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<document id="53E01BF2DF0E87EC61A5E2C077BBEB38" ID-DOI="10.5281/zenodo.10831085" ID-ISSN="1942-1354" ID-Zenodo-Dep="10831085" ID-ZooBank="C1D5B819-A964-4679-B090-84CDBBC59D6A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688389961601" checkinUser="felipe" docAuthor="Otto, Robert L., Muona, Jyrki &amp; Córdoba-Alfaro, Jim" docDate="2023" docId="954287CDB706FFBD69F0FABC46E65995" docLanguage="en" docName="InsectaMundi.2023.0991.1-36.pdf" docOrigin="Insecta Mundi 2023 (991)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Adelothyreus costaricensis Otto, Muona and Cordoba-Alfaro 2023, new species" docType="treatment" docVersion="4" lastPageNumber="9" masterDocId="697BFFB5B70FFFB76930FFD643695A4D" masterDocTitle="A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama" masterLastPageNumber="36" masterPageNumber="1" pageNumber="8" updateTime="1710774778166" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="048D96E69B685B8E20C061C5299EFB27">
<mods:title id="248046471E7559ACB0BC4A9A30E1ECD5">A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama</mods:title>
</mods:titleInfo>
<mods:name id="06B3892A80A4B7A0330B5ED58DBA9FEA" type="personal">
<mods:role id="7256E0FFF83D5BC0E5A2985C5340728D">
<mods:roleTerm id="AC5A130FEA1311D9898FF056AC51E007">Author</mods:roleTerm>
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<mods:namePart id="82688165E8A42B1A8A57A0DD122D42B3">Otto, Robert L.</mods:namePart>
<mods:affiliation id="C3898C5E677004E2079F24A8FDD881FE">W 4806 Chrissie Circle, Shawano, WI 54166, U. S. A.</mods:affiliation>
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<mods:role id="F57DBAA15E637BEBEAA4F0C5E6B124C4">
<mods:roleTerm id="7B397EB2222E5FD001FE64E125B95441">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9796EA455283A9FF9AC4555CA3DA5CF">Muona, Jyrki</mods:namePart>
<mods:affiliation id="D5A735C895BA78DD803F52C4A07A9987">Finnish Museum of Natural History, Zoology Unit PO Box 17, University of Helsinki, Finland FIN- 00014</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="6BF7F1B9B8A1F96B6DF766B46418C3DB">Córdoba-Alfaro, Jim</mods:namePart>
<mods:affiliation id="3CCB66E6A3428C430A367ECC2C249DBC">Insectopia Insect Museum, BioSur Foundation Puerto Jiménez, Península de Osa, Costa Rica jim. cordoba @ biosurcorcovado. org</mods:affiliation>
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<mods:title id="D762270D0FBF9EFE3416403E9E3F6E23">Insecta Mundi</mods:title>
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<mods:part id="ECDC7F93FD695B978A8D53D8C3DD73EE">
<mods:date id="9136D59C9542D362A698531DCC8D88D1">2023</mods:date>
<mods:detail id="9F92FA489144EABCB44D79BF81E9D1BC" type="pubDate">
<mods:number id="510C769AC343439B1C1F059313D87B38">2023-05-26</mods:number>
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<mods:detail id="9CBB02877B05A4E23AD7EC72EDD1B25D" type="volume">
<mods:number id="21D58CF6572AFE1176CE14D07FFF0FD5">2023</mods:number>
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<mods:number id="00A8FD787F333CCD68E5F2C4A95CEA12">991</mods:number>
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<mods:classification id="0BAA8A81EB323A0A10DC7B658DDD2B4A">journal article</mods:classification>
<mods:identifier id="0A53EDB9595122DE673120DA51DE67FF" type="DOI">10.5281/zenodo.10831085</mods:identifier>
<mods:identifier id="72B9027CFAFCDC5C3887976B590CB119" type="ISSN">1942-1354</mods:identifier>
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<treatment id="954287CDB706FFBD69F0FABC46E65995" ID-DOI="http://doi.org/10.5281/zenodo.8109068" ID-Zenodo-Dep="8109068" LSID="urn:lsid:plazi:treatment:954287CDB706FFBD69F0FABC46E65995" httpUri="http://treatment.plazi.org/id/954287CDB706FFBD69F0FABC46E65995" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
<subSubSection id="55F16550B706FFBE69F0FABC470D5FCA" box="[192,1124,1386,1415]" pageId="9" pageNumber="8" type="nomenclature">
<paragraph id="1D5436DBB706FFBE69F0FABC470D5FCA" blockId="9.[192,1124,1386,1449]" box="[192,1124,1386,1415]" pageId="9" pageNumber="8">
<heading id="461C81B7B706FFBE69F0FABC470D5FCA" box="[192,1124,1386,1415]" fontSize="12" level="2" pageId="9" pageNumber="8" reason="2">
<emphasis id="2F9FEAC9B706FFBE69F0FABC470D5FCA" bold="true" box="[192,1124,1386,1415]" pageId="9" pageNumber="8">
<taxonomicName id="DAEB4D58B706FFBE69F0FABC40A85FCA" authority="Otto, Muona and Cordoba-Alfaro" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,961,1386,1415]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
<emphasis id="2F9FEAC9B706FFBE69F0FABC41635FCA" bold="true" box="[192,522,1386,1415]" italics="true" pageId="9" pageNumber="8">Adelothyreus costaricensis</emphasis>
Otto, Muona and Córdoba-Alfaro
</taxonomicName>
,
<taxonomicNameLabel id="34AC57B2B706FFBE6AFFFABD470D5FCA" box="[975,1124,1386,1415]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B706FFBE69F0FA5942345FE4" box="[192,349,1423,1449]" pageId="9" pageNumber="8" type="description">
<paragraph id="1D5436DBB706FFBE69F0FA5942345FE4" blockId="9.[192,1124,1386,1449]" box="[192,349,1423,1449]" pageId="9" pageNumber="8">
<figureCitation id="85D02A5EB706FFBE69F0FA5942345FE4" box="[192,349,1423,1449]" captionStart="Figures 1016" captionStartId="9.[193,280,1019,1045]" captionTargetBox="[449,1184,213,989]" captionTargetId="figure-360@9.[449,1184,213,989]" captionTargetPageId="9" captionText="Figures 1016. Costa Rican Adelothyreus species. 10) Adelothyreus brevis sp. nov., dorsal habitus. 11) Adelothyreus brevis sp. nov., pronotal lateral habitus. 12) Adelothyreus brevis sp. nov., ventral habitus. 13) Adelothyreus costaricensis sp. nov., male holotype, dorsal habitus. 14) Adelothyreus costaricensis sp. nov., male holotype, pronotal lateral habitus. 15) Adelothyreus costaricensis sp. nov., male holotype, ventral habitus. 16) Adelothyreus costaricensis sp. nov. female allotype, dorsal habitus. (Scale: 1016 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831091" httpUri="https://zenodo.org/record/10831091/files/figure.png" pageId="9" pageNumber="8">Figures 1316</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B706FFBE69F0FA6C40625FB5" pageId="9" pageNumber="8" type="diagnosis">
<paragraph id="1D5436DBB706FFBE69F0FA6C40625FB5" blockId="9.[192,1440,1466,1528]" pageId="9" pageNumber="8">
<emphasis id="2F9FEAC9B706FFBE69F0FA6C42555F99" bold="true" box="[192,316,1466,1492]" pageId="9" pageNumber="8">Diagnosis.</emphasis>
This new species is very similar to
<taxonomicName id="DAEB4D58B706FFBE6BE0FA6D40585F99" box="[720,817,1467,1493]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="comes">
<emphasis id="2F9FEAC9B706FFBE6BE0FA6D40585F99" box="[720,817,1467,1493]" italics="true" pageId="9" pageNumber="8">A. comes</emphasis>
</taxonomicName>
. The posteroventrally directed anterior lateral pronotal ridge will distinguish the new species from
<taxonomicName id="DAEB4D58B706FFBE6B93FA0B406C5FBA" box="[675,773,1501,1527]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="comes">
<emphasis id="2F9FEAC9B706FFBE6B93FA0B406C5FBA" box="[675,773,1501,1527]" italics="true" pageId="9" pageNumber="8">A. comes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B706FFBE69F0F9DE47DF5C9D" pageId="9" pageNumber="8" type="materials_examined">
<paragraph id="1D5436DBB706FFBE69F0F9DE47DF5C9D" blockId="9.[192,1441,1543,1744]" pageId="9" pageNumber="8">
<emphasis id="2F9FEAC9B706FFBE69F0F9DE42C55C6F" bold="true" box="[192,428,1543,1570]" pageId="9" pageNumber="8">Type material. Male</emphasis>
<materialsCitation id="AD833C86B706FFBE6883F9DE47DB5C9D" collectingDate="1993-06-02" collectingDateMax="1994-02-01" collectingDateMin="1993-06-02" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" county="Est. Bio. La Selva" elevation="150" latitude="10.433333" location="Est. Bio. La Selva" longLatPrecision="1298" longitude="-84.01667" municipality="Parcelas" pageId="9" pageNumber="8" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
<emphasis id="2F9FEAC9B706FFBE6883F9DE41485C6F" bold="true" box="[435,545,1544,1570]" pageId="9" pageNumber="8">
<typeStatus id="C2508879B706FFBE6883F9DE41755C6F" box="[435,540,1544,1570]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
:
</emphasis>
<collectingCountry id="65FC764BB706FFBE6B02F9DE41BE5C6F" box="[562,727,1544,1570]" name="Costa Rica" pageId="9" pageNumber="8">COSTA RICA</collectingCountry>
:
<collectingRegion id="DF2FF839B706FFBE6BD1F9DE40545C6F" box="[737,829,1544,1570]" country="Costa Rica" name="Heredia" pageId="9" pageNumber="8">Heredia</collectingRegion>
;
<collectingCounty id="F4354E57B706FFBE6A79F9DE476C5C6F" box="[841,1029,1544,1570]" pageId="9" pageNumber="8">Est.Bio. La Selva</collectingCounty>
, 50;
<quantity id="DA139B3EB706FFBE6D79F9DE47F95C6F" box="[1097,1168,1544,1571]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="8" unit="m" value="150.0">
<elevation id="96C6D1E8B706FFBE6D79F9DE47F95C6F" box="[1097,1168,1544,1571]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="8" unit="m" value="150.0">150 m</elevation>
</quantity>
<geoCoordinate id="78DF501CB706FFBE6DA8F9DE479A5C6F" box="[1176,1267,1544,1570]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="9" pageNumber="8" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB706FFBE6DCAF9DE46345C6F" box="[1274,1373,1544,1570]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="9" pageNumber="8" precision="925" value="-84.01667">84°01W</geoCoordinate>
; Proy.
<collectionCode id="7BFAAE1EB706FFBE69F0F9FD426F5C08" box="[192,262,1579,1605]" pageId="9" pageNumber="8">ALAS</collectionCode>
, INBio-OET” / “M/01/332;
<date id="6955101BB706FFBE6B09F9FD41875C08" box="[569,750,1579,1605]" pageId="9" pageNumber="8" value="1994-02-01">
<collectingDate id="7911E9F3B706FFBE6B09F9FD41875C08" box="[569,750,1579,1605]" pageId="9" pageNumber="8" value="1994-02-01">01 Febrero 1994</collectingDate>
</date>
;
<collectingMunicipality id="FD30ACA1B706FFBE6BC8F9FD403B5C08" box="[760,850,1579,1605]" pageId="9" pageNumber="8">Parcelas</collectingMunicipality>
sucesionales” / “
<emphasis id="2F9FEAC9B706FFBE6D3CF9FC47C15C09" bold="true" box="[1036,1192,1578,1604]" pageId="9" pageNumber="8">
<typeStatus id="C2508879B706FFBE6D3CF9FC47CA5C09" box="[1036,1187,1578,1604]" pageId="9" pageNumber="8" type="holotype">HOLOTYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B706FFBE6D84F9FD462B5C08" authorityName="Chevrolat" authorityYear="1868" box="[1204,1346,1579,1605]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B706FFBE6D84F9FD462B5C08" box="[1204,1346,1579,1605]" italics="true" pageId="9" pageNumber="8">Adelothyreus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B706FFBE6C7EF9FD43965C2A" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="2F9FEAC9B706FFBE6C7EF9FD43965C2A" italics="true" pageId="9" pageNumber="8">costaricensis</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B706FFBE6837F98742755C2A" box="[263,284,1617,1639]" pageId="9" pageNumber="8" type="male"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB706FFBE6BE9F99841845C25" box="[729,749,1614,1640]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="8">R</collectionCode>
.
<collectorName id="B01E530DB706FFBE6BC0F998402D5C25" box="[752,836,1614,1640]" pageId="9" pageNumber="8">L. Otto</collectorName>
; 2019” (red printed label).
<emphasis id="2F9FEAC9B706FFBE6D47F99846515C25" bold="true" box="[1143,1336,1614,1640]" pageId="9" pageNumber="8">
Female
<typeStatus id="C2508879B706FFBE6DE3F998465A5C25" box="[1235,1331,1614,1640]" pageId="9" pageNumber="8" type="allotype">allotype</typeStatus>
:
</emphasis>
<collectingCountry id="65FC764BB706FFBE6C78F998426D5CC7" name="Costa Rica" pageId="9" pageNumber="8">COSTA RICA</collectingCountry>
,
<collectingRegion id="DF2FF839B706FFBE683FF9A642025CC7" box="[271,363,1648,1674]" country="Costa Rica" name="Heredia" pageId="9" pageNumber="8">Heredia</collectingRegion>
;
<location id="18346000B706FFBE6845F9A641465CC7" LSID="urn:lsid:plazi:treatment:954287CDB706FFBD69F0FABC46E65995:18346000B706FFBE6845F9A641465CC7" box="[373,559,1648,1675]" country="Costa Rica" county="Est. Bio. La Selva" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Parcelas" name="Est. Bio. La Selva" pageId="9" pageNumber="8" stateProvince="Heredia">Est.Bio. La Selva</location>
, 50;
<quantity id="DA139B3EB706FFBE6B40F9A641D15CC6" box="[624,696,1648,1675]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="8" unit="m" value="150.0">
<elevation id="96C6D1E8B706FFBE6B40F9A641D15CC6" box="[624,696,1648,1675]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="8" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<geoCoordinate id="78DF501CB706FFBE6BF2F9A640725CC6" box="[706,795,1648,1675]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="9" pageNumber="8" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB706FFBE6A12F9A640EB5CC6" box="[802,898,1648,1675]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="9" pageNumber="8" precision="925" value="-84.01667">84°01W</geoCoordinate>
; Proy.
<collectionCode id="7BFAAE1EB706FFBE6AF5F9A647655CC7" box="[965,1036,1648,1674]" pageId="9" pageNumber="8">ALAS</collectionCode>
, INBio-OET” / “M/15/121;
<date id="6955101BB706FFBE6C72F9A643915CE0" pageId="9" pageNumber="8" value="1993-06-02">
<collectingDate id="7911E9F3B706FFBE6C72F9A643915CE0" pageId="9" pageNumber="8" value="1993-06-02">02 Junio 1993</collectingDate>
</date>
; Bosque secundario” / “
<emphasis id="2F9FEAC9B706FFBE6B38F94541F55CE0" bold="true" box="[520,668,1683,1709]" pageId="9" pageNumber="8">
<typeStatus id="C2508879B706FFBE6B38F94541F15CE0" box="[520,664,1683,1709]" pageId="9" pageNumber="8" type="allotype">ALLOTYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B706FFBE6B9AF94540515CE0" authorityName="Chevrolat" authorityYear="1868" box="[682,824,1683,1709]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B706FFBE6B9AF94540515CE0" box="[682,824,1683,1709]" italics="true" pageId="9" pageNumber="8">Adelothyreus</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B706FFBE6A77F94240A75CE0" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[839,974,1684,1709]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="2F9FEAC9B706FFBE6A77F94240A75CE0" box="[839,974,1684,1709]" italics="true" pageId="9" pageNumber="8">costaricensis</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B706FFBE6AE6F94140825CE0" box="[982,1003,1687,1709]" pageId="9" pageNumber="8" type="female"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB706FFBE69F1F96043BC5C9D" box="[193,213,1718,1744]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="8">R</collectionCode>
.
<collectorName id="B01E530DB706FFBE69E9F96042425C9D" box="[217,299,1718,1744]" pageId="9" pageNumber="8">L. Otto</collectorName>
; 2019” (yellow printed label).
<typeStatus id="C2508879B706FFBE6B49F96041885C9D" box="[633,737,1718,1744]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
and
<typeStatus id="C2508879B706FFBE6A28F96040185C9D" box="[792,881,1718,1744]" pageId="9" pageNumber="8" type="allotype">allotype</typeStatus>
are to be deposited in
<collectionCode id="7BFAAE1EB706FFBE6D5DF96047DB5C9D" box="[1133,1202,1718,1744]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="9" pageNumber="8">INBC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B706FFBD69F0F93647B45974" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="description">
<paragraph id="1D5436DBB706FFBD69F0F936402E5886" blockId="9.[191,1440,1760,1927]" lastBlockId="10.[191,1442,214,715]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
<emphasis id="2F9FEAC9B706FFBE69F0F93641695CB6" bold="true" box="[192,512,1760,1787]" pageId="9" pageNumber="8">
Description. Male
<typeStatus id="C2508879B706FFBE68A9F93741695CB6" box="[409,512,1761,1787]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
</emphasis>
: Length,
<quantity id="DA139B3EB706FFBE6B5AF93741AA5CB6" box="[618,707,1761,1788]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="8" unit="mm" value="2.5">2.5 mm</quantity>
. Width,
<quantity id="DA139B3EB706FFBE6A13F93740E05CB6" box="[803,905,1761,1788]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="8" unit="mm" value="0.75">0.75 mm</quantity>
. Body elongate; uniformly dark black; antennae dark brownish black; femur and tibiae dark brownish black; tarsi dark brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (
<figureCitation id="85D02A5EB706FFBE6B5FF8F041B45D0C" box="[623,733,1830,1857]" captionStart="Figures 1016" captionStartId="9.[193,280,1019,1045]" captionTargetBox="[449,1184,213,989]" captionTargetId="figure-360@9.[449,1184,213,989]" captionTargetPageId="9" captionText="Figures 1016. Costa Rican Adelothyreus species. 10) Adelothyreus brevis sp. nov., dorsal habitus. 11) Adelothyreus brevis sp. nov., pronotal lateral habitus. 12) Adelothyreus brevis sp. nov., ventral habitus. 13) Adelothyreus costaricensis sp. nov., male holotype, dorsal habitus. 14) Adelothyreus costaricensis sp. nov., male holotype, pronotal lateral habitus. 15) Adelothyreus costaricensis sp. nov., male holotype, ventral habitus. 16) Adelothyreus costaricensis sp. nov. female allotype, dorsal habitus. (Scale: 1016 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831091" httpUri="https://zenodo.org/record/10831091/files/figure.png" pageId="9" pageNumber="8">Figure 13</figureCitation>
).
<emphasis id="2F9FEAC9B706FFBE6BC0F8F140475D0C" bold="true" box="[752,814,1831,1857]" pageId="9" pageNumber="8">Head</emphasis>
: Surface densely punctate, somewhat dullish, subspherical; frons convex, without median fovea or carina; apical margin of frontoclypeal region rounded, more than 2.0 times wider than base; mandibles stout, bidentate, densely punctate.
<emphasis id="2F9FEAC9B706FFBE6ADEF8BA473B5DCB" bold="true" box="[1006,1106,1900,1926]" pageId="9" pageNumber="8">Antenna</emphasis>
: Pectinate from flagellomeres IIVIII, attaining about 2/3 the length of the body; flagellomere I longer than II; flagellomeres IIIII sub-equal, each shorter than IV; flagellomere IV longer than III, shorter than V; flagellomeres VVIII each sub-equal, longer than wide; flagellomere IX longer than VIII; ramus on flagellomere II arising along entire length, rami on flagellomeres IIIVIII arising at apical end, elongate and about 1.5 times longer than length of segment.
<emphasis id="2F9FEAC9B705FFBD6C13FEEA46F35B1B" bold="true" box="[1315,1434,316,342]" pageId="10" pageNumber="9">Pronotum</emphasis>
: Surface somewhat dullish, closely punctate; longer than wide, with poorly developed hind angles; lateral sides parallel-sided, apically arcuate; disc convex without median groove; anterior lateral pronotal ridge short, arcuate, directed posteroventrally; posterior lateral pronotal ridge elongate, about 3/4 the length of the pronotum (
<figureCitation id="85D02A5EB705FFBD6C43FE7742605B90" captionStart="Figures 1016" captionStartId="9.[193,280,1019,1045]" captionTargetBox="[449,1184,213,989]" captionTargetId="figure-360@9.[449,1184,213,989]" captionTargetPageId="9" captionText="Figures 1016. Costa Rican Adelothyreus species. 10) Adelothyreus brevis sp. nov., dorsal habitus. 11) Adelothyreus brevis sp. nov., pronotal lateral habitus. 12) Adelothyreus brevis sp. nov., ventral habitus. 13) Adelothyreus costaricensis sp. nov., male holotype, dorsal habitus. 14) Adelothyreus costaricensis sp. nov., male holotype, pronotal lateral habitus. 15) Adelothyreus costaricensis sp. nov., male holotype, ventral habitus. 16) Adelothyreus costaricensis sp. nov. female allotype, dorsal habitus. (Scale: 1016 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831091" httpUri="https://zenodo.org/record/10831091/files/figure.png" pageId="10" pageNumber="9">Figure 14</figureCitation>
); base sinuous, with circular strong impressions above scutellar shield.
<emphasis id="2F9FEAC9B705FFBD6D1CFE1547B65B90" bold="true" box="[1068,1247,451,477]" pageId="10" pageNumber="9">Scutellar shield</emphasis>
: Short, quadrate, sub-triangular, shallowly punctate, setose and distally rounded.
<emphasis id="2F9FEAC9B705FFBD6AB5FE3340A45BB2" bold="true" box="[901,973,485,511]" pageId="10" pageNumber="9">Elytra</emphasis>
: Very shallow striate indicated; interstices slightly elevated; surfaces shiny, very closely punctate to rugose at basal 1/2, closely punctate at apical 1/2.
<emphasis id="2F9FEAC9B705FFBD6C1BFDD14609586C" bold="true" box="[1323,1376,519,545]" pageId="10" pageNumber="9">Legs</emphasis>
: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres IIII simple; metatarsomere IV very short, excavate; metatarsomere V short with simple claws.
<emphasis id="2F9FEAC9B705FFBD683CFDBB423358CA" bold="true" box="[268,346,621,647]" pageId="10" pageNumber="9">Venter</emphasis>
(
<figureCitation id="85D02A5EB705FFBD685BFDBB42B458C5" box="[363,477,621,648]" captionStart="Figures 1016" captionStartId="9.[193,280,1019,1045]" captionTargetBox="[449,1184,213,989]" captionTargetId="figure-360@9.[449,1184,213,989]" captionTargetPageId="9" captionText="Figures 1016. Costa Rican Adelothyreus species. 10) Adelothyreus brevis sp. nov., dorsal habitus. 11) Adelothyreus brevis sp. nov., pronotal lateral habitus. 12) Adelothyreus brevis sp. nov., ventral habitus. 13) Adelothyreus costaricensis sp. nov., male holotype, dorsal habitus. 14) Adelothyreus costaricensis sp. nov., male holotype, pronotal lateral habitus. 15) Adelothyreus costaricensis sp. nov., male holotype, ventral habitus. 16) Adelothyreus costaricensis sp. nov. female allotype, dorsal habitus. (Scale: 1016 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831091" httpUri="https://zenodo.org/record/10831091/files/figure.png" pageId="10" pageNumber="9">Figure 15</figureCitation>
): Closely punctate, with short, recumbent yellowish setae; hypomeron with apically widened notosternal antennal grooves; antennal grooves smooth, sparsely punctate; metepisterna caudally wide; elytral epipleura punctate; metacoxal plates parallel-sided.
</paragraph>
<paragraph id="1D5436DBB705FFBD69F0FD0D47B45974" blockId="10.[192,1444,731,825]" pageId="10" pageNumber="9">
<emphasis id="2F9FEAC9B705FFBD69F0FD0D421358B8" bold="true" box="[192,378,731,757]" pageId="10" pageNumber="9">
Female
<typeStatus id="C2508879B705FFBD682CFD0D421358B8" box="[284,378,731,757]" pageId="10" pageNumber="9" type="allotype">allotype</typeStatus>
</emphasis>
(
<figureCitation id="85D02A5EB705FFBD68BCFD0D429458B8" box="[396,509,731,757]" captionStart="Figures 1016" captionStartId="9.[193,280,1019,1045]" captionTargetBox="[449,1184,213,989]" captionTargetId="figure-360@9.[449,1184,213,989]" captionTargetPageId="9" captionText="Figures 1016. Costa Rican Adelothyreus species. 10) Adelothyreus brevis sp. nov., dorsal habitus. 11) Adelothyreus brevis sp. nov., pronotal lateral habitus. 12) Adelothyreus brevis sp. nov., ventral habitus. 13) Adelothyreus costaricensis sp. nov., male holotype, dorsal habitus. 14) Adelothyreus costaricensis sp. nov., male holotype, pronotal lateral habitus. 15) Adelothyreus costaricensis sp. nov., male holotype, ventral habitus. 16) Adelothyreus costaricensis sp. nov. female allotype, dorsal habitus. (Scale: 1016 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831091" httpUri="https://zenodo.org/record/10831091/files/figure.png" pageId="10" pageNumber="9">Figure 16</figureCitation>
): Length, 3.0 mm. Width, 1.0 mm. Antennae serriform, about 1/2 the length of the body; flagellomere I longer than wide, longer than II; flagellomere II quadrate, serrate, shorter than III; flagellomeres IIIVIII sub-equal, transverse, asymmetrically serrate; flagellomere IX longer than VIII.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B705FFBD69F0FC9E4748592E" box="[192,1057,840,867]" pageId="10" pageNumber="9" type="distribution">
<paragraph id="1D5436DBB705FFBD69F0FC9E4748592E" blockId="10.[192,1057,840,867]" box="[192,1057,840,867]" pageId="10" pageNumber="9">
<emphasis id="2F9FEAC9B705FFBD69F0FC9E4232592F" bold="true" box="[192,347,840,866]" pageId="10" pageNumber="9">Distribution.</emphasis>
This species is only known from its
<typeStatus id="C2508879B705FFBD6BDDFC9F4074592E" box="[749,797,841,867]" pageId="10" pageNumber="9">type</typeStatus>
location in
<collectingCountry id="65FC764BB705FFBD6A92FC9F4774592E" box="[930,1053,841,867]" name="Costa Rica" pageId="10" pageNumber="9">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B705FFBD69F0FCA441A359E3" pageId="10" pageNumber="9" type="biology_ecology">
<paragraph id="1D5436DBB705FFBD69F0FCA441A359E3" blockId="10.[192,1439,882,943]" pageId="10" pageNumber="9">
<emphasis id="2F9FEAC9B705FFBD69F0FCA4424959C1" bold="true" box="[192,288,882,908]" pageId="10" pageNumber="9">Biology.</emphasis>
The
<typeStatus id="C2508879B705FFBD686CFCA442D759C1" box="[348,446,882,908]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
was taken from a successional plot of trees. The
<typeStatus id="C2508879B705FFBD6ACAFCA4473A59C1" box="[1018,1107,882,908]" pageId="10" pageNumber="9" type="allotype">allotype</typeStatus>
was taken from a secondary growth forest. Larvae and pupae are unknown.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B705FFBD69F0FC6946E65995" box="[192,1423,958,985]" pageId="10" pageNumber="9" type="etymology">
<paragraph id="1D5436DBB705FFBD69F0FC6946E65995" blockId="10.[192,1423,958,985]" box="[192,1423,958,985]" pageId="10" pageNumber="9">
<emphasis id="2F9FEAC9B705FFBD69F0FC69422D5994" bold="true" box="[192,324,959,985]" pageId="10" pageNumber="9">Etymology.</emphasis>
The specific epithet is named for the country of
<collectingCountry id="65FC764BB705FFBD6A6FFC6940BE5995" box="[863,983,958,985]" name="Costa Rica" pageId="10" pageNumber="9">Costa Rica</collectingCountry>
from which the new species was taken.
</paragraph>
</subSubSection>
</treatment>
</document>