478 lines
52 KiB
XML
478 lines
52 KiB
XML
<document id="BA7AE2B63D8BA3FAF40EAC661F152659" ID-DOI="10.5281/zenodo.10831085" ID-ISSN="1942-1354" ID-Zenodo-Dep="10831085" ID-ZooBank="C1D5B819-A964-4679-B090-84CDBBC59D6A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688389961601" checkinUser="felipe" docAuthor="Otto, Robert L., Muona, Jyrki & Córdoba-Alfaro, Jim" docDate="2023" docId="954287CDB71CFFA269F0FB65471E5888" docLanguage="en" docName="InsectaMundi.2023.0991.1-36.pdf" docOrigin="Insecta Mundi 2023 (991)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Isarthrus striatus Otto, Muona and Cordoba-Alfaro 2023, new species" docType="treatment" docVersion="4" lastPageNumber="20" masterDocId="697BFFB5B70FFFB76930FFD643695A4D" masterDocTitle="A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama" masterLastPageNumber="36" masterPageNumber="1" pageNumber="18" updateTime="1710774778166" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="257FA08DF7B5B5DFCCA694453CCF329F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DF6988CB1F3C1D170B5F5D7F3611E838">
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<mods:title id="C2961CE317AACD61F622896B25ABEB33">A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama</mods:title>
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</mods:titleInfo>
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<mods:name id="01ACF97449E66C051AE6BF287A2B4848" type="personal">
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<mods:role id="E08A404955BADE0936F3AE50FB1345A4">
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<mods:roleTerm id="109BA0669E53D681A6C3F927612F5A11">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DF27C9DF4C9DB0D18667C6BB0B3839FC">Otto, Robert L.</mods:namePart>
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<mods:affiliation id="C8A34A8380DEC3FE126D80999A209F05">W 4806 Chrissie Circle, Shawano, WI 54166, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:name id="B02EA7B392FE256409AE4AFF8AA68C24" type="personal">
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<mods:role id="EEA608E0EF428270C87FF9BC341E558D">
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<mods:roleTerm id="A915ED8453805237962D9F1EEE80A0D9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="625B3893EF0E20F725C455A58430E72A">Muona, Jyrki</mods:namePart>
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<mods:affiliation id="884D52C52DE420FB9B8390577E14C03C">Finnish Museum of Natural History, Zoology Unit PO Box 17, University of Helsinki, Finland FIN- 00014</mods:affiliation>
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</mods:name>
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<mods:name id="D0BB4425C5C47D53808410EE4AE762AE" type="personal">
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<mods:roleTerm id="0F69F3C0AB96E132E51EB857DAF62F85">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="109B1A3D07E5A4282D2FD3FE8DD7E6C6">Córdoba-Alfaro, Jim</mods:namePart>
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<mods:affiliation id="6A22610FE9C5CC5560DE185E41A06710">Insectopia Insect Museum, BioSur Foundation Puerto Jiménez, Península de Osa, Costa Rica jim. cordoba @ biosurcorcovado. org</mods:affiliation>
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<mods:typeOfResource id="3824D49E8F021C99B73E37B3315C081A">text</mods:typeOfResource>
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<mods:title id="E7E98C33595C643D848CA50CB4924057">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="95CD79CFCF6E3AC7C72D9E0EF73DACB0">
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<mods:date id="5A19FFA3FB06B18E690552EFDAB70C0E">2023</mods:date>
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<mods:detail id="BD4CB76EFE072CE1BDBDC8EA9E3648AD" type="pubDate">
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<mods:number id="CF95CDA993D1E544674A97BDF95523B4">2023-05-26</mods:number>
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<mods:detail id="76C5663882E8267FE3073624A5387633" type="volume">
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<mods:number id="F8B3913CA77EF4553D42C5B9296E59B8">2023</mods:number>
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<mods:detail id="86294556034F141D808E9E145F5F937B" type="issue">
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<mods:number id="1ED773D78DA04BED0E692A6610FB4257">991</mods:number>
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<mods:start id="0BFBE74ADD7701604E87F6C8349B0385">1</mods:start>
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<mods:classification id="2C3FAB39CD7616E502A0FA98532732EB">journal article</mods:classification>
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<mods:identifier id="48CBEE3AC5798CF0110F8BFF813DFB48" type="DOI">10.5281/zenodo.10831085</mods:identifier>
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<mods:identifier id="727F7350B769B230713BD7720C2A6EFC" type="ISSN">1942-1354</mods:identifier>
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<treatment id="954287CDB71CFFA269F0FB65471E5888" ID-DOI="http://doi.org/10.5281/zenodo.8109095" ID-Zenodo-Dep="8109095" LSID="urn:lsid:plazi:treatment:954287CDB71CFFA269F0FB65471E5888" httpUri="http://treatment.plazi.org/id/954287CDB71CFFA269F0FB65471E5888" lastPageId="21" lastPageNumber="20" pageId="19" pageNumber="18">
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<subSubSection id="55F16550B71CFFA469F0FB65409D5E9C" box="[192,1012,1203,1233]" pageId="19" pageNumber="18" type="nomenclature">
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<paragraph id="1D5436DBB71CFFA469F0FB65409D5E9C" blockId="19.[192,1012,1203,1267]" box="[192,1012,1203,1233]" pageId="19" pageNumber="18">
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<heading id="461C81B7B71CFFA469F0FB65409D5E9C" box="[192,1012,1203,1233]" fontSize="12" level="2" pageId="19" pageNumber="18" reason="2">
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<emphasis id="2F9FEAC9B71CFFA469F0FB65409D5E9C" bold="true" box="[192,1012,1203,1233]" pageId="19" pageNumber="18">
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<taxonomicName id="DAEB4D58B71CFFA469F0FB6540385E9D" authority="Otto, Muona and Cordoba-Alfaro" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,849,1203,1233]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
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<emphasis id="2F9FEAC9B71CFFA469F0FB6542F35E9D" bold="true" box="[192,410,1203,1232]" italics="true" pageId="19" pageNumber="18">Isarthrus striatus</emphasis>
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Otto, Muona and Córdoba-Alfaro
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="34AC57B2B71CFFA46A6FFB62409D5E9C" box="[863,1012,1204,1233]" pageId="19" pageNumber="18" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="55F16550B71CFFA469F0FB0E42345EBE" box="[192,349,1240,1267]" pageId="19" pageNumber="18" type="description">
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<paragraph id="1D5436DBB71CFFA469F0FB0E42345EBE" blockId="19.[192,1012,1203,1267]" box="[192,349,1240,1267]" pageId="19" pageNumber="18">
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<figureCitation id="85D02A5EB71CFFA469F0FB0E42345EBE" box="[192,349,1240,1267]" captionStart="Figures 35–39" captionStartId="20.[193,280,1172,1198]" captionTargetBox="[455,1178,213,1142]" captionTargetId="figure-344@20.[455,1178,213,1142]" captionTargetPageId="20" captionText="Figures 35–39.Costa Rican macraulacine eucnemids. 35) Isarthrus striatus sp. nov., dorsal habitus. 36) Isarthrus striatus sp. nov., ventral habitus. 37) Absensiugum brunneum sp. nov., male holotype, dorsal habitus. 38) Absensiugum brunneum sp. nov., male holotype, ventral habitus. 39) Absensiugum brunneum sp. nov., Male aedeagus, dorsal view. (Scale: 35–36 = 5.0 mm; 37–39 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831101" httpUri="https://zenodo.org/record/10831101/files/figure.png" pageId="19" pageNumber="18">Figures 35–36</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="55F16550B71CFFA469F0FAD740A05F8B" pageId="19" pageNumber="18" type="diagnosis">
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||
<paragraph id="1D5436DBB71CFFA469F0FAD740A05F8B" blockId="19.[192,1440,1281,1478]" pageId="19" pageNumber="18">
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<emphasis id="2F9FEAC9B71CFFA469F0FAD742555F56" bold="true" box="[192,316,1281,1307]" pageId="19" pageNumber="18">Diagnosis.</emphasis>
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The new eucnemid species can be readily distinguished from any macraulacine members of
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<taxonomicName id="DAEB4D58B71CFFA46C03FAD4438E5F70" authorityName="Eschscholtz" authorityYear="1829" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="family">Eucnemidae</taxonomicName>
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present in
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<collectingCountry id="65FC764BB71CFFA4685BFAF2428C5F70" box="[363,485,1315,1342]" name="Costa Rica" pageId="19" pageNumber="18">Costa Rica</collectingCountry>
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by the presence of an interantennal carina, a wide lateral antennal groove which is widest in the middle, simple tarsal claws and dark colored dorsum with reddish-brown antennae and legs. The new species differs from the Nearctic
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<taxonomicName id="DAEB4D58B71CFFA46B5CFAB140F75FCF" authority="LeConte" authorityName="LeConte" box="[620,926,1383,1410]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="calceatus">
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<emphasis id="2F9FEAC9B71CFFA46B5CFAB140505FCC" box="[620,825,1383,1409]" italics="true" pageId="19" pageNumber="18">Isarthrus calceatus</emphasis>
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LeConte
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</taxonomicName>
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by its antennal grooves widest in the middle, striate elytra and shorter flagellomeres II–III.
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<taxonomicName id="DAEB4D58B71CFFA46BFFFA5F40F75FEE" box="[719,926,1417,1443]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="calceatus">
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||
<emphasis id="2F9FEAC9B71CFFA46BFFFA5F40F75FEE" box="[719,926,1417,1443]" italics="true" pageId="19" pageNumber="18">Isarthrus calceatus</emphasis>
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</taxonomicName>
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have parallel-sided antennal grooves, poorly developed striae on the elytra, and slightly longer flagellomeres II–III.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="55F16550B71CFFA369F0FA0342A65FEB" lastPageId="20" lastPageNumber="19" pageId="19" pageNumber="18" type="materials_examined">
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<paragraph id="1D5436DBB71CFFA469F0FA03412C5C18" blockId="19.[191,1442,1493,1927]" pageId="19" pageNumber="18">
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<emphasis id="2F9FEAC9B71CFFA469F0FA0342AB5FA2" bold="true" box="[192,450,1493,1519]" pageId="19" pageNumber="18">Type material. Female</emphasis>
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<materialsCitation id="AD833C86B71CFFA468F8FA03412C5C18" collectingDate="2004-02-24" collectionCode="R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Est. Bio. La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="19" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Heredia" typeStatus="holotype">
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<emphasis id="2F9FEAC9B71CFFA468F8FA0341465FA2" bold="true" box="[456,559,1493,1519]" pageId="19" pageNumber="18">
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<typeStatus id="C2508879B71CFFA468F8FA0341465FA2" box="[456,559,1493,1519]" pageId="19" pageNumber="18" type="holotype">holotype</typeStatus>
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||
</emphasis>
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||
: “
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||
<collectingCountry id="65FC764BB71CFFA46B75FA0341815FA2" box="[581,744,1493,1519]" name="Costa Rica" pageId="19" pageNumber="18">COSTA RICA</collectingCountry>
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||
: Prov.
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||
<collectingRegion id="DF2FF839B71CFFA46A00FA0340E55FA2" box="[816,908,1493,1519]" country="Costa Rica" name="Heredia" pageId="19" pageNumber="18">Heredia</collectingRegion>
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;
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<location id="18346000B71CFFA46AA6FA0347275FA2" LSID="urn:lsid:plazi:treatment:954287CDB71CFFA269F0FB65471E5888:18346000B71CFFA46AA6FA0347275FA2" box="[918,1102,1493,1520]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="Est. Bio. La Selva" pageId="19" pageNumber="18" stateProvince="Heredia">Est.Bio. La Selva</location>
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, 50–;
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<quantity id="DA139B3EB71CFFA46DBFFA0347BD5FBD" box="[1167,1236,1493,1520]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="18" unit="m" value="150.0">
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<elevation id="96C6D1E8B71CFFA46DBFFA0347BD5FBD" box="[1167,1236,1493,1520]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="18" unit="m" value="150.0">150 m</elevation>
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</quantity>
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<geoCoordinate id="78DF501CB71CFFA46DEAFA03465C5FBD" box="[1242,1333,1493,1520]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="19" pageNumber="18" precision="925" value="10.433333">10°26′N</geoCoordinate>
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<geoCoordinate id="78DF501CB71CFFA46C0BFA0346F45FBD" box="[1339,1437,1493,1520]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="19" pageNumber="18" precision="925" value="-84.01667">84°01′W</geoCoordinate>
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;
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<date id="6955101BB71CFFA469F0FA21422F5C5C" box="[192,326,1527,1553]" pageId="19" pageNumber="18" value="2004-02-24">
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<collectingDate id="7911E9F3B71CFFA469F0FA21422F5C5C" box="[192,326,1527,1553]" pageId="19" pageNumber="18" value="2004-02-24">24 Feb 2004</collectingDate>
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</date>
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||
; INBio-OET-ALAS transect” (“24 Feb” handwritten on label) / “
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<date id="6955101BB71CFFA46D36FA2147D25C5C" box="[1030,1211,1527,1553]" pageId="19" pageNumber="18" value="2004-02-24">
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||
<collectingDate id="7911E9F3B71CFFA46D36FA2147D25C5C" box="[1030,1211,1527,1553]" pageId="19" pageNumber="18" value="2004-02-24">24 Febrero 2004</collectingDate>
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||
</date>
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||
; Malaise; M/26/768” / “
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||
<emphasis id="2F9FEAC9B71CFFA469EBF9CF421E5C7E" bold="true" box="[219,375,1561,1587]" pageId="19" pageNumber="18">
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<typeStatus id="C2508879B71CFFA469EBF9CF421B5C7E" box="[219,370,1561,1587]" pageId="19" pageNumber="18" type="holotype">HOLOTYPE</typeStatus>
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:
|
||
</emphasis>
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||
;
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||
<taxonomicName id="DAEB4D58B71CFFA468B3F9CF428F5C7E" authorityName="LeConte" authorityYear="1852" box="[387,486,1561,1587]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
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||
<emphasis id="2F9FEAC9B71CFFA468B3F9CF428F5C7E" box="[387,486,1561,1587]" italics="true" pageId="19" pageNumber="18">Isarthrus</emphasis>
|
||
</taxonomicName>
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||
;
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||
<taxonomicName id="DAEB4D58B71CFFA468C3F9CC412C5C7E" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[499,581,1562,1587]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="2F9FEAC9B71CFFA468C3F9CC412C5C7E" box="[499,581,1562,1587]" italics="true" pageId="19" pageNumber="18">striatus</emphasis>
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||
</taxonomicName>
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||
<specimenCount id="0BEDFD52B71CFFA46B7BF9CA41095C7F" box="[587,608,1564,1586]" pageId="19" pageNumber="18" type="female">♀</specimenCount>
|
||
; Otto, Muona & Córdoba-Alfaro; Det.
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||
<collectionCode id="7BFAAE1EB71CFFA46D23F9CF474E5C7E" box="[1043,1063,1561,1587]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="18">R</collectionCode>
|
||
.
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||
<collectorName id="B01E530DB71CFFA46D1AF9CF47155C7E" box="[1066,1148,1561,1587]" pageId="19" pageNumber="18">L. Otto</collectorName>
|
||
; 2019” (red printed label).
|
||
<typeStatus id="C2508879B71CFFA469F0F9ED42415C18" box="[192,296,1595,1621]" pageId="19" pageNumber="18" type="holotype">Holotype</typeStatus>
|
||
to be deposited in
|
||
<collectionCode id="7BFAAE1EB71CFFA468CCF9ED41285C18" box="[508,577,1595,1621]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="19" pageNumber="18">INBC</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="1D5436DBB71CFFA36830F98B42A65FEB" blockId="19.[191,1442,1493,1927]" lastBlockId="20.[192,1440,1352,1446]" lastPageId="20" lastPageNumber="19" pageId="19" pageNumber="18">
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||
<materialsCitation id="AD833C86B71CFFA46830F98B40995CF6" collectingDate="1993-02" collectingDateMax="1993-04-31" collectingDateMin="1993-02" collectionCode="R, INBC" collectorName="P. Hanson & L. Otto" country="Costa Rica" county="Pr" latitude="10.433333" location="Pto. Viejo" longLatPrecision="1298" longitude="-84.01667" municipality="La Selva Bio. Sta." pageId="19" pageNumber="18" specimenCount="6" specimenCount-female="6" stateProvince="Heredia" typeStatus="paratype">
|
||
<emphasis id="2F9FEAC9B71CFFA46830F98B42135C3A" bold="true" box="[256,378,1629,1655]" pageId="19" pageNumber="18">
|
||
<typeStatus id="C2508879B71CFFA46830F98B421C5C3A" box="[256,373,1629,1655]" pageId="19" pageNumber="18" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="0BEDFD52B71CFFA468B1F98B42A95C3B" box="[385,448,1629,1655]" pageId="19" pageNumber="18" type="female">4 ♀♀</specimenCount>
|
||
:
|
||
<emphasis id="2F9FEAC9B71CFFA468FEF98A41135C3B" bold="true" box="[462,634,1628,1654]" pageId="19" pageNumber="18">
|
||
<collectingCountry id="65FC764BB71CFFA468FEF98A411C5C3B" box="[462,629,1628,1654]" name="Costa Rica" pageId="19" pageNumber="18">COSTA RICA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="0BEDFD52B71CFFA46BB2F98B41C55C3B" box="[642,684,1629,1655]" pageId="19" pageNumber="18" type="female">1 ♀</specimenCount>
|
||
, “COSTA RICA:
|
||
<collectingRegion id="DF2FF839B71CFFA46A42F98B40A75C3A" box="[882,974,1629,1655]" country="Costa Rica" name="Heredia" pageId="19" pageNumber="18">Heredia</collectingRegion>
|
||
;
|
||
<collectingCounty id="F4354E57B71CFFA46AE9F98B409F5C3A" box="[985,1014,1629,1655]" pageId="19" pageNumber="18">Pr</collectingCounty>
|
||
:
|
||
<collectingMunicipality id="FD30ACA1B71CFFA46D32F98B47AC5C3A" box="[1026,1221,1629,1655]" pageId="19" pageNumber="18">La Selva Bio. Sta.</collectingMunicipality>
|
||
;
|
||
<quantity id="DA139B3EB71CFFA46DE0F98B46625C3A" box="[1232,1291,1629,1655]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="18" unit="km" value="3.0">3 km</quantity>
|
||
s.
|
||
<location id="18346000B71CFFA46C1BF98B46F55C3A" LSID="urn:lsid:plazi:treatment:954287CDB71CFFA269F0FB65471E5888:18346000B71CFFA46C1BF98B46F55C3A" box="[1323,1436,1629,1655]" country="Costa Rica" county="Pr" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="La Selva Bio. Sta." name="Pto. Viejo" pageId="19" pageNumber="18" stateProvince="Heredia">Pto. Viejo</location>
|
||
;
|
||
<geoCoordinate id="78DF501CB71CFFA469F1F9A942735CD4" box="[193,282,1663,1689]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="19" pageNumber="18" precision="925" value="10.433333">10°26′N</geoCoordinate>
|
||
<geoCoordinate id="78DF501CB71CFFA46814F9A942EC5CD4" box="[292,389,1663,1689]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="19" pageNumber="18" precision="925" value="-84.01667">84°01′W</geoCoordinate>
|
||
” / “
|
||
<date id="6955101BB71CFFA46885F9A9414C5CD4" box="[437,549,1663,1689]" pageId="19" pageNumber="18" value="1993-02" valueMax="1993-04-31" valueMin="1993-02-00">
|
||
<collectingDate id="7911E9F3B71CFFA46885F9A9414C5CD4" box="[437,549,1663,1689]" pageId="19" pageNumber="18" value="1993-02" valueMax="1993-04-31" valueMin="1993-02-00">ii–iv.1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="B01E530DB71CFFA46B02F9A941C15CD4" box="[562,680,1663,1689]" pageId="19" pageNumber="18">P. Hanson</collectorName>
|
||
,
|
||
<collectingMethod id="C4AA4ECCB71CFFA46B85F9A940215CD4" box="[693,840,1663,1689]" pageId="19" pageNumber="18">Malaise Trap</collectingMethod>
|
||
” / “
|
||
<emphasis id="2F9FEAC9B71CFFA46A48F9A847635CD5" bold="true" box="[888,1034,1662,1688]" pageId="19" pageNumber="18">
|
||
<typeStatus id="C2508879B71CFFA46A48F9A8476F5CD5" box="[888,1030,1662,1688]" pageId="19" pageNumber="18" type="paratype">PARATYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
;
|
||
<taxonomicName id="DAEB4D58B71CFFA46D2AF9A947145CD4" authorityName="LeConte" authorityYear="1852" box="[1050,1149,1663,1689]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2F9FEAC9B71CFFA46D2AF9A947145CD4" box="[1050,1149,1663,1689]" italics="true" pageId="19" pageNumber="18">Isarthrus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="DAEB4D58B71CFFA46DBDF9A947B65CD5" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[1165,1247,1663,1688]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="2F9FEAC9B71CFFA46DBDF9A947B65CD5" box="[1165,1247,1663,1688]" italics="true" pageId="19" pageNumber="18">striatus</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="0BEDFD52B71CFFA46DD8F95447945CD5" box="[1256,1277,1666,1688]" pageId="19" pageNumber="18" type="female">♀</specimenCount>
|
||
; Otto, Muona & Córdoba-Alfaro; Det.
|
||
<collectionCode id="7BFAAE1EB71CFFA468E8F97742855CF6" box="[472,492,1697,1723]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="18">R</collectionCode>
|
||
.
|
||
<collectorName id="B01E530DB71CFFA468C0F977412C5CF6" box="[496,581,1697,1723]" pageId="19" pageNumber="18">L. Otto</collectorName>
|
||
; 2019” (yellow printed label) (
|
||
<collectionCode id="7BFAAE1EB71CFFA46A90F97740815CF6" box="[928,1000,1697,1723]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="19" pageNumber="18">INBC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="AD833C86B71CFFA46ACDF977460C5CB2" collectingDate="1993-04" collectingDateMax="1993-05-31" collectingDateMin="1993-04" collectionCode="R, INBC" collectorName="P. Hanson & L. Otto" country="Costa Rica" county="Pr" latitude="10.433333" location="Pto. Viejo" longLatPrecision="1298" longitude="-84.01667" municipality="La Selva Bio. Sta." pageId="19" pageNumber="18" specimenCount="2" specimenCount-female="2" stateProvince="Heredia" typeStatus="paratype">
|
||
<specimenCount id="0BEDFD52B71CFFA46ACDF97747415CF7" box="[1021,1064,1697,1723]" pageId="19" pageNumber="18" type="female">1 ♀</specimenCount>
|
||
, “COSTA RICA:
|
||
<collectingRegion id="DF2FF839B71CFFA46DC2F97646275CF7" box="[1266,1358,1696,1722]" country="Costa Rica" name="Heredia" pageId="19" pageNumber="18">Heredia</collectingRegion>
|
||
;
|
||
<collectingCounty id="F4354E57B71CFFA46C6AF977461E5CF6" box="[1370,1399,1697,1723]" pageId="19" pageNumber="18">Pr</collectingCounty>
|
||
:
|
||
<collectingMunicipality id="FD30ACA1B71CFFA46CB4F977420A5C90" pageId="19" pageNumber="18">La Selva Bio. Sta.</collectingMunicipality>
|
||
;
|
||
<quantity id="DA139B3EB71CFFA46840F91542C55C91" box="[368,428,1730,1757]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="18" unit="km" value="3.0">3 km</quantity>
|
||
s.
|
||
<location id="18346000B71CFFA468FFF915412A5C91" LSID="urn:lsid:plazi:treatment:954287CDB71CFFA269F0FB65471E5888:18346000B71CFFA468FFF915412A5C91" box="[463,579,1730,1757]" country="Costa Rica" county="Pr" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="La Selva Bio. Sta." name="Pto. Viejo" pageId="19" pageNumber="18" stateProvince="Heredia">Pto. Viejo</location>
|
||
;
|
||
<geoCoordinate id="78DF501CB71CFFA46B60F91441C35C90" box="[592,682,1730,1757]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="19" pageNumber="18" precision="925" value="10.433333">10°26′N</geoCoordinate>
|
||
<geoCoordinate id="78DF501CB71CFFA46B83F915407C5C90" box="[691,789,1731,1757]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="19" pageNumber="18" precision="925" value="-84.01667">84°01′W</geoCoordinate>
|
||
” / “
|
||
<date id="6955101BB71CFFA46A75F91440DA5C91" box="[837,947,1730,1756]" pageId="19" pageNumber="18" value="1993-04" valueMax="1993-05-31" valueMin="1993-04-00">
|
||
<collectingDate id="7911E9F3B71CFFA46A75F91440DA5C91" box="[837,947,1730,1756]" pageId="19" pageNumber="18" value="1993-04" valueMax="1993-05-31" valueMin="1993-04-00">iv–v.1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="B01E530DB71CFFA46AF0F915475F5C90" box="[960,1078,1731,1757]" pageId="19" pageNumber="18">P. Hanson</collectorName>
|
||
,
|
||
<collectingMethod id="C4AA4ECCB71CFFA46D73F91447BE5C90" box="[1091,1239,1730,1757]" pageId="19" pageNumber="18">Malaise Trap</collectingMethod>
|
||
” / “
|
||
<emphasis id="2F9FEAC9B71CFFA46C37F91446F05C91" bold="true" box="[1287,1433,1730,1756]" pageId="19" pageNumber="18">
|
||
<typeStatus id="C2508879B71CFFA46C37F91446FC5C91" box="[1287,1429,1730,1756]" pageId="19" pageNumber="18" type="paratype">PARATYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
;
|
||
<taxonomicName id="DAEB4D58B71CFFA469F0F932424A5CB3" authorityName="LeConte" authorityYear="1852" box="[192,291,1764,1790]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2F9FEAC9B71CFFA469F0F932424A5CB3" box="[192,291,1764,1790]" italics="true" pageId="19" pageNumber="18">Isarthrus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="DAEB4D58B71CFFA46801F93342EA5CB3" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[305,387,1765,1790]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="2F9FEAC9B71CFFA46801F93342EA5CB3" box="[305,387,1765,1790]" italics="true" pageId="19" pageNumber="18">striatus</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="0BEDFD52B71CFFA468BAF93E42F65CB3" box="[394,415,1768,1790]" pageId="19" pageNumber="18" type="female">♀</specimenCount>
|
||
; Otto, Muona & Córdoba-Alfaro; Det.
|
||
<collectionCode id="7BFAAE1EB71CFFA46A67F93340025CB2" box="[855,875,1765,1791]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="18">R</collectionCode>
|
||
.
|
||
<collectorName id="B01E530DB71CFFA46A5FF93340AB5CB2" box="[879,962,1765,1791]" pageId="19" pageNumber="18">L. Otto</collectorName>
|
||
; 2019” (yellow printed label) (
|
||
<collectionCode id="7BFAAE1EB71CFFA46C25F93346345CB2" box="[1301,1373,1765,1791]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="19" pageNumber="18">INBC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="AD833C86B71CFFA46C40F93341F05D29" collectingDate="1993-06-01" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Proy." longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" pageId="19" pageNumber="18" specimenCount="2" specimenCount-female="2" stateProvince="Heredia" typeStatus="paratype">
|
||
<specimenCount id="0BEDFD52B71CFFA46C40F93346F05CB3" box="[1392,1433,1765,1791]" pageId="19" pageNumber="18" type="female">1 ♀</specimenCount>
|
||
, “COSTA RICA:
|
||
<collectingRegion id="DF2FF839B71CFFA46846F8D042BB5D6D" box="[374,466,1798,1824]" country="Costa Rica" name="Heredia" pageId="19" pageNumber="18">Heredia</collectingRegion>
|
||
;
|
||
<collectingMunicipality id="FD30ACA1B71CFFA468ECF8D041FF5D6D" box="[476,662,1798,1825]" pageId="19" pageNumber="18">Est.Bio. La Selva</collectingMunicipality>
|
||
, 50–;
|
||
<quantity id="DA139B3EB71CFFA46BE7F8D040745D6C" box="[727,797,1798,1825]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="18" unit="m" value="150.0">
|
||
<elevation id="96C6D1E8B71CFFA46BE7F8D040745D6C" box="[727,797,1798,1825]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="18" unit="m" value="150.0">150 m</elevation>
|
||
</quantity>
|
||
<geoCoordinate id="78DF501CB71CFFA46A14F8D040165D6C" box="[804,895,1798,1825]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="19" pageNumber="18" precision="925" value="10.433333">10°26′N</geoCoordinate>
|
||
<geoCoordinate id="78DF501CB71CFFA46AB5F8D0408F5D6C" box="[901,998,1798,1825]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="19" pageNumber="18" precision="925" value="-84.01667">84°01′W</geoCoordinate>
|
||
;
|
||
<location id="18346000B71CFFA46AC1F8D047405D6D" LSID="urn:lsid:plazi:treatment:954287CDB71CFFA269F0FB65471E5888:18346000B71CFFA46AC1F8D047405D6D" box="[1009,1065,1798,1824]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" name="Proy." pageId="19" pageNumber="18" stateProvince="Heredia">Proy.</location>
|
||
<collectionCode id="7BFAAE1EB71CFFA46D18F8D047065D6D" box="[1064,1135,1798,1824]" pageId="19" pageNumber="18">ALAS</collectionCode>
|
||
, INBio-OET” / “M/04/111;
|
||
<date id="6955101BB71CFFA469F0F8FE42325D0F" box="[192,347,1832,1858]" pageId="19" pageNumber="18" value="1993-06-01">
|
||
<collectingDate id="7911E9F3B71CFFA469F0F8FE42325D0F" box="[192,347,1832,1858]" pageId="19" pageNumber="18" value="1993-06-01">01 Junio 1993</collectingDate>
|
||
</date>
|
||
; Bosque primario” / “
|
||
<emphasis id="2F9FEAC9B71CFFA46B79F8FE41B25D0F" bold="true" box="[585,731,1832,1858]" pageId="19" pageNumber="18">
|
||
<typeStatus id="C2508879B71CFFA46B79F8FE41BE5D0F" box="[585,727,1832,1858]" pageId="19" pageNumber="18" type="paratype">PARATYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
;
|
||
<taxonomicName id="DAEB4D58B71CFFA46BD7F8FE40235D0F" authorityName="LeConte" authorityYear="1852" box="[743,842,1832,1858]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2F9FEAC9B71CFFA46BD7F8FE40235D0F" box="[743,842,1832,1858]" italics="true" pageId="19" pageNumber="18">Isarthrus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="DAEB4D58B71CFFA46A67F8FF40C05D0F" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[855,937,1833,1858]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="2F9FEAC9B71CFFA46A67F8FF40C05D0F" box="[855,937,1833,1858]" italics="true" pageId="19" pageNumber="18">striatus</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="0BEDFD52B71CFFA46A9EF8FA40AA5D0F" box="[942,963,1836,1858]" pageId="19" pageNumber="18" type="female">♀</specimenCount>
|
||
; Otto, Muona & Córdoba-Alfaro; Det.
|
||
<collectionCode id="7BFAAE1EB71CFFA46C42F8FF46EF5D0E" box="[1394,1414,1833,1859]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="18">R</collectionCode>
|
||
.
|
||
<collectorName id="B01E530DB71CFFA46CBAF8FF439C5D29" pageId="19" pageNumber="18">L. Otto</collectorName>
|
||
; 2019” (yellow printed label) (
|
||
<collectionCode id="7BFAAE1EB71CFFA46B79F89C41F85D29" box="[585,657,1866,1892]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="19" pageNumber="18">INBC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="AD833C86B71CFFA36B94F89C47735F2F" collectionCode="ALAS" collectorName="Robert L. Otto" country="Costa Rica" elevation="150" lastPageId="20" lastPageNumber="19" latitude="10.433333" location="Est. Bio. La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="19" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Heredia" typeStatus="paratype">
|
||
<specimenCount id="0BEDFD52B71CFFA46B94F89C41A65D29" box="[676,719,1866,1892]" pageId="19" pageNumber="18" type="female">1 ♀</specimenCount>
|
||
, “COSTA RICA:
|
||
<collectingRegion id="DF2FF839B71CFFA46AA3F89C40865D29" box="[915,1007,1866,1892]" country="Costa Rica" name="Heredia" pageId="19" pageNumber="18">Heredia</collectingRegion>
|
||
;
|
||
<location id="18346000B71CFFA46ACAF89C47DF5D29" LSID="urn:lsid:plazi:treatment:954287CDB71CFFA269F0FB65471E5888:18346000B71CFFA46ACAF89C47DF5D29" box="[1018,1206,1866,1893]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="Est. Bio. La Selva" pageId="19" pageNumber="18" stateProvince="Heredia">Est.Bio. La Selva</location>
|
||
, 50–;
|
||
<quantity id="DA139B3EB71CFFA46DC8F89C46565D28" box="[1272,1343,1866,1893]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="18" unit="m" value="150.0">
|
||
<elevation id="96C6D1E8B71CFFA46DC8F89C46565D28" box="[1272,1343,1866,1893]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="18" unit="m" value="150.0">150 m</elevation>
|
||
</quantity>
|
||
<geoCoordinate id="78DF501CB71CFFA46C76F89C46C95D28" box="[1350,1440,1866,1893]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="19" pageNumber="18" precision="925" value="10.433333">10°26′N</geoCoordinate>
|
||
<geoCoordinate id="78DF501CB71CFFA469F0F8BA42495DCA" box="[192,288,1900,1927]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="19" pageNumber="18" precision="925" value="-84.01667">84°01′W</geoCoordinate>
|
||
; Proy.
|
||
<collectionCode id="7BFAAE1EB71CFFA46853F8BA42C35DCB" box="[355,426,1900,1926]" pageId="19" pageNumber="18">ALAS</collectionCode>
|
||
, INBio-OET” / “M/10/172; 03 Augosto 1993; Bosque primario” / “Collection of the Global; Eucnemid Research Project; (
|
||
<collectorName id="B01E530DB71BFFA36B3AFA9E41DA5F2F" box="[522,691,1352,1379]" pageId="20" pageNumber="19">Robert L. Otto</collectorName>
|
||
)” (green framed white label) / “
|
||
</materialsCitation>
|
||
<emphasis id="2F9FEAC9B71BFFA36D2BFA9E47C45F2F" bold="true" box="[1051,1197,1352,1378]" pageId="20" pageNumber="19">
|
||
<typeStatus id="C2508879B71BFFA36D2BFA9E47C05F2F" box="[1051,1193,1352,1378]" pageId="20" pageNumber="19" type="paratype">PARATYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
;
|
||
<taxonomicName id="DAEB4D58B71BFFA36D8BFA9E46775F2F" authorityName="LeConte" authorityYear="1852" box="[1211,1310,1352,1378]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2F9FEAC9B71BFFA36D8BFA9E46775F2F" box="[1211,1310,1352,1378]" italics="true" pageId="20" pageNumber="19">Isarthrus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="DAEB4D58B71BFFA36C1CFA9F46175F2F" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[1324,1406,1353,1378]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="2F9FEAC9B71BFFA36C1CFA9F46175F2F" box="[1324,1406,1353,1378]" italics="true" pageId="20" pageNumber="19">striatus</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="0BEDFD52B71BFFA36CB5FA9A46F35F2F" box="[1413,1434,1356,1378]" pageId="20" pageNumber="19" type="female">♀</specimenCount>
|
||
;
|
||
<materialsCitation id="AD833C86B71BFFA369F0FABC414F5FC9" box="[192,550,1386,1412]" collectorName="Muona & Cordoba-Alfaro" country="Costa Rica" location="Otto" pageId="20" pageNumber="19" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
|
||
<location id="18346000B71BFFA369F0FABC439C5FC9" LSID="urn:lsid:plazi:treatment:954287CDB71CFFA269F0FB65471E5888:18346000B71BFFA369F0FABC439C5FC9" box="[192,245,1386,1412]" country="Costa Rica" name="Otto" pageId="20" pageNumber="19" stateProvince="Heredia">Otto</location>
|
||
,
|
||
<collectorName id="B01E530DB71BFFA36830FABC42395FC9" box="[256,336,1386,1412]" pageId="20" pageNumber="19">Muona</collectorName>
|
||
&
|
||
<collectorName id="B01E530DB71BFFA36840FABC414F5FC9" box="[368,550,1386,1412]" pageId="20" pageNumber="19">Córdoba-Alfaro</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="AD833C86B71BFFA36B01FABC41B95FC9" box="[561,720,1386,1412]" collectionCode="R" collectorName="L. Otto" country="Costa Rica" location="Det." pageId="20" pageNumber="19" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
|
||
<location id="18346000B71BFFA36B01FABC41085FC9" LSID="urn:lsid:plazi:treatment:954287CDB71CFFA269F0FB65471E5888:18346000B71BFFA36B01FABC41085FC9" box="[561,609,1386,1412]" country="Costa Rica" name="Det." pageId="20" pageNumber="19" stateProvince="Heredia">Det.</location>
|
||
<collectionCode id="7BFAAE1EB71BFFA36B57FABC41125FC9" box="[615,635,1386,1412]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="19">R</collectionCode>
|
||
.
|
||
<collectorName id="B01E530DB71BFFA36B4EFABC41B95FC9" box="[638,720,1386,1412]" pageId="20" pageNumber="19">L. Otto</collectorName>
|
||
</materialsCitation>
|
||
; 2019” (yellow printed label) (
|
||
<collectionCode id="7BFAAE1EB71BFFA36D2FFABC47025FC9" box="[1055,1131,1386,1412]" pageId="20" pageNumber="19">GERP</collectionCode>
|
||
).
|
||
<typeStatus id="C2508879B71BFFA36D4CFABC47815FC9" box="[1148,1256,1386,1412]" pageId="20" pageNumber="19" type="paratype">Paratypes</typeStatus>
|
||
are to be deposited in
|
||
<collectionCode id="7BFAAE1EB71BFFA3683EFA5A42395FEB" box="[270,336,1420,1446]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="20" pageNumber="19">INBC</collectionCode>
|
||
and
|
||
<collectionCode id="7BFAAE1EB71BFFA368B7FA5A42A55FEB" box="[391,460,1420,1446]" pageId="20" pageNumber="19">GERP</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="49946653B71BFFA369F1FB4240745F5E" ID-DOI="http://doi.org/10.5281/zenodo.10831101" ID-Zenodo-Dep="10831101" httpUri="https://zenodo.org/record/10831101/files/figure.png" pageId="20" pageNumber="19" startId="20.[193,280,1172,1198]" targetBox="[455,1178,213,1142]" targetPageId="20" targetType="figure">
|
||
<paragraph id="1D5436DBB71BFFA369F1FB4240745F5E" blockId="20.[192,1441,1172,1300]" pageId="20" pageNumber="19">
|
||
<emphasis id="2F9FEAC9B71BFFA369F1FB4242025EE3" bold="true" box="[193,363,1172,1198]" pageId="20" pageNumber="19">Figures 35–39.</emphasis>
|
||
Costa Rican macraulacine eucnemids.
|
||
<emphasis id="2F9FEAC9B71BFFA36A2BFB4240285EE3" bold="true" box="[795,833,1172,1198]" pageId="20" pageNumber="19">35)</emphasis>
|
||
<taxonomicName id="DAEB4D58B71BFFA36A77FB4247685EE3" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[839,1025,1172,1198]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B71BFFA36A77FB4247685EE3" box="[839,1025,1172,1198]" italics="true" pageId="20" pageNumber="19">Isarthrus striatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B71BFFA36D37FB4347345EE2" bold="true" box="[1031,1117,1173,1199]" pageId="20" pageNumber="19">
|
||
<taxonomicNameLabel id="34AC57B2B71BFFA36D37FB4347345EE2" box="[1031,1117,1173,1199]" pageId="20" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, dorsal habitus.
|
||
<emphasis id="2F9FEAC9B71BFFA36C20FB42465F5EE3" bold="true" box="[1296,1334,1172,1198]" pageId="20" pageNumber="19">36)</emphasis>
|
||
<taxonomicName id="DAEB4D58B71BFFA36C0DFB4242785E9D" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B71BFFA36C0DFB4242785E9D" italics="true" pageId="20" pageNumber="19">Isarthrus striatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B71BFFA36828FB6042065E9D" bold="true" box="[280,367,1206,1232]" pageId="20" pageNumber="19">
|
||
<taxonomicNameLabel id="34AC57B2B71BFFA36828FB6042065E9D" box="[280,367,1206,1232]" pageId="20" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ventral habitus.
|
||
<emphasis id="2F9FEAC9B71BFFA36B1FFB60413C5E9D" bold="true" box="[559,597,1206,1232]" pageId="20" pageNumber="19">37)</emphasis>
|
||
<taxonomicName id="DAEB4D58B71BFFA36B6BFB60400E5E9D" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[603,871,1206,1232]" class="Insecta" family="Eucnemidae" genus="Absensiugum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="brunneum" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B71BFFA36B6BFB60400E5E9D" box="[603,871,1206,1232]" italics="true" pageId="20" pageNumber="19">Absensiugum brunneum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B71BFFA36A5EFB6040AC5E9D" bold="true" box="[878,965,1206,1232]" pageId="20" pageNumber="19">
|
||
<taxonomicNameLabel id="34AC57B2B71BFFA36A5EFB6040AC5E9D" box="[878,965,1206,1232]" pageId="20" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, dorsal habitus.
|
||
<emphasis id="2F9FEAC9B71BFFA36C18FB6046275E9D" bold="true" box="[1320,1358,1206,1232]" pageId="20" pageNumber="19">38)</emphasis>
|
||
<taxonomicName id="DAEB4D58B71BFFA36C64FB6042E05EBF" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" class="Insecta" family="Eucnemidae" genus="Absensiugum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="brunneum" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B71BFFA36C64FB6042E05EBF" italics="true" pageId="20" pageNumber="19">Absensiugum brunneum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B71BFFA368A0FB0E428E5EBF" bold="true" box="[400,487,1240,1266]" pageId="20" pageNumber="19">
|
||
<taxonomicNameLabel id="34AC57B2B71BFFA368A0FB0E428E5EBF" box="[400,487,1240,1266]" pageId="20" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, ventral habitus.
|
||
<emphasis id="2F9FEAC9B71BFFA36A63FB0140105EBC" bold="true" box="[851,889,1239,1265]" pageId="20" pageNumber="19">39)</emphasis>
|
||
<taxonomicName id="DAEB4D58B71BFFA36A4FFB0E47E25EBF" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[895,1163,1240,1266]" class="Insecta" family="Eucnemidae" genus="Absensiugum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="brunneum" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B71BFFA36A4FFB0E47E25EBF" box="[895,1163,1240,1266]" italics="true" pageId="20" pageNumber="19">Absensiugum brunneum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B71BFFA36DA2FB0E47805EBF" bold="true" box="[1170,1257,1240,1266]" pageId="20" pageNumber="19">
|
||
<taxonomicNameLabel id="34AC57B2B71BFFA36DA2FB0E47805EBF" box="[1170,1257,1240,1266]" pageId="20" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Male aedeagus, dorsal view. (Scale: 35–36 = 5.0 mm; 37–39 = 1.0 mm)
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="55F16550B71BFFA269F0FA6342B95BAE" lastPageId="21" lastPageNumber="20" pageId="20" pageNumber="19" type="description">
|
||
<paragraph id="1D5436DBB71BFFA269F0FA6341C95B19" blockId="20.[192,1443,1461,1924]" lastBlockId="21.[191,1441,214,340]" lastPageId="21" lastPageNumber="20" pageId="20" pageNumber="19">
|
||
<emphasis id="2F9FEAC9B71BFFA369F0FA6341705F82" bold="true" box="[192,537,1461,1487]" pageId="20" pageNumber="19">
|
||
Description. Female
|
||
<typeStatus id="C2508879B71BFFA36882FA6341705F82" box="[434,537,1461,1487]" pageId="20" pageNumber="19" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: Length,
|
||
<quantity id="DA139B3EB71BFFA36BB2FA6341B25F9D" box="[642,731,1461,1488]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="20" pageNumber="19" unit="mm" value="8.5">8.5 mm</quantity>
|
||
. Width,
|
||
<quantity id="DA139B3EB71BFFA36A09FA6340F85F9D" box="[825,913,1461,1488]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="19" unit="mm" value="2.5">2.5 mm</quantity>
|
||
. Body elongate; dark brownish-black; antennae and legs dark reddish-brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (
|
||
<figureCitation id="85D02A5EB71BFFA36C41FA0142635C5E" captionStart="Figures 35–39" captionStartId="20.[193,280,1172,1198]" captionTargetBox="[455,1178,213,1142]" captionTargetId="figure-344@20.[455,1178,213,1142]" captionTargetPageId="20" captionText="Figures 35–39.Costa Rican macraulacine eucnemids. 35) Isarthrus striatus sp. nov., dorsal habitus. 36) Isarthrus striatus sp. nov., ventral habitus. 37) Absensiugum brunneum sp. nov., male holotype, dorsal habitus. 38) Absensiugum brunneum sp. nov., male holotype, ventral habitus. 39) Absensiugum brunneum sp. nov., Male aedeagus, dorsal view. (Scale: 35–36 = 5.0 mm; 37–39 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831101" httpUri="https://zenodo.org/record/10831101/files/figure.png" pageId="20" pageNumber="19">Figure 35</figureCitation>
|
||
).
|
||
<emphasis id="2F9FEAC9B71BFFA3682FFA2E42355C5F" bold="true" box="[287,348,1528,1554]" pageId="20" pageNumber="19">Head</emphasis>
|
||
: Surface somewhat shiny, subspherical with crowded punctures; frons convex with slight circular impression above interantennal carina, poorly developed median carina present; interantennal carina present; apical margin of frontoclypeal region weakly trilobed, more than 2 times wider than base; mandibles stout, bidentate, densely punctate.
|
||
<emphasis id="2F9FEAC9B71BFFA368F7F98B41455C3A" bold="true" box="[455,556,1629,1655]" pageId="20" pageNumber="19">Antenna</emphasis>
|
||
: Filiform from flagellomeres I–IX, attaining about 1/3 the length of the body; flagellomere I longer than wide, longer than II; flagellomeres II–III each slightly longer than wide, sub-equal, each shorter than I and IV; flagellomeres IV–V each longer than wide, sub-equal, longer than III, shorter than VI; flagellomeres VI–VIII each longer than wide, sub-equal; flagellomere IX slightly longer than VIII.
|
||
<emphasis id="2F9FEAC9B71BFFA36C10F91446F15C91" bold="true" box="[1312,1432,1730,1756]" pageId="20" pageNumber="19">Pronotum</emphasis>
|
||
: Surface somewhat shiny, with deep, crowded punctures; quadrate, with well-developed, sharp hind angles; sides parallel-sided at basal 1/2, arcuate at apical 1/2; disc convex, slightly depressed near base above scutellar shield; base sinuous.
|
||
<emphasis id="2F9FEAC9B71BFFA36869F8F041655D0D" bold="true" box="[345,524,1830,1856]" pageId="20" pageNumber="19">Scutellar shield</emphasis>
|
||
: Quadrate, sub-triangular, shallowly punctate, setose and distally rounded.
|
||
<emphasis id="2F9FEAC9B71BFFA36C62F8F146F35D0C" bold="true" box="[1362,1434,1831,1857]" pageId="20" pageNumber="19">Elytra</emphasis>
|
||
: Very shallow striae present; interstices slightly elevated; surfaces shiny, closely punctate.
|
||
<emphasis id="2F9FEAC9B71BFFA36D80F89E478C5D2F" bold="true" box="[1200,1253,1864,1890]" pageId="20" pageNumber="19">Legs</emphasis>
|
||
: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I–III simple; metatarsomere IV excavated; metatarsomere V elongate with simple claws.
|
||
<emphasis id="2F9FEAC9B71AFFA26C63FF0046C85ABD" bold="true" box="[1363,1441,214,240]" pageId="21" pageNumber="20">Venter</emphasis>
|
||
(
|
||
<figureCitation id="85D02A5EB71AFFA269F7FF21425F5B5C" box="[199,310,247,273]" captionStart="Figures 35–39" captionStartId="20.[193,280,1172,1198]" captionTargetBox="[455,1178,213,1142]" captionTargetId="figure-344@20.[455,1178,213,1142]" captionTargetPageId="20" captionText="Figures 35–39.Costa Rican macraulacine eucnemids. 35) Isarthrus striatus sp. nov., dorsal habitus. 36) Isarthrus striatus sp. nov., ventral habitus. 37) Absensiugum brunneum sp. nov., male holotype, dorsal habitus. 38) Absensiugum brunneum sp. nov., male holotype, ventral habitus. 39) Absensiugum brunneum sp. nov., Male aedeagus, dorsal view. (Scale: 35–36 = 5.0 mm; 37–39 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831101" httpUri="https://zenodo.org/record/10831101/files/figure.png" pageId="21" pageNumber="20">Figure 36</figureCitation>
|
||
): Closely punctate, with short, recumbent silver-colored setae; hypomeron with basally opened, lateral antennal grooves, widest in the middle; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially 3.0–6.0 times wider than laterally.
|
||
</paragraph>
|
||
<paragraph id="1D5436DBB71AFFA269F0FEB242B95BAE" blockId="21.[192,1440,356,483]" pageId="21" pageNumber="20">
|
||
<emphasis id="2F9FEAC9B71AFFA269F0FEB242285B33" bold="true" box="[192,321,356,382]" pageId="21" pageNumber="20">Variations.</emphasis>
|
||
Four female
|
||
<typeStatus id="C2508879B71AFFA268E3FEB341565B32" box="[467,575,357,383]" pageId="21" pageNumber="20" type="paratype">paratypes</typeStatus>
|
||
were examined. These
|
||
<typeStatus id="C2508879B71AFFA26A71FEB340C45B32" box="[833,941,357,383]" pageId="21" pageNumber="20" type="paratype">paratypes</typeStatus>
|
||
measured 7.0–
|
||
<quantity id="DA139B3EB71AFFA26D67FEB247C55B32" box="[1111,1196,356,383]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="21" pageNumber="20" unit="mm" value="8.5">8.5 mm</quantity>
|
||
long and 2.0–
|
||
<quantity id="DA139B3EB71AFFA26C7BFEB246C95B32" box="[1355,1440,356,383]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="20" unit="mm" value="2.5">2.5 mm</quantity>
|
||
wide. One female
|
||
<typeStatus id="C2508879B71AFFA268A2FE50429A5BED" box="[402,499,390,416]" pageId="21" pageNumber="20" type="paratype">paratype</typeStatus>
|
||
is as large as and just as wide as the
|
||
<typeStatus id="C2508879B71AFFA26A95FE5047635BED" box="[933,1034,390,416]" pageId="21" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
. The remaining
|
||
<specimenCount id="0BEDFD52B71AFFA26DF7FE50461C5BED" box="[1223,1397,390,416]" pageId="21" pageNumber="20" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
|
||
are shorter and narrower than the
|
||
<typeStatus id="C2508879B71AFFA26B2BFE7141E95B8C" box="[539,640,423,449]" pageId="21" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
. Median carina on the frons are evident in
|
||
<specimenCount id="0BEDFD52B71AFFA26D55FE7E466B5B8F" box="[1125,1282,424,450]" pageId="21" pageNumber="20" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
and absent in the other
|
||
<specimenCount id="0BEDFD52B71AFFA2681DFE1F42A55BAE" box="[301,460,457,483]" pageId="21" pageNumber="20" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55F16550B71AFFA269F0FE24424B5863" pageId="21" pageNumber="20" type="distribution">
|
||
<paragraph id="1D5436DBB71AFFA269F0FE24424B5863" blockId="21.[192,1440,498,558]" pageId="21" pageNumber="20">
|
||
<emphasis id="2F9FEAC9B71AFFA269F0FE2442325841" bold="true" box="[192,347,498,524]" pageId="21" pageNumber="20">Distribution.</emphasis>
|
||
This eucnemid species is known from
|
||
<specimenCount id="0BEDFD52B71AFFA26A22FE2440DF5841" box="[786,950,498,524]" pageId="21" pageNumber="20" type="generic">five specimens</specimenCount>
|
||
taken from La Selva within the province of
|
||
<collectingRegion id="DF2FF839B71AFFA269F1FDC242745863" box="[193,285,532,558]" country="Costa Rica" name="Heredia" pageId="21" pageNumber="20">Heredia</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55F16550B71AFFA269F0FDEB41685834" pageId="21" pageNumber="20" type="biology_ecology">
|
||
<paragraph id="1D5436DBB71AFFA269F0FDEB41685834" blockId="21.[192,1440,573,634]" pageId="21" pageNumber="20">
|
||
<emphasis id="2F9FEAC9B71AFFA269F0FDEB4249581A" bold="true" box="[192,288,573,599]" pageId="21" pageNumber="20">Biology.</emphasis>
|
||
Two
|
||
<typeStatus id="C2508879B71AFFA2686DFDE842A05815" box="[349,457,574,600]" pageId="21" pageNumber="20" type="paratype">paratypes</typeStatus>
|
||
were taken from a primary forest. Three specimens were taken from a Malaise trap. Larvae and pupae are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55F16550B71AFFA269F0FD5F471E5888" pageId="21" pageNumber="20" type="etymology">
|
||
<paragraph id="1D5436DBB71AFFA269F0FD5F471E5888" blockId="21.[192,1440,649,709]" pageId="21" pageNumber="20">
|
||
<emphasis id="2F9FEAC9B71AFFA269F0FD5F422D58EE" bold="true" box="[192,324,649,675]" pageId="21" pageNumber="20">Etymology.</emphasis>
|
||
The specific epithet is derived from the presence of well-developed striae on the elytra which serves as one of the best feature to distinguish the new species from the Nearctic
|
||
<taxonomicName id="DAEB4D58B71AFFA26AC9FD7D47185889" box="[1017,1137,682,708]" class="Insecta" family="Eucnemidae" genus="Isarthrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="calceatus">
|
||
<emphasis id="2F9FEAC9B71AFFA26AC9FD7D47185889" box="[1017,1137,682,708]" italics="true" pageId="21" pageNumber="20">I. calceatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |