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<mods:title id="6BCC995BBE4001C99127E3C55647C84D">Unexpected discovery of a new Lycocerus (Coleoptera, Cantharidae) from Okinawa Island, Ryukyu Islands, southwest Japan</mods:title>
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<treatment id="7742879BFFA6FFC0C2BFFC15F6ECFCC4" ID-DOI="http://doi.org/10.5281/zenodo.13758009" ID-Zenodo-Dep="13758009" LSID="urn:lsid:plazi:treatment:7742879BFFA6FFC0C2BFFC15F6ECFCC4" httpUri="http://treatment.plazi.org/id/7742879BFFA6FFC0C2BFFC15F6ECFCC4" lastPageId="5" lastPageNumber="494" pageId="4" pageNumber="493">
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<paragraph id="FF54368DFFA6FFC1C2BFFC15F45BFBC1" blockId="4.[151,606,1014,1077]" box="[151,606,1014,1041]" pageId="4" pageNumber="493">
<heading id="A41C81E1FFA6FFC1C2BFFC15F45BFBC1" bold="true" box="[151,606,1014,1041]" fontSize="11" level="1" pageId="4" pageNumber="493" reason="1">
<taxonomicName id="38EB4D0EFFA6FFC1C2BFFC15F45BFBC1" authority="(Ishida, 1985)" baseAuthorityName="Ishida" baseAuthorityYear="1985" box="[151,606,1014,1041]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFC15F45BFBC1" bold="true" box="[151,606,1014,1041]" pageId="4" pageNumber="493">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFC15F7A9FBC0" bold="true" box="[151,428,1014,1041]" italics="true" pageId="4" pageNumber="493">Lycocerus okinawanus</emphasis>
(Ishida, 1985)
</emphasis>
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<paragraph id="FF54368DFFA6FFC1C2BFFBF9F73EFBE4" blockId="4.[151,606,1014,1077]" box="[151,315,1051,1077]" pageId="4" pageNumber="493">
(
<figureCitation id="67D02A08FFA6FFC1C2B7FBF9F6EAFBE4" box="[159,239,1051,1077]" captionStart="FIGURES 16" captionStartId="2.[151,264,1763,1787]" captionTargetBox="[175,1412,194,1726]" captionTargetId="figure-28@2.[162,1425,181,1739]" captionTargetPageId="2" captionText="FIGURES 16. Habitus of Lycocerus spp., male. 1 &amp; 46, L. todai sp. nov., 1: holotype, dorsal; 4: paratype from Jana, Oppadale, ventral; 5: ditto, alive, dorsal; 6: ditto, alive, lateral; 2, L. okinawanus, dorsal; 3, L. yonaensis, dorsal. Scale bars: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13758005" httpUri="https://zenodo.org/record/13758005/files/figure.png" pageId="4" pageNumber="493">Figs. 2</figureCitation>
,
<figureCitation id="67D02A08FFA6FFC1C2D3FBF9F70FFBE4" box="[251,266,1051,1077]" captionStart="FIGURES 79" captionStartId="3.[151,264,1924,1948]" captionTargetBox="[172,1415,195,1888]" captionTargetId="figure-15@3.[159,1427,181,1901]" captionTargetPageId="3" captionText="FIGURES 79.Aedeagus, male tergite IX and male sternite IX of Lycocerus spp. 7, L. todai sp. nov., paratype; 8, L. okinawanus; 9, L. yonaensis. A: aedeagus, ventral; B: ditto, lateral; C: ditto, dorsal; D: tergite IX and sternite IX, ventral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13758007" httpUri="https://zenodo.org/record/13758007/files/figure.png" pageId="4" pageNumber="493">8</figureCitation>
,
<figureCitation id="67D02A08FFA6FFC1C33FFBF9F737FBE4" box="[279,306,1051,1077]" captionStart="FIGURE 11" captionStartId="6.[151,250,1652,1676]" captionTargetBox="[169,1422,980,1627]" captionTargetId="figure-289@6.[166,1422,977,1627]" captionTargetPageId="6" captionText="FIGURE 11. Distribution map of three black-colored Lycocerus species in the Ryukyu Islands, created based on the distributional information presented in this paper." figureDoi="http://doi.org/10.5281/zenodo.13758013" httpUri="https://zenodo.org/record/13758013/files/figure.png" pageId="4" pageNumber="493">11</figureCitation>
)
</paragraph>
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<subSubSection id="B7F16506FFA6FFC1C2BFFBBDF335FB02" pageId="4" pageNumber="493" type="reference_group">
<paragraph id="FF54368DFFA6FFC1C2BFFBBDF238FBA6" blockId="4.[151,1436,1119,1235]" box="[151,1085,1119,1143]" pageId="4" pageNumber="493">
<treatmentCitationGroup id="DFFB11A3FFA6FFC1C2BFFBBDF238FBA6" box="[151,1085,1119,1143]" pageId="4" pageNumber="493">
<taxonomicName id="38EB4D0EFFA6FFC1C2BFFBBDF7B7FBA6" authority="Ishida" authorityName="Ishida" box="[151,434,1119,1143]" class="Insecta" family="Cantharidae" genus="Athemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFBBDF76BFBA6" box="[151,366,1119,1143]" italics="true" pageId="4" pageNumber="493">Athemus okinawanus</emphasis>
Ishida
</taxonomicName>
in
<treatmentCitation id="7E4A109CFFA6FFC1C3FAFBBDF476FBA6" author="Sato, M." box="[466,627,1119,1143]" page="113" pageId="4" pageNumber="493" year="1985">
<bibRefCitation id="9B7A4B7CFFA6FFC1C3FAFBBDF476FBA6" author="Sato, M." box="[466,627,1119,1143]" pageId="4" pageNumber="493" refId="ref5030" refString="Sato, M. (1985) Cantharidae. In: Kurosawa, Y., Hisamatsu, S. &amp; Sasaji, H. (Eds.), The Coleoptera of Japan in Color. Vol. 3. Hoikusha, Osaka, pp. 107 - 119. [in Japanese, with English book title]" type="book" year="1985">Satô, 1985: 113</bibRefCitation>
</treatmentCitation>
, pl. 18, fig. 23 (Okinawa-hontô Is., Ryukyu).
</treatmentCitationGroup>
</paragraph>
<paragraph id="FF54368DFFA6FFC1C2BFFB9CF79EFB64" blockId="4.[151,1436,1119,1235]" pageId="4" pageNumber="493">
<treatmentCitationGroup id="DFFB11A3FFA6FFC1C2BFFB9CF79EFB64" pageId="4" pageNumber="493">
<treatmentCitation id="7E4A109CFFA6FFC1C2BFFB9CF433FB47" author="Ishida, K." box="[151,566,1150,1174]" page="196" pageId="4" pageNumber="493" year="1986">
<taxonomicName id="38EB4D0EFFA6FFC1C2BFFB9CF433FB47" authority="Ishida, 1986: 196" authorityName="Ishida" authorityPageNumber="196" authorityYear="1986" box="[151,566,1150,1174]" class="Insecta" family="Cantharidae" genus="Athemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFB9CF777FB47" box="[151,370,1150,1174]" italics="true" pageId="4" pageNumber="493">Athemus okinawanus</emphasis>
<bibRefCitation id="9B7A4B7CFFA6FFC1C354FB9CF433FB47" author="Ishida, K." box="[380,566,1150,1174]" pageId="4" pageNumber="493" pagination="193 - 213" refId="ref4551" refString="Ishida, K. (1986) A revision of the two genera Athemus and Athemellus, of Japan (Coleoptera: Cantharidae). Transactions of the Shikoku Entomological Society, 17, 193 - 213." type="journal article" year="1986">Ishida, 1986: 196</bibRefCitation>
</taxonomicName>
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, figs. 47 (Ginama-mura, Kunigami,
<collectingRegion id="3D2FF86FFFA6FFC1C1EEFB9CF224FB47" box="[966,1057,1150,1174]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">Okinawa</collectingRegion>
Is.);
<treatmentCitation id="7E4A109CFFA6FFC1C676FB9CF39DFB47" author="Sato, M. &amp; Okushima, Y." box="[1118,1432,1150,1174]" page="858" pageId="4" pageNumber="493" year="1992">
<bibRefCitation id="9B7A4B7CFFA6FFC1C676FB9CF39DFB47" author="Sato, M. &amp; Okushima, Y." box="[1118,1432,1150,1174]" pageId="4" pageNumber="493" pagination="857 - 865" refId="ref5090" refString="Sato, M. &amp; Okushima, Y. (1992) The genus Athemus (Coleoptera, Cantharidae) of the Ryukyu Islands, southwest Japan. Japanese Journal of Entomology, 60, 857 - 865." type="journal article" year="1992">Satô &amp; Okushima, 1992: 858</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="7E4A109CFFA6FFC1C2EFFB7EF793FB64" author="Takahashi, K." box="[199,406,1180,1205]" page="40" pageId="4" pageNumber="493" year="1998">
<bibRefCitation id="9B7A4B7CFFA6FFC1C2EFFB7EF793FB64" author="Takahashi, K." box="[199,406,1180,1205]" pageId="4" pageNumber="493" pagination="29 - 48" refId="ref5129" refString="Takahashi, K. (1998) Check-list of Cantharidae of Japan. Kanagawa-Chuho, 122, 29 - 48. [in Japanese, with English title]" type="journal article" year="1998">Takahashi, 1998: 40</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="FF54368DFFA6FFC1C2BFFB5EF335FB02" blockId="4.[151,1436,1119,1235]" box="[151,1328,1211,1235]" pageId="4" pageNumber="493">
<treatmentCitationGroup id="DFFB11A3FFA6FFC1C2BFFB5EF335FB02" box="[151,1328,1211,1235]" pageId="4" pageNumber="493">
<taxonomicName id="38EB4D0EFFA6FFC1C2BFFB5EF413FB02" authority="(Ishida, 1986)" baseAuthorityName="Ishida" baseAuthorityYear="1986" box="[151,534,1211,1235]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA6FFC1C2BFFB5EF77BFB02" box="[151,382,1211,1235]" italics="true" pageId="4" pageNumber="493">Lycocerus okinawanus</emphasis>
(
<bibRefCitation id="9B7A4B7CFFA6FFC1C3A3FB59F40BFB02" author="Ishida, K." box="[395,526,1211,1235]" pageId="4" pageNumber="493" pagination="193 - 213" refId="ref4551" refString="Ishida, K. (1986) A revision of the two genera Athemus and Athemellus, of Japan (Coleoptera: Cantharidae). Transactions of the Shikoku Entomological Society, 17, 193 - 213." type="journal article" year="1986">Ishida, 1986</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="7E4A109CFFA6FFC1C008FB59F4F8FB02" author="Okushima, Y." box="[544,765,1211,1235]" page="262" pageId="4" pageNumber="493" year="2005">
<bibRefCitation id="9B7A4B7CFFA6FFC1C008FB59F4F8FB02" author="Okushima, Y." box="[544,765,1211,1235]" pageId="4" pageNumber="493" pagination="1 - 383" refId="ref4882" refString="Okushima, Y. (2005) A taxonomic study on the genus Lycocerus (Coleoptera, Cantharidae) from Japan, with zoogeographical considerations. Japanese Journal of Systematic Entomology, Monographic Series, 2, 1 - 383." type="journal article" year="2005">Okushima, 2005: 262</bibRefCitation>
</treatmentCitation>
, figs. 65, 136, 179;
<treatmentCitation id="7E4A109CFFA6FFC1C1EEFB5EF329FB02" author="Kazantsev, S. &amp; Brancucci, M." box="[966,1324,1211,1235]" page="252" pageId="4" pageNumber="493" year="2007">
<bibRefCitation id="9B7A4B7CFFA6FFC1C1EEFB5EF329FB02" author="Kazantsev, S. &amp; Brancucci, M." box="[966,1324,1211,1235]" pageId="4" pageNumber="493" refId="ref4627" refString="Kazantsev, S. &amp; Brancucci, M. (2007) Cantharidae. In Lo ¨ bl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 234 - 298." type="book" year="2007">Kazantsev &amp; Brancucci, 2007: 252</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<emphasis id="CD9FEA9FFFA6FFC1C2BFFAE3F7CCFACA" bold="true" box="[151,457,1281,1307]" pageId="4" pageNumber="493">Type materials examined.</emphasis>
<materialsCitation id="4F833CD0FFA6FFC1C3F8FAE3F6F3FAEE" collectingDate="1978-04-01" collectionCode="EUMJ" collectorName="N. Kanie" location="Kunigami" municipality="Ginama Vil." pageId="4" pageNumber="493" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<specimenCount id="E9EDFD04FFA6FFC1C3F8FAE3F41FFACA" box="[464,538,1281,1307]" count="1" pageId="4" pageNumber="493" type="male">1 male</specimenCount>
(
<typeStatus id="2050882FFFA6FFC1C001FAE3F494FACA" box="[553,657,1281,1307]" pageId="4" pageNumber="493" type="holotype">holotype</typeStatus>
), “
<collectingMunicipality id="1F30ACF7FFA6FFC1C086FAE3F53EFACA" box="[686,827,1281,1307]" pageId="4" pageNumber="493">Ginama Vil.</collectingMunicipality>
,
<location id="FA346056FFA6FFC1C16EFAE3F5BCFACA" LSID="urn:lsid:plazi:treatment:7742879BFFA6FFC0C2BFFC15F6ECFCC4:FA346056FFA6FFC1C16EFAE3F5BCFACA" box="[838,953,1281,1307]" municipality="Ginama Vil." name="Kunigami" pageId="4" pageNumber="493" stateProvince="Okinawa">Kunigami</location>
,
<collectingRegion id="3D2FF86FFFA6FFC1C1EBFAE3F26DFACA" box="[963,1128,1281,1307]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">Okinawa Pref.</collectingRegion>
,
<date id="8B55104DFFA6FFC1C65AFAE3F2F4FACA" box="[1138,1265,1281,1308]" pageId="4" pageNumber="493" value="1978-04-01">
<collectingDate id="9B11E9A5FFA6FFC1C65AFAE3F2F4FACA" box="[1138,1265,1281,1308]" pageId="4" pageNumber="493" value="1978-04-01">1. IV. 1978</collectingDate>
</date>
,
<collectorName id="521E535BFFA6FFC1C6D3FAE0F365FACA" box="[1275,1376,1281,1308]" pageId="4" pageNumber="493">N. Kanie</collectorName>
leg.” (
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)
</materialsCitation>
;
<materialsCitation id="4F833CD0FFA6FFC1C32AFAC7F4BAFAEE" box="[258,703,1317,1344]" collectingDate="1978-04-01" collectionCode="EUMJ" collectorName="N. Kanie" location="Kunigami" municipality="Ginama Vil." pageId="4" pageNumber="493" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
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(
<typeStatus id="2050882FFFA6FFC1C359FAC7F7D2FAEE" box="[369,471,1317,1343]" pageId="4" pageNumber="493" type="paratype">paratype</typeStatus>
), same data (
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;
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), “
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,
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,
<collectingRegion id="3D2FF86FFFA6FFC1C6ADFAC7F2EFFAEE" box="[1157,1258,1317,1343]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">Okinawa</collectingRegion>
<location id="FA346056FFA6FFC1C6D9FAC4F308FA91" LSID="urn:lsid:plazi:treatment:7742879BFFA6FFC0C2BFFC15F6ECFCC4:FA346056FFA6FFC1C6D9FAC4F308FA91" box="[1265,1293,1318,1344]" county="Yona" municipality="Kunigami" name="Is." pageId="4" pageNumber="493" stateProvince="Okinawa">Is.</location>
,
<date id="8B55104DFFA6FFC1C732FAC7F39DFAEE" box="[1306,1432,1317,1344]" pageId="4" pageNumber="493" value="1978-04-02">
<collectingDate id="9B11E9A5FFA6FFC1C732FAC7F39DFAEE" box="[1306,1432,1317,1344]" pageId="4" pageNumber="493" value="1978-04-02">2. IV. 1978</collectingDate>
</date>
,
<collectorName id="521E535BFFA6FFC1C2BFFAA8F6F9FAB2" box="[151,252,1353,1380]" pageId="4" pageNumber="493">N. Kanie</collectorName>
leg.” (
<collectionCode id="99FAAE48FFA6FFC1C36FFAABF793FAB2" box="[327,406,1353,1379]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4" pageNumber="493">EUMJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4F833CD0FFA6FFC1C382FAABF6F2FA56" collectingDate="1981-06-07" collectionCode="EUMJ" location="OKINAWA" pageId="4" pageNumber="493" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="E9EDFD04FFA6FFC1C382FAABF40FFAB2" box="[426,522,1353,1379]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
(
<typeStatus id="2050882FFFA6FFC1C031FAABF47AFAB2" box="[537,639,1353,1379]" pageId="4" pageNumber="493" type="paratype">paratype</typeStatus>
), “IERINDOH, KUNIGAMI V,
<collectingRegion id="3D2FF86FFFA6FFC1C1C1FAABF275FAB2" box="[1001,1136,1353,1379]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">OKINAWA</collectingRegion>
,
<date id="8B55104DFFA6FFC1C653FAA8F2E8FAB2" box="[1147,1261,1353,1380]" pageId="4" pageNumber="493" value="1981-06-07">
<collectingDate id="9B11E9A5FFA6FFC1C653FAA8F2E8FAB2" box="[1147,1261,1353,1380]" pageId="4" pageNumber="493" value="1981-06-07">7 VI 1981</collectingDate>
</date>
, T. OHMOTO” (
<collectionCode id="99FAAE48FFA6FFC1C2B7FA8FF6EBFA56" box="[159,238,1389,1415]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4" pageNumber="493">EUMJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="FF54368DFFA6FFC0C2EFFA73F27FFECC" blockId="4.[151,1437,1281,2028]" lastBlockId="5.[151,1437,151,789]" lastPageId="5" lastPageNumber="494" pageId="4" pageNumber="493">
<emphasis id="CD9FEA9FFFA6FFC1C2EFFA73F444FA7A" bold="true" box="[199,577,1425,1451]" pageId="4" pageNumber="493">Additional materials examined.</emphasis>
<specimenCount id="E9EDFD04FFA6FFC1C061FA73F4A6FA7A" box="[585,675,1425,1451]" count="3" pageId="4" pageNumber="493" type="male">3 males</specimenCount>
, “Yona, Kunigami-son, Okinawa Pref.,
<date id="8B55104DFFA6FFC1C64DFA73F2FBFA7D" box="[1125,1278,1425,1452]" pageId="4" pageNumber="493" value="1995-02-28">28 Feb. 1995</date>
, H. Sato leg.” (
<collectionCode id="99FAAE48FFA6FFC1C2B7FA57F6F0FA1E" box="[159,245,1461,1487]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C321FA57F75DFA1E" box="[265,344,1461,1487]" count="1" pageId="4" pageNumber="493" type="male">1 male</specimenCount>
,
<specimenCount id="E9EDFD04FFA6FFC1C34DFA57F7CFFA1E" box="[357,458,1461,1487]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
, same data but “
<date id="8B55104DFFA6FFC1C0AEFA57F51AFA1E" box="[646,799,1461,1487]" pageId="4" pageNumber="493" value="1994-02-13">13 Feb. 1994</date>
, T. Shimizu leg.” (
<collectionCode id="99FAAE48FFA6FFC1C1DCFA57F24FFA1E" box="[1012,1098,1461,1487]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C676FA57F2C6FA1E" box="[1118,1219,1461,1487]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
, same data but “
<date id="8B55104DFFA6FFC1C757FA57F70EFA22" pageId="4" pageNumber="493" value="1994-02-25">25 Feb. 1994</date>
, T. Shimizu leg.” (
<collectionCode id="99FAAE48FFA6FFC1C3F7FA3BF430FA22" box="[479,565,1497,1523]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C061FA3BF492FA22" box="[585,663,1497,1523]" count="1" pageId="4" pageNumber="493" type="male">1 male</specimenCount>
,
<specimenCount id="E9EDFD04FFA6FFC1C08CFA3BF516FA22" box="[676,787,1497,1523]" count="3" pageId="4" pageNumber="493" type="female">3 females</specimenCount>
, same data but “
<date id="8B55104DFFA6FFC1C1E6FA3BF25EFA22" box="[974,1115,1497,1524]" pageId="4" pageNumber="493" value="1991-03-06">6 Mar. 1991</date>
, Y. Okushima leg. (
<collectionCode id="99FAAE48FFA6FFC1C712FA3BF395FA22" box="[1338,1424,1497,1523]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C2BFFA1FF70DF9C6" box="[151,264,1533,1559]" count="2" pageId="4" pageNumber="493" type="female">2 females</specimenCount>
, same data but “
<date id="8B55104DFFA6FFC1C3EEFA1FF451F9C6" box="[454,596,1533,1560]" pageId="4" pageNumber="493" value="1991-03-07">7 Mar. 1991</date>
, Y. Okushima leg.” (
<collectionCode id="99FAAE48FFA6FFC1C16BFA1FF59CF9C6" box="[835,921,1533,1559]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C185FA1FF216F9C6" box="[941,1043,1533,1559]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
, same data but “
<date id="8B55104DFFA6FFC1C6F8FA1FF39DF9C6" box="[1232,1432,1533,1560]" pageId="4" pageNumber="493" value="1988-03-12" valueMax="1988-03-14" valueMin="1988-03-12">1214 Mar. 1988</date>
, Coll. Teruhisa Ueno” (
<collectionCode id="99FAAE48FFA6FFC1C3BEF9C3F7E9F9EA" box="[406,492,1569,1595]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C028F9C3F44AF9EA" box="[512,591,1569,1595]" count="1" pageId="4" pageNumber="493" type="male">1 male</specimenCount>
,
<specimenCount id="E9EDFD04FFA6FFC1C073F9C3F4C5F9EA" box="[603,704,1569,1595]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
, “Yona, Kunigami-son, Okinawa, Ryukyus,
<collectingCountry id="87FC761DFFA6FFC1C69DF9C0F2FDF9ED" box="[1205,1272,1570,1596]" name="Japan" pageId="4" pageNumber="493">Japan</collectingCountry>
,
<date id="8B55104DFFA6FFC1C72BF9C3F37EF9ED" box="[1283,1403,1569,1596]" pageId="4" pageNumber="493" value="1991-03-05">5-III-1991</date>
, T. Ueno leg.” (
<collectionCode id="99FAAE48FFA6FFC1C336F9A7F771F98E" box="[286,372,1605,1631]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C3AFF9A7F7EFF98E" box="[391,490,1605,1631]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
, “
<collectingRegion id="3D2FF86FFFA6FFC1C3D6F9A7F486F98E" box="[510,643,1605,1631]" country="Japan" name="Okinawa" pageId="4" pageNumber="493">OKINAWA</collectingRegion>
IS., Yona,
<date id="8B55104DFFA6FFC1C0D5F9A7F571F98E" box="[765,884,1605,1631]" pageId="4" pageNumber="493" value="1988-03-13">1988-3-13</date>
, T. Ueno leg.” (
<collectionCode id="99FAAE48FFA6FFC1C60AF9A7F27DF98E" box="[1058,1144,1605,1631]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C6A3F9A7F2D7F98E" box="[1163,1234,1605,1631]" count="1" pageId="4" pageNumber="493" type="male">1male</specimenCount>
, “Yona, Okinawa, Ryukyus,
<date id="8B55104DFFA6FFC1C322F98BF78AF952" box="[266,399,1641,1667]" pageId="4" pageNumber="493" value="1994-04-20">20-IV-1994</date>
, M. Kawanabe” (
<collectionCode id="99FAAE48FFA6FFC1C070F98BF4A2F952" box="[600,679,1641,1667]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4" pageNumber="493">EUMJ</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C095F98BF51DF952" box="[701,792,1641,1668]" count="5" pageId="4" pageNumber="493" type="male">5 males</specimenCount>
,
<specimenCount id="E9EDFD04FFA6FFC1C10DF98BF593F952" box="[805,918,1641,1667]" count="2" pageId="4" pageNumber="493" type="female">2 females</specimenCount>
, “Mt. Yonaha-dake, Kunigami Son, Okinawa Pref.,
<date id="8B55104DFFA6FFC1C2F5F96FF768F976" box="[221,365,1677,1704]" pageId="4" pageNumber="493" value="1991-03-08">8 Mar. 1991</date>
, Y. Okushima leg. (
<collectionCode id="99FAAE48FFA6FFC1C07BF96FF4ACF976" box="[595,681,1677,1703]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C097F96FF51FF976" box="[703,794,1677,1703]" count="2" pageId="4" pageNumber="493" type="male">2 males</specimenCount>
, same data but “
<date id="8B55104DFFA6FFC1C1F5F96FF26BF979" box="[989,1134,1677,1704]" pageId="4" pageNumber="493" value="1995-03-03">3 Mar. 1995</date>
, H. Sato leg.” (
<collectionCode id="99FAAE48FFA6FFC1C70BF96FF37CF976" box="[1315,1401,1677,1703]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C7A6F96FF6D5F91A" count="1" pageId="4" pageNumber="493" type="male">1 male</specimenCount>
,
<specimenCount id="E9EDFD04FFA6FFC1C2F6F953F741F91A" box="[222,324,1713,1739]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
, same data but “
<date id="8B55104DFFA6FFC1C02EF953F492F91A" box="[518,663,1713,1740]" pageId="4" pageNumber="493" value="1999-03-02">2 Mar. 1999</date>
, Coll. T. Nonaka” (
<collectionCode id="99FAAE48FFA6FFC1C151F953F5CAF91A" box="[889,975,1713,1739]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C1CCF953F24FF91A" box="[996,1098,1713,1739]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
, same data but “
<date id="8B55104DFFA6FFC1C724F953F39DF91A" box="[1292,1432,1713,1740]" pageId="4" pageNumber="493" value="1990-04-05">5 Apr. 1990</date>
, Coll. Y. Okushima” (
<collectionCode id="99FAAE48FFA6FFC1C3AFF937F7D8F93E" box="[391,477,1749,1775]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C3DAF937F466F93E" box="[498,611,1749,1775]" count="4" pageId="4" pageNumber="493" type="female">4 females</specimenCount>
, “Ohkuni-ridou, Okinawa, Ryukyus,
<date id="8B55104DFFA6FFC1C625F937F28CF93E" box="[1037,1161,1749,1775]" pageId="4" pageNumber="493" value="2003-02-10">10-II-2003</date>
, M. Sato leg.” (
<collectionCode id="99FAAE48FFA6FFC1C769F937F395F93E" box="[1345,1424,1749,1775]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="4" pageNumber="493">EUMJ</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C2BFF91BF6F6F8C2" box="[151,243,1785,1811]" count="2" pageId="4" pageNumber="493" type="male">2 males</specimenCount>
,
<specimenCount id="E9EDFD04FFA6FFC1C32AF91BF771F8C2" box="[258,372,1785,1811]" count="4" pageId="4" pageNumber="493" type="female">4 females</specimenCount>
, “
<collectingCountry id="87FC761DFFA6FFC1C3A5F91BF7ECF8C2" box="[397,489,1785,1811]" name="Japan" pageId="4" pageNumber="493">JAPAN</collectingCountry>
: Okinawa-jima Is., Kunigami-son, Ura, Ôkuni-rindô,
<date id="8B55104DFFA6FFC1C67DF91BF2E7F8C2" box="[1109,1250,1785,1812]" pageId="4" pageNumber="493" value="2024-02-14">14. II. 2024</date>
, Ryo Nakamura leg.
<emphasis id="CD9FEA9FFFA6FFC1C2E0F8FCF7EDF8E9" bold="true" box="[200,488,1822,1848]" pageId="4" pageNumber="493">ṗäŝ 国DZť大国ṳã</emphasis>
” (
<collectionCode id="99FAAE48FFA6FFC1C02BF8FFF44AF8E6" box="[515,591,1821,1847]" pageId="4" pageNumber="493">FZUT</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C04BF8FFF4D7F8E6" box="[611,722,1821,1847]" count="2" pageId="4" pageNumber="493" type="female">2 females</specimenCount>
, “[
<emphasis id="CD9FEA9FFFA6FFC1C0DBF8FCF5A6F8E9" bold="true" box="[755,931,1822,1848]" pageId="4" pageNumber="493">ṗäŝ ạüƜ</emphasis>
]
<emphasis id="CD9FEA9FFFA6FFC1C19CF8FCF301F8E9" bold="true" box="[948,1284,1822,1848]" pageId="4" pageNumber="493">ṗäḁ国DZō国DZťṉǭẹ</emphasis>
,
<date id="8B55104DFFA6FFC1C73AF8FFF39DF8E9" box="[1298,1432,1821,1848]" pageId="4" pageNumber="493" value="2015-02-28">28. II. 2015</date>
, leg.
<emphasis id="CD9FEA9FFFA6FFC1C2E2F8A0F73FF88D" bold="true" box="[202,314,1858,1884]" pageId="4" pageNumber="493">DƂ尚*</emphasis>
” (
<collectionCode id="99FAAE48FFA6FFC1C370F8A3F7A1F88A" box="[344,420,1857,1883]" pageId="4" pageNumber="493">FZUT</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C392F8A3F40FF88A" box="[442,522,1857,1883]" count="1" pageId="4" pageNumber="493" type="male">1 male</specimenCount>
,
<specimenCount id="E9EDFD04FFA6FFC1C030F8A3F48CF88A" box="[536,649,1857,1883]" count="2" pageId="4" pageNumber="493" type="female">2 females</specimenCount>
, “Benoki-dam, Kunigami Son, Okinawa Pref.,
<date id="8B55104DFFA6FFC1C680F8A3F33DF88D" box="[1192,1336,1857,1884]" pageId="4" pageNumber="493" value="1995-03-02">2 Mar. 1995</date>
, H. Sato leg.” (
<collectionCode id="99FAAE48FFA6FFC1C2F3F887F734F8AE" box="[219,305,1893,1919]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C36BF887F794F8AE" box="[323,401,1893,1919]" count="1" pageId="4" pageNumber="493" type="male">1 male</specimenCount>
, “
<emphasis id="CD9FEA9FFFA6FFC1C380F884F549F851" bold="true" box="[424,844,1894,1920]" pageId="4" pageNumber="493">mṵṗäḛŝṗä×ŝṉǭẹダム</emphasis>
, [
<collectingCountry id="87FC761DFFA6FFC1C148F884F5BAF851" box="[864,959,1894,1920]" name="Japan" pageId="4" pageNumber="493">JAPAN</collectingCountry>
]: the Ryukyus, Okinawa Isls., Okinawa Is., Benoki,
<geoCoordinate id="9ADF504AFFA6FFC1C2DDF86BF796F872" box="[245,403,1929,1955]" degrees="26" direction="north" minutes="48" orientation="latitude" pageId="4" pageNumber="493" precision="1" seconds="01.1" value="26.800304">26°4801.1”N</geoCoordinate>
128°16106.7”E,
<date id="8B55104DFFA6FFC1C074F86BF4DEF872" box="[604,731,1929,1956]" pageId="4" pageNumber="493" value="2023-03-05">5. III. 2023</date>
. J. Souma leg.” (
<collectionCode id="99FAAE48FFA6FFC1C1B9F86BF5D8F872" box="[913,989,1929,1955]" pageId="4" pageNumber="493">FZUT</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C1C7F86BF257F872" box="[1007,1106,1929,1955]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
, “
<emphasis id="CD9FEA9FFFA6FFC1C640F868F6CAF819" bold="true" pageId="4" pageNumber="493">mṵṗäḛŝṗä×ŝṉ ǭẹ</emphasis>
, [
<collectingCountry id="87FC761DFFA6FFC1C2CBF84CF747F819" box="[227,322,1966,1992]" name="Japan" pageId="4" pageNumber="493">JAPAN</collectingCountry>
]: the Ryukyus, Okinawa Isls., Okinawa Is., Benoki Dam,
<geoCoordinate id="9ADF504AFFA6FFC1C196F84FF258F816" box="[958,1117,1965,1991]" degrees="26" direction="north" minutes="47" orientation="latitude" pageId="4" pageNumber="493" precision="1" seconds="09.9" value="26.786083">26°4709.9”N</geoCoordinate>
<geoCoordinate id="9ADF504AFFA6FFC1C64BF84FF30BF819" box="[1123,1294,1965,1992]" degrees="128" direction="east" minutes="15" orientation="longitude" pageId="4" pageNumber="493" precision="1" seconds="15.9" value="128.25441">128°1515.9”E</geoCoordinate>
,
<date id="8B55104DFFA6FFC1C731F84FF39CF816" box="[1305,1433,1965,1992]" pageId="4" pageNumber="493" value="2023-03-05">5. III. 2023</date>
. J. Souma leg.” (
<collectionCode id="99FAAE48FFA6FFC1C364F833F79DF83A" box="[332,408,2001,2027]" pageId="4" pageNumber="493">FZUT</collectionCode>
);
<specimenCount id="E9EDFD04FFA6FFC1C384F833F414F83A" box="[428,529,2001,2027]" count="1" pageId="4" pageNumber="493" type="female">1 female</specimenCount>
, “Haneji, Nago City, Okinawa Pref.,
<date id="8B55104DFFA6FFC1C19EF833F24AF83A" box="[950,1103,2001,2028]" pageId="4" pageNumber="493" value="1991-03-11">11 Mar. 1991</date>
, Y. Okushima leg.” (
<collectionCode id="99FAAE48FFA6FFC1C712F833F395F83A" box="[1338,1424,2001,2027]" pageId="4" pageNumber="493">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA7FFC0C2BFFF75F6E3FF60" box="[151,230,151,177]" count="1" pageId="5" pageNumber="494" type="male">1 male</specimenCount>
,
<specimenCount id="E9EDFD04FFA7FFC0C2DBFF75F761FF60" box="[243,356,151,177]" count="2" pageId="5" pageNumber="494" type="female">2 females</specimenCount>
, “Haneji, Okinawa,
<date id="8B55104DFFA7FFC0C063FF75F4D9FF60" box="[587,732,151,177]" pageId="5" pageNumber="494" value="1986-03-23">23. III. 1986</date>
, M. Sawai leg.” (
<collectionCode id="99FAAE48FFA7FFC0C18CFF75F5F6FF60" box="[932,1011,151,177]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="5" pageNumber="494">EUMJ</collectionCode>
);
<specimenCount id="E9EDFD04FFA7FFC0C62FFF75F252FF60" box="[1031,1111,151,177]" count="1" pageId="5" pageNumber="494" type="male">1 male</specimenCount>
, “HANEJI, Is. Okinawa,
<date id="8B55104DFFA7FFC0C751FF75F6F9FF04" pageId="5" pageNumber="494" value="1987-03-16">16. III. 1987</date>
, M. SAWAI” (
<collectionCode id="99FAAE48FFA7FFC0C3B7FF59F7EEFF04" box="[415,491,187,213]" pageId="5" pageNumber="494">FZUT</collectionCode>
);
<specimenCount id="E9EDFD04FFA7FFC0C3D6FF59F467FF04" box="[510,610,187,213]" count="1" pageId="5" pageNumber="494" type="female">1 female</specimenCount>
, “Afuso, Onna Son, Okinawa Pref.,
<date id="8B55104DFFA7FFC0C1D0FF59F27BFF04" box="[1016,1150,187,213]" pageId="5" pageNumber="494" value="1989-04-03">3 Apr. 1989</date>
, Coll. T. Ueno” (
<collectionCode id="99FAAE48FFA7FFC0C712FF59F395FF04" box="[1338,1424,187,213]" pageId="5" pageNumber="494">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA7FFC0C2BFFF3DF6FBFF28" box="[151,254,223,249]" count="1" pageId="5" pageNumber="494" type="female">1 female</specimenCount>
, “Afuso, Okinawa-hontô Is., Okinawa Pref.,
<date id="8B55104DFFA7FFC0C122FF3DF5A0FF28" box="[778,933,223,249]" pageId="5" pageNumber="494" value="1989-04-21">21 Apr. 1989</date>
, Coll. Teruhisa Ueno” (
<collectionCode id="99FAAE48FFA7FFC0C692FF3DF315FF28" box="[1210,1296,223,249]" pageId="5" pageNumber="494">KURA</collectionCode>
);
<specimenCount id="E9EDFD04FFA7FFC0C70EFF3DF39DFF28" box="[1318,1432,223,249]" count="2" pageId="5" pageNumber="494" type="female">2 females</specimenCount>
, “Kenminnomori, Onna Son, Okinawa Pref.,
<date id="8B55104DFFA7FFC0C0A2FEE1F526FECC" box="[650,803,259,285]" pageId="5" pageNumber="494" value="1990-03-31">31 Mar. 1990</date>
, Coll. Y. Okushima” (
<collectionCode id="99FAAE48FFA7FFC0C630FEE1F26BFECC" box="[1048,1134,259,285]" pageId="5" pageNumber="494">KURA</collectionCode>
).
</paragraph>
<paragraph id="FF54368DFFA7FFC0C2EFFEC4F7ECFE58" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="494">
<emphasis id="CD9FEA9FFFA7FFC0C2EFFEC4F417FE91" bold="true" box="[199,530,294,320]" pageId="5" pageNumber="494">Supplementary description.</emphasis>
Male sternite IX (fig. 8D) cymbiform, slightly shorter than tergite IX, 2.80 times longer than wide, lateral margins parallel in the middle, abruptly narrowing in apical 1/4 to form a triangle with a rounded acute angle at the tip.
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA7FFC0C2EFFE70F545FE7C" box="[199,832,402,429]" pageId="5" pageNumber="494" type="distribution">
<paragraph id="FF54368DFFA7FFC0C2EFFE70F545FE7C" blockId="5.[151,1437,151,789]" box="[199,832,402,429]" pageId="5" pageNumber="494">
<emphasis id="CD9FEA9FFFA7FFC0C2EFFE70F764FE7D" bold="true" box="[199,353,402,428]" pageId="5" pageNumber="494">Distribution.</emphasis>
<collectingCountry id="87FC761DFFA7FFC0C34FFE71F7A2FE7C" box="[359,423,403,429]" name="Japan" pageId="5" pageNumber="494">Japan</collectingCountry>
(Ryukyu:
<collectingRegion id="3D2FF86FFFA7FFC0C036FE71F486FE7C" box="[542,643,403,429]" country="Japan" name="Okinawa" pageId="5" pageNumber="494">Okinawa</collectingRegion>
Island) (fig. 11).
</paragraph>
</subSubSection>
<subSubSection id="B7F16506FFA7FFC0C2EFFE54F6ECFCC4" pageId="5" pageNumber="494" type="biology_ecology">
<paragraph id="FF54368DFFA7FFC0C2EFFE54F729FDC8" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="494">
<emphasis id="CD9FEA9FFFA7FFC0C2EFFE54F722FE01" bold="true" box="[199,295,438,464]" pageId="5" pageNumber="494">Biology.</emphasis>
This species can be caught by sweeping blossoms and green leaves in the primitive or secondary broad-leaved forest (
<bibRefCitation id="9B7A4B7CFFA7FFC0C3A9FE39F43AFE24" author="Okushima, Y." box="[385,575,474,501]" pageId="5" pageNumber="494" pagination="1 - 383" refId="ref4882" refString="Okushima, Y. (2005) A taxonomic study on the genus Lycocerus (Coleoptera, Cantharidae) from Japan, with zoogeographical considerations. Japanese Journal of Systematic Entomology, Monographic Series, 2, 1 - 383." type="journal article" year="2005">Okushima, 2005</bibRefCitation>
; the authors personal observation). Adults are collected from mid-February to mid-April.
</paragraph>
<paragraph id="FF54368DFFA7FFC0C2EFFDC1F38EFD1C" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="494">
<emphasis id="CD9FEA9FFFA7FFC0C2EFFDC1F404FDEC" bold="true" box="[199,513,547,573]" pageId="5" pageNumber="494">Nomenclatural comments.</emphasis>
<bibRefCitation id="9B7A4B7CFFA7FFC0C020FDC1F49BFDEC" author="Ishida, K." box="[520,670,547,573]" pageId="5" pageNumber="494" pagination="193 - 213" refId="ref4551" refString="Ishida, K. (1986) A revision of the two genera Athemus and Athemellus, of Japan (Coleoptera: Cantharidae). Transactions of the Shikoku Entomological Society, 17, 193 - 213." type="journal article" year="1986">Ishida (1986)</bibRefCitation>
has hitherto been regarded as the original description of this species (
<bibRefCitation id="9B7A4B7CFFA7FFC0C2B7FDA5F7A8FDB0" author="Sato, M. &amp; Okushima, Y." box="[159,429,583,609]" pageId="5" pageNumber="494" pagination="857 - 865" refId="ref5090" refString="Sato, M. &amp; Okushima, Y. (1992) The genus Athemus (Coleoptera, Cantharidae) of the Ryukyu Islands, southwest Japan. Japanese Journal of Entomology, 60, 857 - 865." type="journal article" year="1992">Satô &amp; Okushima, 1992</bibRefCitation>
;
<bibRefCitation id="9B7A4B7CFFA7FFC0C39EFDA5F46AFDB0" author="Takahashi, K." box="[438,623,583,609]" pageId="5" pageNumber="494" pagination="29 - 48" refId="ref5129" refString="Takahashi, K. (1998) Check-list of Cantharidae of Japan. Kanagawa-Chuho, 122, 29 - 48. [in Japanese, with English title]" type="journal article" year="1998">Takahashi, 1998</bibRefCitation>
;
<bibRefCitation id="9B7A4B7CFFA7FFC0C051FDA5F536FDB0" author="Okushima, Y." box="[633,819,582,609]" pageId="5" pageNumber="494" pagination="1 - 383" refId="ref4882" refString="Okushima, Y. (2005) A taxonomic study on the genus Lycocerus (Coleoptera, Cantharidae) from Japan, with zoogeographical considerations. Japanese Journal of Systematic Entomology, Monographic Series, 2, 1 - 383." type="journal article" year="2005">Okushima, 2005</bibRefCitation>
;
<bibRefCitation id="9B7A4B7CFFA7FFC0C115FDA5F28EFDB0" author="Kazantsev, S. &amp; Brancucci, M." box="[829,1163,583,609]" pageId="5" pageNumber="494" refId="ref4627" refString="Kazantsev, S. &amp; Brancucci, M. (2007) Cantharidae. In Lo ¨ bl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 234 - 298." type="book" year="2007">Kazantsev &amp; Brancucci, 2007</bibRefCitation>
), but the name “
<taxonomicName id="38EB4D0EFFA7FFC0C714FDA5F769FD54" authority="Ishida" authorityName="Ishida" class="Insecta" family="Cantharidae" genus="Athemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA7FFC0C714FDA5F718FD54" italics="true" pageId="5" pageNumber="494">Athemus okinawanus</emphasis>
Ishida
</taxonomicName>
” did appear precedently in
<bibRefCitation id="9B7A4B7CFFA7FFC0C08EFD89F52FFD54" author="Sato, M." box="[678,810,618,645]" pageId="5" pageNumber="494" refId="ref5030" refString="Sato, M. (1985) Cantharidae. In: Kurosawa, Y., Hisamatsu, S. &amp; Sasaji, H. (Eds.), The Coleoptera of Japan in Color. Vol. 3. Hoikusha, Osaka, pp. 107 - 119. [in Japanese, with English book title]" type="book" year="1985">Satô (1985)</bibRefCitation>
along with color photograph and a short description in the key. Based on ICZN Article 10 and 13 (
<bibRefCitation id="9B7A4B7CFFA7FFC0C0BCFD6DF35FFD78" author="International Commission on Zoological Nomenclature &amp; Tipografia La" box="[660,1370,655,681]" pageId="5" pageNumber="494" refId="ref4522" refString="International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. Tipografia La Garangola, Padova, xxix + 306 pp." type="book" year="1999">International Commission on Zoological Nomenclature, 1999</bibRefCitation>
), it is considered that the name
<taxonomicName id="38EB4D0EFFA7FFC0C39DFD51F4E8FD1C" authority="Ishida" authorityName="Ishida" box="[437,749,691,717]" class="Insecta" family="Cantharidae" genus="Athemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="okinawanus">
<emphasis id="CD9FEA9FFFA7FFC0C39DFD51F4A6FD1C" box="[437,675,691,717]" italics="true" pageId="5" pageNumber="494">Athemus okinawanus</emphasis>
Ishida
</taxonomicName>
was available at the time of the publication by
<bibRefCitation id="9B7A4B7CFFA7FFC0C728FD51F38DFD1C" author="Sato, M." box="[1280,1416,690,717]" pageId="5" pageNumber="494" refId="ref5030" refString="Sato, M. (1985) Cantharidae. In: Kurosawa, Y., Hisamatsu, S. &amp; Sasaji, H. (Eds.), The Coleoptera of Japan in Color. Vol. 3. Hoikusha, Osaka, pp. 107 - 119. [in Japanese, with English book title]" type="book" year="1985">Satô (1985)</bibRefCitation>
.
</paragraph>
<paragraph id="FF54368DFFA7FFC0C2EFFD35F6ECFCC4" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="494">
<emphasis id="CD9FEA9FFFA7FFC0C2EFFD35F703FD20" bold="true" box="[199,262,727,753]" pageId="5" pageNumber="494">Note.</emphasis>
This species is placed in the
<taxonomicName id="38EB4D0EFFA7FFC0C05AFD35F554FD21" box="[626,849,727,752]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="aegrotus">
<emphasis id="CD9FEA9FFFA7FFC0C05AFD35F554FD21" box="[626,849,727,752]" italics="true" pageId="5" pageNumber="494">Lycocerus aegrotus</emphasis>
</taxonomicName>
subgroup of the
<taxonomicName id="38EB4D0EFFA7FFC0C60CFD35F2A2FD21" box="[1060,1191,727,752]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="494" phylum="Arthropoda" rank="species" species="aegrotus">
<emphasis id="CD9FEA9FFFA7FFC0C60CFD35F2A2FD21" box="[1060,1191,727,752]" italics="true" pageId="5" pageNumber="494">L. aegrotus</emphasis>
</taxonomicName>
group by
<bibRefCitation id="9B7A4B7CFFA7FFC0C701FD35F6E0FCC4" author="Okushima, Y." pageId="5" pageNumber="494" pagination="1 - 383" refId="ref4882" refString="Okushima, Y. (2005) A taxonomic study on the genus Lycocerus (Coleoptera, Cantharidae) from Japan, with zoogeographical considerations. Japanese Journal of Systematic Entomology, Monographic Series, 2, 1 - 383." type="journal article" year="2005">Okushima (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>