385 lines
31 KiB
XML
385 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.222.98275" ID-Pensoft-Pub="1314-2003-222-19" ID-Pensoft-UUID="ADEB9E9C6CD65BDFA9FA09D5A02E9AE4" ModsDocID="1314-2003-222-19" checkinTime="1679436444134" checkinUser="pensoft" docAuthor="De Leon, Miguel David, Cabactulan, Derek, Cuerdo, Ryu D. & Rodda, Michele" docDate="2023" docId="77B9CB2B847B5F4FB42BC8E18EA42DF7" docLanguage="en" docName="PhytoKeys 222: 19-26" docOrigin="PhytoKeys 222" docPubDate="2023-03-20" docSource="http://dx.doi.org/10.3897/phytokeys.222.98275" docTitle="Hoya medusa M. D. De Leon, Cabactulan, Cuerdo & Rodda 2023, sp. nov." docType="treatment" docVersion="1" id="ADEB9E9C6CD65BDFA9FA09D5A02E9AE4" lastPageNumber="19" masterDocId="ADEB9E9C6CD65BDFA9FA09D5A02E9AE4" masterDocTitle="A new species of Hoya R. Br. (Apocynaceae, Asclepiadoideae) from the Philippines" masterLastPageNumber="26" masterPageNumber="19" pageNumber="19" updateTime="1679436444134" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of Hoya R. Br. (Apocynaceae, Asclepiadoideae) from the Philippines</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>De Leon, Miguel David</mods:namePart>
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<mods:affiliation>Cagayan de Oro Medical Center, Tiano cor. Nacalaban Sts., Cagayan de Oro, Misamis Oriental 9000, Philippines</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cabactulan, Derek</mods:namePart>
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<mods:affiliation>48 Corrales and 1 st Streets, Nazareth, Cagayan de Oro City, Misamis Oriental 9000, Philippines</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cuerdo, Ryu D.</mods:namePart>
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<mods:affiliation>Tejero, San Luis, Batangas 4210, Philippines</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rodda, Michele</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4130-6685</mods:nameIdentifier>
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<mods:affiliation>Singapore Botanic Gardens, National Parks Board, 1 Cluny Road, Singapore 259569, Singapore</mods:affiliation>
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<mods:nameIdentifier type="email">rodda.michele@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>222</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>26</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.222.98275</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.222.98275</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-222-19</mods:identifier>
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<mods:identifier type="Pensoft-UUID">ADEB9E9C6CD65BDFA9FA09D5A02E9AE4</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:77B9CB2B847B5F4FB42BC8E18EA42DF7" httpUri="http://treatment.plazi.org/id/77B9CB2B847B5F4FB42BC8E18EA42DF7" lastPageNumber="19" pageId="0" pageNumber="19">
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<subSubSection pageId="0" pageNumber="19" type="nomenclature">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName LSID="77B9CB2B-847B-5F4F-B42B-C8E18EA42DF7" authority="M. D. De Leon, Cabactulan, Cuerdo & Rodda" authorityName="M. D. De Leon, Cabactulan, Cuerdo & Rodda" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa" status="sp. nov.">Hoya medusa M.D.De Leon, Cabactulan, Cuerdo & Rodda</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="19">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya medusa photographed in cultivation A flower, top view B corolla, from underneath C flower, side view two corolla lobes removed D flower, side view E corona, top view F corona, from underneath G, H corona, side view I calyx and ovaries J pollinarium. (Photographs by R. D. Cuerdo)." figureDoi="10.3897/phytokeys.222.98275.figure1" httpUri="https://binary.pensoft.net/fig/827584" pageId="0" pageNumber="19">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya medusa flowering branch. (Photograph by M. D. De Leon)." figureDoi="10.3897/phytokeys.222.98275.figure2" httpUri="https://binary.pensoft.net/fig/827585" pageId="0" pageNumber="19">, 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="diagnosis">
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<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Similar to
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya edeni" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="edeni">
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<emphasis italics="true" pageId="0" pageNumber="19">Hoya edeni</emphasis>
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</taxonomicName>
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King & Hook.f. in its shrubby habit and caudate and curved inner corona lobe processes, distinct by the corona processes (wavy to serpentine in
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
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<emphasis italics="true" pageId="0" pageNumber="19">H. medusa</emphasis>
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</taxonomicName>
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<emphasis italics="true" pageId="0" pageNumber="19">versus</emphasis>
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linear and curved at the tip in
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. edeni" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="edeni">
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<emphasis italics="true" pageId="0" pageNumber="19">H. edeni</emphasis>
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</taxonomicName>
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), corolla shape (urceolate in
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
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<emphasis italics="true" pageId="0" pageNumber="19">H. medusa</emphasis>
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</taxonomicName>
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<emphasis italics="true" pageId="0" pageNumber="19">versus</emphasis>
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rotate with reflexed lobes in
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. edeni" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="edeni">
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<emphasis italics="true" pageId="0" pageNumber="19">H. edeni</emphasis>
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</taxonomicName>
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) and size (8.5-10 mm in
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
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<emphasis italics="true" pageId="0" pageNumber="19">H. medusa</emphasis>
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</taxonomicName>
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<emphasis italics="true" pageId="0" pageNumber="19">versus</emphasis>
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c. 20 mm in
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. edeni" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="edeni">
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<emphasis italics="true" pageId="0" pageNumber="19">H. edeni</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="materials_examined">
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<paragraph pageId="0" pageNumber="19">
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<typeStatus>Type</typeStatus>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<materialsCitation collectingDate="2022-02-07" collectorName="M. D. De Leon" country="Philippines" elevation="750" location="Mt. Cetaceo" municipality="Luzon" specimenCount="1" stateProvince="Cagayan Province" typeStatus="holotype">
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<collectingCountry name="Philippines">Philippines</collectingCountry>
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.
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<collectingMunicipality>Luzon</collectingMunicipality>
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,
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<collectingRegion country="Philippines" name="Cagayan">Cagayan Province</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:77B9CB2B847B5F4FB42BC8E18EA42DF7:78FD63AFAE08AB20A7D70AB45CFE5835" country="Philippines" municipality="Luzon" name="Mt. Cetaceo" stateProvince="Cagayan Province">Mt. Cetaceo</location>
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<quantity metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">500 to 1,000 m</elevation>
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</quantity>
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, vouchered in cultivation,
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<collectingDate value="2022-02-07">07 Feb 2022</collectingDate>
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,
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<emphasis italics="true" pageId="0" pageNumber="19">
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<collectorName>M.D. De Leon</collectorName>
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s.n.
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</emphasis>
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(
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<typeStatus>holotype</typeStatus>
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: PNH, sheet no. 258696)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">Description.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Plant</emphasis>
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epiphytic pendent shrub, with white exudate in all vegetative parts.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Roots</emphasis>
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basal, fibrous.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Stems</emphasis>
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slender, terete, 1.5-7 mm in diam., green, sparsely puberulent when immature, turning lignified, brown and glabrescent when mature,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">internodes</emphasis>
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2.8-6 cm long.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Leaves</emphasis>
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: petiole stout, usually curved, canaliculate above, flattened towards lamina base, 3-7 mm long, 1.5-2.0 mm in diameter, dark green, sparsely pubescent turning glabrescent when old; blades coriaceous, stiff, flat to slightly curved, variable in shape, oblong, ovate, to elliptic-ovate, 55-90 mm long
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<normalizedToken originalValue="×">x</normalizedToken>
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20-34 mm wide; base acute to obtuse, apex acute to acuminate, with a caudate tip, adaxially mid-green, abaxially light green; margins entire, occasionally slightly undulate; adaxially and abaxially sparsely pubescent to glabrescent in older leaves; venation pinnate-brochidodromous, with 4-8 lateral veins on each sides of mid vein, prominent (very pale green) on younger leaves and obscure on older leaves yet clearly visible on dried specimens; colleters one at each lamina base, conical, c. 0.20 mm long, grayish brown.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Inflorescence</emphasis>
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extra-axillary, umbelliform, convex, with up to 11 flowers (occasionally up to 16 flowers in particularly strong cultivated specimens).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Peduncle</emphasis>
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short stout or sometimes sessile, cylindrical, 1.0-3.0 mm long and 1.0-2.0 mm in diam., perennial, sparsely pubescent.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Pedicels</emphasis>
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terete, outer pedicels slightly curved, otherwise straight, 8-10 mm long and 0.5-0.8 mm in diam., light green, lenticellate except the base of the calyx which is pubescent.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Calyx</emphasis>
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lobes oblong, 1.8-2 mm long, c. 0.50 mm wide, reddish, abaxially strigose, adaxially glabrous, basal colleters 1 between each calyx lobes, oblong, c. 0.02 mm long
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<normalizedToken originalValue="×">x</normalizedToken>
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c. 0.02 mm wide.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Corolla</emphasis>
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basally urceolate, with flat spreading lobes, white, corolla lobes spreading, 8.5 to 10 mm in diam. across the corolla lobes, corolla tube 2.0-3.0 mm long, 4-5 mm in diameter, lobes triangular ovate, 2.5-3.0 mm long, 2.5-3.5 mm wide, apex acute, slightly revolute, margins revolute, corolla lobes adaxially densely strigose, densely hirsute towards the rim and interior of the tube and the column, abaxially glabrous;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">gynostegium</emphasis>
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stipitate;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">column</emphasis>
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cylindrical, 0.50-0.70 mm high
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<normalizedToken originalValue="×">x</normalizedToken>
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c. 0.50 mm in diam.;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">corona</emphasis>
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staminal 4-5 mm high, c. 3 mm in diam.;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">corona lobe</emphasis>
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bulbous-obpyriform, c. 3.5 mm long
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<normalizedToken originalValue="×">x</normalizedToken>
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0.5-0.7 mm wide, inner (apical) process caudate, upright, curved, wavy to serpentine, meeting at the center and overlapping, erect above the gynostegium, outer (basal) process obovate, with basal revolute margins, guide rail raised, laterally compressed, prominent at the base of the corona lobes c. 0.80 mm long and extending c. 0.20 mm laterally.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Pollinia</emphasis>
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erect, elliptic-oblong, c. 0.50 mm long, c. 0.22 mm wide;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">caudicle</emphasis>
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obovoid, c. 0.10 mm diam;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">corpusculum</emphasis>
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oblong, c. 3.0 mm long by c. 1.0 mm wide;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">ovary</emphasis>
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ovoid, c. 1.5 mm long, each carpel c. 0.9 mm wise at the base, glabrous.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">Fruit and seed</emphasis>
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not seen. Flowers vespertine lasting up to 15 days in cultivation, drying of the inner corona processes begins on the second day causing brown to black discoloration, flowers slightly fragrant, fragrance floral, powdery scent or mild jasmine persistent throughout the day and night.
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</paragraph>
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<caption doi="10.3897/phytokeys.222.98275.figure1" httpUri="https://binary.pensoft.net/fig/827584" pageId="0" pageNumber="19" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">Figure 1.</emphasis>
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<taxonomicName authorityName="M. D. De Leon, Cabactulan, Cuerdo & Rodda" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
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<emphasis italics="true" pageId="0" pageNumber="19">Hoya medusa</emphasis>
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</taxonomicName>
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photographed in cultivation
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<emphasis bold="true" pageId="0" pageNumber="19">A</emphasis>
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flower, top view
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<emphasis bold="true" pageId="0" pageNumber="19">B</emphasis>
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corolla, from underneath
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<emphasis bold="true" pageId="0" pageNumber="19">C</emphasis>
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flower, side view two corolla lobes removed
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<emphasis bold="true" pageId="0" pageNumber="19">D</emphasis>
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flower, side view
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<emphasis bold="true" pageId="0" pageNumber="19">E</emphasis>
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corona, top view
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<emphasis bold="true" pageId="0" pageNumber="19">F</emphasis>
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corona, from underneath
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<emphasis bold="true" pageId="0" pageNumber="19">G, H</emphasis>
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corona, side view
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<emphasis bold="true" pageId="0" pageNumber="19">I</emphasis>
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calyx and ovaries
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<emphasis bold="true" pageId="0" pageNumber="19">J</emphasis>
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pollinarium. (Photographs by R.D. Cuerdo).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="etymology">
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<paragraph pageId="0" pageNumber="19">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="19">The specific epithet refers to the serpentine processes of the inner corona reminiscent of the snake headdress of Medusa in Greek mythology.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="distribution">
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<paragraph pageId="0" pageNumber="19">Distribution and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName authorityName="M. D. De Leon, Cabactulan, Cuerdo & Rodda" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
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<emphasis italics="true" pageId="0" pageNumber="19">Hoya medusa</emphasis>
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</taxonomicName>
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was collected by local collectors in Luzon Island, Mt. Cetaceo and has been in cultivation, circulated by local plant nurseries and plant hobbyists. It was first collected in low montane forest at 500 to 1,000 m where it was growing as an epiphyte in disturbed primary broadly leaf and mossy forest in full sun to part shade.
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</paragraph>
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<caption doi="10.3897/phytokeys.222.98275.figure2" httpUri="https://binary.pensoft.net/fig/827585" pageId="0" pageNumber="19" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">Figure 2.</emphasis>
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<taxonomicName authorityName="M. D. De Leon, Cabactulan, Cuerdo & Rodda" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
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<emphasis italics="true" pageId="0" pageNumber="19">Hoya medusa</emphasis>
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</taxonomicName>
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flowering branch. (Photograph by M.D. De Leon).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="conservation status">
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<paragraph pageId="0" pageNumber="19">Conservation status.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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The forested area where
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<taxonomicName authorityName="M. D. De Leon, Cabactulan, Cuerdo & Rodda" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
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<emphasis italics="true" pageId="0" pageNumber="19">Hoya medusa</emphasis>
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</taxonomicName>
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||
was collected is threatened by habitat destruction due to extensive harvesting of trees for local housing and wood-fuel use, destructive farming practices such as "
|
||
<emphasis italics="true" pageId="0" pageNumber="19">kaingin</emphasis>
|
||
" (slash and burn farming in Filipino), land conversion and illegal commercial logging. The type locality is a heavily logged forest. Parts of the deforested areas had been converted to agricultural land for crops such as corn, cassava, and vegetables. The western portion of the area is near a protected forested area and remains intact but it is endangered by rapid expansion of agricultural lands by the lowland settlements.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
<taxonomicName authorityName="M. D. De Leon, Cabactulan, Cuerdo & Rodda" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya medusa</emphasis>
|
||
</taxonomicName>
|
||
is only known from a single collection and we do not have information on the extant population size; therefore, the conservation status is proposed as Data Deficient (DD,
|
||
<bibRefCitation author="Camara-Leret, R" journalOrPublisher="Nature" pageId="0" pageNumber="19" publicationUrl="http://iucnredlist.org/resources/redlistguidelines" refId="B2" refString="2022. . http://iucnredlist.org/resources/redlistguidelines" url="http://iucnredlist.org/resources/redlistguidelines" year="2022">IUCN Standards and Petitions Committee (2022)</bibRefCitation>
|
||
) until more information is known about its area of occurrence.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="19" type="notes">
|
||
<paragraph pageId="0" pageNumber="19">Notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
<taxonomicName authorityName="M. D. De Leon, Cabactulan, Cuerdo & Rodda" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya medusa</emphasis>
|
||
</taxonomicName>
|
||
displays an unusual combination of characters, in particular shrubby habit and corolla urceolate, with flat spreading lobes. This sets it apart from all other shrubby species of
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya</emphasis>
|
||
</taxonomicName>
|
||
. In the diagnosis we compared it to
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya edeni" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="edeni">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya edeni</emphasis>
|
||
</taxonomicName>
|
||
because both species are shrubs and present somewhat similar inner corona lobe processes, yet they present very different corolla morphology. The only other species with a similar corolla shape (salverform, with tube narrowly campanulate) is
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya kachinensis" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="kachinensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya kachinensis</emphasis>
|
||
</taxonomicName>
|
||
Rodda & K.Armstr. from Myanmar, which is a climber with oblanceolate, ca. 25 long leaves and therefore clearly distinct from
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. medusa</emphasis>
|
||
</taxonomicName>
|
||
. Other species that have a somewhat similar corolla shape are
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya telosmoides" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="telosmoides">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya telosmoides</emphasis>
|
||
</taxonomicName>
|
||
Omlor from Borneo, and
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. versteegii" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="versteegii">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. versteegii</emphasis>
|
||
</taxonomicName>
|
||
Simonsson & Rodda from New Guinea, which are both twining climbers. The only species with a tubular/urceolate corolla, somewhat spreading but much reduced corolla lobes as well as a shrubby habit is
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya manipurensis" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="manipurensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya manipurensis</emphasis>
|
||
</taxonomicName>
|
||
Deb from India, Myanmar, China and Thailand.
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya manipurensis" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="manipurensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya manipurensis</emphasis>
|
||
</taxonomicName>
|
||
has obcordate to triangular laminas and has corolla lobes much shorter than corolla tube, making it once again very easily separated from
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. medusa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="19">
|
||
Among other species of
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya</emphasis>
|
||
</taxonomicName>
|
||
occurring in the Philippines
|
||
<taxonomicName authorityName="M. D. De Leon, Cabactulan, Cuerdo & Rodda" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya medusa</emphasis>
|
||
</taxonomicName>
|
||
is somewhat similar to
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya linavergarae" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="linavergarae">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya linavergarae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya odorata" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="odorata">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">Hoya odorata</emphasis>
|
||
</taxonomicName>
|
||
. The leaf margins of
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. medusa</emphasis>
|
||
</taxonomicName>
|
||
are slightly undulate, whereas undulate in
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. linavergarae" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="linavergarae">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. linavergarae</emphasis>
|
||
</taxonomicName>
|
||
, and flat in
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. odorata" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="odorata">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. odorata</emphasis>
|
||
</taxonomicName>
|
||
. The flowers of
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. medusa</emphasis>
|
||
</taxonomicName>
|
||
are smaller (8.5 to 10 mm) and basally urceolate with spreading lobes, whereas
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. linavergarae" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="linavergarae">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. linavergarae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. odorata" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="odorata">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. odorata</emphasis>
|
||
</taxonomicName>
|
||
have larger flowers (15-23 mm and 13-18 mm respectively), and basally campanulate (
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. linavergarae" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="linavergarae">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. linavergarae</emphasis>
|
||
</taxonomicName>
|
||
), or rotate (
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. odorata" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="odorata">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. odorata</emphasis>
|
||
</taxonomicName>
|
||
) with inflexed corolla lobes. Further, the corolla of
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. medusa</emphasis>
|
||
</taxonomicName>
|
||
is densely hirsute inside, whereas puberulent in
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. linavergarae" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="linavergarae">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. linavergarae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. odorata" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="odorata">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. odorata</emphasis>
|
||
</taxonomicName>
|
||
. The inner corona processes of
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. medusa" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="medusa">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. medusa</emphasis>
|
||
</taxonomicName>
|
||
have a caudate, upright, curved, wavy to serpentine appendage, whereas the inner corona processes of
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. linavergarae" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="linavergarae">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. linavergarae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. odorata" order="Gentianales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="odorata">
|
||
<emphasis italics="true" pageId="0" pageNumber="19">H. odorata</emphasis>
|
||
</taxonomicName>
|
||
do not have an appendage.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |