CE8" authority="Temminck & Schlegel, 1846" authorityName="Temminck & Schlegel" authorityYear="1846" box="[662,1210,933,955]" class="Actinopterygii" family="Scophthalmidae" genus="Rhombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="species" species="grandisquama">
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<emphasis id="0886517DFFB3FF9D32E0FC0EFC3C9CE8" box="[662,910,933,955]" italics="true" pageId="1" pageNumber="450">Rhombus grandisquama</emphasis>
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<bibRefCitation id="5E63F09EFFB3FF9D33E5FC0EFB089CE8" author="Temminck, C. J. & Schlegel, H." bookContentInfo="325 pp." box="[915,1210,933,955]" editor="von Siebold, P. F." journalOrPublisher="Lugduni-Batavorum, Leiden" pageId="1" pageNumber="450" refId="ref16125" refString="Temminck, C. J. & Schlegel, H. (1846) Pisces. In: von Siebold, P. F. (Ed.), Fauna Japonica sive descriptio animalium, quae in itinere per Japoniam jussu et auspiciis superiorum, qui summum in Indian Batavia Imperium tenent suscepto annis 1823 - 1850 collegit, notis, observationibus et ad umbra. Lugduni-Batavorum, Leiden, 325 pp." title="Pisces" type="book chapter" volumeTitle="Fauna Japonica sive descriptio animalium, quae in itinere per Japoniam jussu et auspiciis superiorum, qui summum in Indian Batavia Imperium tenent suscepto annis 1823 - 1850 collegit, notis, observationibus et ad umbra." year="1846">Temminck & Schlegel, 1846</bibRefCitation>
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</taxonomicName>
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).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="72E8DEE4FFB3FF9D30E1FC4CFEFE9AE3" pageId="1" pageNumber="450" type="diagnosis">
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<paragraph id="3A4D8D6FFFB3FF9D30E1FC4CFB5A9BE6" blockId="1.[151,1437,999,1816]" pageId="1" pageNumber="450">
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<emphasis id="0886517DFFB3FF9D30E1FC4CFEA39B53" bold="true" box="[151,273,999,1024]" pageId="1" pageNumber="450">Diagnosis.</emphasis>
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Body shallow to deeply ovate. Anterior tip of isthmus ventral to middle of lower eye. A strong, or feeble (rarely absent) rostral spine in males; rostral spine feeble or absent in females and juveniles of both sexes. Anterior margin of head steeper in mature males than in females and juveniles, or rarely with the same profile in both sexes. Interorbital space wide to narrow, concave (to almost ridge-like), becoming wider with growth, and wider in males than in females and juveniles. Mouth usually moderate or sometime large in size, upper jaw length on ocular side 2.0–
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<quantity id="FD0A208AFFB3FF9D311AFB37FE1E9BE6" box="[364,428,1180,1205]" metricMagnitude="-2" metricUnit="m" metricValue="9.144" pageId="1" pageNumber="450" unit="in" value="3.6">3.6 in</quantity>
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HL. Teeth on upper jaw uniserial or biserial; teeth uniserial on lower jaw.
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</paragraph>
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<paragraph id="3A4D8D6FFFB3FF9D30B1FB14FEFE9AE3" blockId="1.[151,1437,999,1816]" pageId="1" pageNumber="450">Scales ctenoid on ocular side with row of feeble ctenii along apical margin; scales cycloid on blind side; 36–61 scales in lateral line. Lateral line on ocular side curved above pectoral fin, absent on blind side. Dorsal-fin origin on blind side anterior to dorsal margin of lower eye. Anal-fin origin below anterior base of ocular-side pectoral fin. Pelvic fin on ocular side originating at tip of isthmus, that on blind side equal to third to fifth fin ray on ocular side. Anus on blind side, just anterior to origin of anal fin; urogenital pore on ocular side, opposite to vent. Caudal skeleton of four plates including a parhypural and three hypurals, all with deep clefts from their distal margins to near their bases.</paragraph>
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</subSubSection>
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<subSubSection id="72E8DEE4FFB3FF9D30B1FA17FDFE984B" pageId="1" pageNumber="450" type="discussion">
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<paragraph id="3A4D8D6FFFB3FF9D30B1FA17FDFE984B" blockId="1.[151,1437,999,1816]" pageId="1" pageNumber="450">
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<emphasis id="0886517DFFB3FF9D30B1FA17FE899A86" bold="true" box="[199,315,1468,1493]" pageId="1" pageNumber="450">Remarks.</emphasis>
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<taxonomicName id="FDF2F6ECFFB3FF9D3134FA16FE569A87" authorityName="Gunther" authorityYear="1862" box="[322,484,1469,1492]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="genus">
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<emphasis id="0886517DFFB3FF9D3134FA16FE569A87" box="[322,484,1469,1492]" italics="true" pageId="1" pageNumber="450">Engyprosopon</emphasis>
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</taxonomicName>
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shares many characters with
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<taxonomicName id="FDF2F6ECFFB3FF9D3346FA17FC689A87" baseAuthorityName="Hensley" baseAuthorityYear="2005" box="[816,986,1468,1492]" class="Actinopterygii" family="Bothidae" genus="Asterorhombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="genus">
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<emphasis id="0886517DFFB3FF9D3346FA17FC689A87" box="[816,986,1468,1492]" italics="true" pageId="1" pageNumber="450">Asterorhombus</emphasis>
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</taxonomicName>
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including the cleft parhypural and three hypurals. Members of
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<taxonomicName id="FDF2F6ECFFB3FF9D31E3FA4AFD8A9AAB" authorityName="Gunther" authorityYear="1862" box="[405,568,1505,1528]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="genus">
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<emphasis id="0886517DFFB3FF9D31E3FA4AFD8A9AAB" box="[405,568,1505,1528]" italics="true" pageId="1" pageNumber="450">Engyprosopon</emphasis>
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</taxonomicName>
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have been distinguished from specimens of the latter genus by having sexual differences in interorbital width, length of the ocular-side pectoral fin, presence of a rostral spine, and sometimes an orbital spine and orbital flap, and by differences in body coloration on the blind side (
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<bibRefCitation id="5E63F09EFFB3FF9D34E8F98CFA3D9913" author="Amaoka, K. & Mihara, E. & Rivaton, J." box="[1182,1423,1575,1600]" editor="Crosnier, A." journalOrPublisher="Memoires du Museum National d'histoire naturelle" pageId="1" pageNumber="450" pagination="377 - 426" part="158" refId="ref14327" refString="Amaoka, K., Mihara, E. & Rivaton, J. (1993) Pisces, Pleuronectiformes: flatfishes from the waters around New Caledonia. A revision of the genus Engyprosopon. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM, vol 11. Memoires du Museum National d'histoire naturelle, 158, 377 - 426." title="Pisces, Pleuronectiformes: flatfishes from the waters around New Caledonia. A revision of the genus Engyprosopon" type="journal article" volumeTitle="Resultats des Campagnes MUSORSTOM, vol 11" year="1993">
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||
Amaoka
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<emphasis id="0886517DFFB3FF9D357EF982FA8E9913" box="[1288,1340,1576,1600]" italics="true" pageId="1" pageNumber="450">et al</emphasis>
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., 1993
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</bibRefCitation>
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||
). However,
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<bibRefCitation id="5E63F09EFFB3FF9D317BF9E7FE0E9936" author="Hensley, D. A." box="[269,444,1612,1637]" journalOrPublisher="Copeia, 2005" pageId="1" pageNumber="450" pagination="445 – 460" part="3" publicationUrl="https://doi.org/10.1643/CI-03-186R3" refId="ref14956" refString="Hensley, D. A. (2005) Revision of the genus Asterorhombu s (Pleuronectiformes: Bothidae). Copeia, 2005 (3), 445 - 460. https: // doi. org / 10.1643 / CI- 03 - 186 R 3" title="Revision of the genus Asterorhombu s (Pleuronectiformes: Bothidae)" type="journal article" year="2005">Hensley (2005)</bibRefCitation>
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found that sexes of
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<taxonomicName id="FDF2F6ECFFB3FF9D32DCF9E7FC299937" authorityName="Hensley & Randall" authorityYear="2003" box="[682,923,1612,1636]" class="Actinopterygii" family="Bothidae" genus="Asterorhombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="species" species="filifer">
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<emphasis id="0886517DFFB3FF9D32DCF9E7FC299937" box="[682,923,1612,1636]" italics="true" pageId="1" pageNumber="450">Asterorhombus filifer</emphasis>
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</taxonomicName>
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differ in interorbital width. In this paper, no sexual differences in the interorbital width and the coloration on the blind side were found for
|
||
<taxonomicName id="FDF2F6ECFFB3FF9D348CF9DAFF5199FF" baseAuthorityName="Bleeker" baseAuthorityYear="1854" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="species" species="mogkii">
|
||
<emphasis id="0886517DFFB3FF9D348CF9DAFF5199FF" italics="true" pageId="1" pageNumber="450">Engyprosopon mogkii</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="FDF2F6ECFFB3FF9D317BF93EFE6699FF" authority="Amaoka & Ho, 2018" authorityName="Amaoka & Ho" authorityYear="2018" box="[269,468,1684,1708]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="species" species="parvipectorale" status="sp. nov.">
|
||
<emphasis id="0886517DFFB3FF9D317BF93EFE6699FF" box="[269,468,1684,1708]" italics="true" pageId="1" pageNumber="450">E. parvipectorale</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0886517DFFB3FF9D31A8F93FFD8899FE" bold="true" box="[478,570,1684,1709]" pageId="1" pageNumber="450">
|
||
<taxonomicNameLabel id="13B5EC06FFB3FF9D31A8F93FFD8899FE" box="[478,570,1684,1709]" pageId="1" pageNumber="450" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Based on these results, these two genera cannot be definitely separated by presence or absence of these sexual characters. Therefore,
|
||
<taxonomicName id="FDF2F6ECFFB3FF9D332AF912FC4C9983" authorityName="Gunther" authorityYear="1862" box="[860,1022,1721,1744]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="genus">
|
||
<emphasis id="0886517DFFB3FF9D332AF912FC4C9983" box="[860,1022,1721,1744]" italics="true" pageId="1" pageNumber="450">Engyprosopon</emphasis>
|
||
</taxonomicName>
|
||
can be clearly distinguished from
|
||
<taxonomicName id="FDF2F6ECFFB3FF9D30E1F977FEF299A7" baseAuthorityName="Hensley" baseAuthorityYear="2005" box="[151,320,1756,1780]" class="Actinopterygii" family="Bothidae" genus="Asterorhombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="genus">
|
||
<emphasis id="0886517DFFB3FF9D30E1F977FEF299A7" box="[151,320,1756,1780]" italics="true" pageId="1" pageNumber="450">Asterorhombus</emphasis>
|
||
</taxonomicName>
|
||
only in lacking a lure on the first dorsal-fin ray and in lacking palmate gill rakers, both features that characterize species of
|
||
<taxonomicName id="FDF2F6ECFFB3FF9D31EDF8ABFDF7984B" baseAuthorityName="Hensley" baseAuthorityYear="2005" box="[411,581,1792,1816]" class="Actinopterygii" family="Bothidae" genus="Asterorhombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="1" pageNumber="450" phylum="Chordata" rank="genus">
|
||
<emphasis id="0886517DFFB3FF9D31EDF8ABFDF7984B" box="[411,581,1792,1816]" italics="true" pageId="1" pageNumber="450">Asterorhombus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |