182 lines
22 KiB
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182 lines
22 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1080.76876" ID-GBIF-Dataset="fd3ac82d-e15e-4f72-b60d-ea3e1e54f50f" ID-PMC="PMC8752575" ID-Pensoft-Pub="1313-2970-1080-1" ID-Pensoft-UUID="57FCF8054EB55191B95D363607C2F202" ID-PubMed="35068962" ModsDocID="1313-2970-1080-1" checkinTime="1641398968081" checkinUser="pensoft" docAuthor="Songsangchote, Chaowalit, Sippawat, Zongtum, Khaikaew, Wuttikrai & Chomphuphuang, Narin" docDate="2022" docId="2AA74850A8ED5C59AF201172A240B263" docLanguage="en" docName="ZooKeys 1080: 1-19" docOrigin="ZooKeys 1080" docPubDate="2022-01-04" docSource="http://dx.doi.org/10.3897/zookeys.1080.76876" docTitle="Taksinus Songsangchote, Sippawat, Khaikaew & Chomphuphuang 2022, gen. nov." docType="treatment" docVersion="4" id="57FCF8054EB55191B95D363607C2F202" lastPageNumber="1" masterDocId="57FCF8054EB55191B95D363607C2F202" masterDocTitle="A new genus of bamboo culm tarantula from Thailand (Araneae, Mygalomorphae, Theraphosidae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="1" updateTime="1668151279985" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus of bamboo culm tarantula from Thailand (Araneae, Mygalomorphae, Theraphosidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Songsangchote, Chaowalit</mods:namePart>
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<mods:affiliation>Faculty of Forestry, Kasetsart University, Bangkok, 10900, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sippawat, Zongtum</mods:namePart>
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<mods:affiliation>160 village no. 7, Mae Tho, Mueang Tak district, Tak, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Khaikaew, Wuttikrai</mods:namePart>
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<mods:affiliation>5 village no. 8, Angthong sub-district, Chiang Kham district, Phayao, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chomphuphuang, Narin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0738-3879</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology and Plant Pathology, Faculty of Agriculture, Khon Kaen University, Khon Kaen, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">narich@kku.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-01-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1080</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>19</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1080.76876</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1080.76876</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1080-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">57FCF8054EB55191B95D363607C2F202</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="191887085" LSID="urn:lsid:zoobank.org:act:AB5E52BE-415F-415E-91AC-1155EA142B1D" httpUri="http://treatment.plazi.org/id/2AA74850A8ED5C59AF201172A240B263" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/AB5E52BE-415F-415E-91AC-1155EA142B1D" authority="Songsangchote, Sippawat, Khaikaew & Chomphuphuang" authorityName="Songsangchote, Sippawat, Khaikaew & Chomphuphuang" authorityYear="2022" family="Theraphosidae" genus="Taksinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taksinus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">Taksinus Songsangchote, Sippawat, Khaikaew & Chomphuphuang</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type species">
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<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Songsangchote, Sippawat, Khaikaew & Chomphuphuang" authorityYear="2022" family="Theraphosidae" genus="Taksinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taksinus bambus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bambus">
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<emphasis italics="true" pageId="0" pageNumber="1">Taksinus bambus</emphasis>
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</taxonomicName>
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Songsangchote, Sippawat, Khaikaew & Chomphuphuang, 2021 from Tak, Thailand.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The characteristics of
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<taxonomicName authorityName="Songsangchote, Sippawat, Khaikaew & Chomphuphuang" authorityYear="2022" family="Theraphosidae" genus="Taksinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taksinus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Taksinus</emphasis>
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</taxonomicName>
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gen. nov. that differ from
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<taxonomicName authorityName="Pocock" authorityYear="1892" class="Arachnida" family="Theraphosidae" genus="Ornithoctonus" kingdom="Animalia" lsidName="Ornithoctonus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Ornithoctonus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Simon" authorityYear="1887" class="Arachnida" family="Theraphosidae" genus="Cyriopagopus" kingdom="Animalia" lsidName="Cyriopagopus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Cyriopagopus</emphasis>
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</taxonomicName>
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are: a low caput, a clypeus that is less than the width of the median ocular quadrangle (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Taksinus bambus sp. nov. Paratype ♀ TAK 3 (A, B, D) Paratype ♀ TAK 2 (C). A carapace, dorsal view B sternum, labium, maxilla, and coxae, ventral view C maxilla, prolateral view D maxillae and labium with cuspules. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1080.76876.figure6" httpUri="https://binary.pensoft.net/fig/629895" pageId="0" pageNumber="1">6A</figureCitation>
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), and spermathecae with twin seminal receptacles (Fig.
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<figureCitation captionStart="Figures 7" captionStartId="F7" captionText="Figures 7. Taksinus bambus sp. nov. paratype ♀ TAK 2 (A-E) paratype ♀ TAK 3 (F). A chelicerae, prolateral view B chelicerae, retrolateral view C plumose hairs outer chelicerae, retrolateral view D chelicerae strikers, retrolateral view E spermathecae, dorsal view F spermathecae, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1080.76876.figures7" httpUri="https://binary.pensoft.net/fig/629896" pageId="0" pageNumber="1">7E, F</figureCitation>
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) (
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<bibRefCitation author="Raven, R" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="1" pagination="1 - 180" refId="B6" refString="Raven, R, 1985. The spider infraorder Mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History 182: 1 - 180" title="The spider infraorder Mygalomorphae (Araneae): cladistics and systematics." volume="182" year="1985">Raven 1985</bibRefCitation>
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;
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<bibRefCitation author="von Wirth, V" journalOrPublisher="Arthropoda" pageId="0" pageNumber="1" pagination="2 - 27" refId="B11" refString="von Wirth, V, Striffler, BF, 2005. Neue Erkenntnisse zur Vogelspinnen-Unterfamilie Ornithoctoninae, mit Beschreibung von Ornithoctonus aureotibialis sp. n. und Haplopelma longipes sp. n. (Araneae, Theraphosidae). Arthropoda 13: 2 - 27" title="Neue Erkenntnisse zur Vogelspinnen-Unterfamilie Ornithoctoninae, mit Beschreibung von Ornithoctonus aureotibialis sp. n. und Haplopelma longipes sp. n. (Araneae, Theraphosidae)." volume="13" year="2005">von Wirth and Striffler 2005</bibRefCitation>
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;
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<bibRefCitation author="Smith, AM" journalOrPublisher="British Tarantula Society Journal" pageId="0" pageNumber="1" pagination="25 - 48" refId="B9" refString="Smith, AM, Jacobi, MA, 2015. Revision of the genus Phormingochilus with the description of three new species from Sulawesi and Sarawak and notes on the placement of the genera Cyriopagopus, Lampropelma and Omothymus. British. British Tarantula Society Journal 30: 25 - 48" title="Revision of the genus Phormingochilus with the description of three new species from Sulawesi and Sarawak and notes on the placement of the genera Cyriopagopus, Lampropelma and Omothymus. British." volume="30" year="2015">Smith and Jacobi 2015</bibRefCitation>
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). The new genus differs from
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<taxonomicName authorityName="Pocock" authorityYear="1895" class="Arachnida" family="Theraphosidae" genus="Citharognathus" kingdom="Animalia" lsidName="Citharognathus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Citharognathus</emphasis>
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</taxonomicName>
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by the lack of incrassate tibia and metatarsus IV.
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<taxonomicName authorityName="Songsangchote, Sippawat, Khaikaew & Chomphuphuang" authorityYear="2022" family="Theraphosidae" genus="Taksinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taksinus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Taksinus</emphasis>
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</taxonomicName>
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gen. nov. differs from
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<taxonomicName authorityName="Simon" authorityYear="1892" class="Arachnida" family="Theraphosidae" genus="Lampropelma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lampropelma" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Lampropelma</emphasis>
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</taxonomicName>
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by the absence of a dense brush of hair on the retrolateral side of the femora of the front limbs (
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<bibRefCitation author="von Wirth, V" journalOrPublisher="Arthropoda" pageId="0" pageNumber="1" pagination="2 - 27" refId="B11" refString="von Wirth, V, Striffler, BF, 2005. Neue Erkenntnisse zur Vogelspinnen-Unterfamilie Ornithoctoninae, mit Beschreibung von Ornithoctonus aureotibialis sp. n. und Haplopelma longipes sp. n. (Araneae, Theraphosidae). Arthropoda 13: 2 - 27" title="Neue Erkenntnisse zur Vogelspinnen-Unterfamilie Ornithoctoninae, mit Beschreibung von Ornithoctonus aureotibialis sp. n. und Haplopelma longipes sp. n. (Araneae, Theraphosidae)." volume="13" year="2005">von Wirth and Striffler 2005</bibRefCitation>
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) and males by lack of apical embolus swelling (Fig.
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<figureCitation captionStart="Figures 5" captionStartId="F5" captionText="Figures 5. Taksinus bambus sp. nov. holotype ♂ TAK 1 (A-E) and Omothymus sp. ♂ (F-J). Palpal bulb A prolateral view B retrolateral view C dorsal view D ventral view E apical view F prolateral view G retrolateral view H dorsal view I ventral view J apical view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1080.76876.figures5" httpUri="https://binary.pensoft.net/fig/629894" pageId="0" pageNumber="1">5A-E</figureCitation>
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; see
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<bibRefCitation DOI="https://doi.org/10.13156/arac.2018.18.2.137" author="Gabriel, R" journalOrPublisher="Arachnology" pageId="0" pageNumber="1" pagination="137 - 147" refId="B3" refString="Gabriel, R, Sherwood, D, 2019. The revised taxonomic placement of some arboreal Ornithoctoninae Pocock, 1895 with description of a new species of Omothymus Thorell, 1891 (Araneae: Theraphosidae). Arachnology 18 (2): 137 - 147, DOI: https://doi.org/10.13156/arac.2018.18.2.137" title="The revised taxonomic placement of some arboreal Ornithoctoninae Pocock, 1895 with description of a new species of Omothymus Thorell, 1891 (Araneae: Theraphosidae)." url="https://doi.org/10.13156/arac.2018.18.2.137" volume="18" year="2019">Gabriel and Sherwood 2019</bibRefCitation>
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: 143, figs 17, 18).
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<taxonomicName authorityName="Songsangchote, Sippawat, Khaikaew & Chomphuphuang" authorityYear="2022" family="Theraphosidae" genus="Taksinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taksinus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Taksinus</emphasis>
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</taxonomicName>
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gen. nov. can be distinguished from
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<taxonomicName authorityName="Thorell" authorityYear="1891" class="Arachnida" family="Theraphosidae" genus="Omothymus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omothymus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Omothymus</emphasis>
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</taxonomicName>
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by male palpal bulb with a gently curved embolus with rounded embolic apex (Fig.
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<figureCitation captionStart="Figures 5" captionStartId="F5" captionText="Figures 5. Taksinus bambus sp. nov. holotype ♂ TAK 1 (A-E) and Omothymus sp. ♂ (F-J). Palpal bulb A prolateral view B retrolateral view C dorsal view D ventral view E apical view F prolateral view G retrolateral view H dorsal view I ventral view J apical view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1080.76876.figures5" httpUri="https://binary.pensoft.net/fig/629894" pageId="0" pageNumber="1">5A-E</figureCitation>
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) vs palpal bulb steep angle embolus and apex with a sharp point in
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<taxonomicName authorityName="Thorell" authorityYear="1891" class="Arachnida" family="Theraphosidae" genus="Omothymus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omothymus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Omothymus</emphasis>
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</taxonomicName>
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(Fig.
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<figureCitation captionStart="Figures 5" captionStartId="F5" captionText="Figures 5. Taksinus bambus sp. nov. holotype ♂ TAK 1 (A-E) and Omothymus sp. ♂ (F-J). Palpal bulb A prolateral view B retrolateral view C dorsal view D ventral view E apical view F prolateral view G retrolateral view H dorsal view I ventral view J apical view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1080.76876.figures5" httpUri="https://binary.pensoft.net/fig/629894" pageId="0" pageNumber="1">5F-J</figureCitation>
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; see
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<bibRefCitation DOI="https://doi.org/10.13156/arac.2018.18.2.137" author="Gabriel, R" journalOrPublisher="Arachnology" pageId="0" pageNumber="1" pagination="137 - 147" refId="B3" refString="Gabriel, R, Sherwood, D, 2019. The revised taxonomic placement of some arboreal Ornithoctoninae Pocock, 1895 with description of a new species of Omothymus Thorell, 1891 (Araneae: Theraphosidae). Arachnology 18 (2): 137 - 147, DOI: https://doi.org/10.13156/arac.2018.18.2.137" title="The revised taxonomic placement of some arboreal Ornithoctoninae Pocock, 1895 with description of a new species of Omothymus Thorell, 1891 (Araneae: Theraphosidae)." url="https://doi.org/10.13156/arac.2018.18.2.137" volume="18" year="2019">Gabriel and Sherwood 2019</bibRefCitation>
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: 139, figs 1-5).
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<taxonomicName authorityName="Songsangchote, Sippawat, Khaikaew & Chomphuphuang" authorityYear="2022" family="Theraphosidae" genus="Taksinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taksinus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Taksinus</emphasis>
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</taxonomicName>
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gen. nov. differs from
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<taxonomicName authorityName="Pocock" authorityYear="1895" class="Arachnida" family="Theraphosidae" genus="Phormingochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phormingochilus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Phormingochilus</emphasis>
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</taxonomicName>
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by the lack of a single megaspine on the inside of the male tibial apophyses (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Taksinus bambus sp. nov. Holotype ♂ TAK 1. Tibial apophysis A ventral view B apical view C prolateral view D dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1080.76876.figure4" httpUri="https://binary.pensoft.net/fig/629893" pageId="0" pageNumber="1">4A, B</figureCitation>
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; see
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<bibRefCitation author="Smith, AM" journalOrPublisher="British Tarantula Society Journal" pageId="0" pageNumber="1" pagination="25 - 48" refId="B9" refString="Smith, AM, Jacobi, MA, 2015. Revision of the genus Phormingochilus with the description of three new species from Sulawesi and Sarawak and notes on the placement of the genera Cyriopagopus, Lampropelma and Omothymus. British. British Tarantula Society Journal 30: 25 - 48" title="Revision of the genus Phormingochilus with the description of three new species from Sulawesi and Sarawak and notes on the placement of the genera Cyriopagopus, Lampropelma and Omothymus. British." volume="30" year="2015">Smith and Jacobi 2015</bibRefCitation>
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: 41, fig. 38;
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<bibRefCitation DOI="https://doi.org/10.13156/arac.2018.18.2.137" author="Gabriel, R" journalOrPublisher="Arachnology" pageId="0" pageNumber="1" pagination="137 - 147" refId="B3" refString="Gabriel, R, Sherwood, D, 2019. The revised taxonomic placement of some arboreal Ornithoctoninae Pocock, 1895 with description of a new species of Omothymus Thorell, 1891 (Araneae: Theraphosidae). Arachnology 18 (2): 137 - 147, DOI: https://doi.org/10.13156/arac.2018.18.2.137" title="The revised taxonomic placement of some arboreal Ornithoctoninae Pocock, 1895 with description of a new species of Omothymus Thorell, 1891 (Araneae: Theraphosidae)." url="https://doi.org/10.13156/arac.2018.18.2.137" volume="18" year="2019">Gabriel and Sherwood 2019</bibRefCitation>
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: 142, figs 14-16), a short embolus compared to palpal bulb length (1:1) (Fig.
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<figureCitation captionStart="Figures 5" captionStartId="F5" captionText="Figures 5. Taksinus bambus sp. nov. holotype ♂ TAK 1 (A-E) and Omothymus sp. ♂ (F-J). Palpal bulb A prolateral view B retrolateral view C dorsal view D ventral view E apical view F prolateral view G retrolateral view H dorsal view I ventral view J apical view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1080.76876.figures5" httpUri="https://binary.pensoft.net/fig/629894" pageId="0" pageNumber="1">5A-E</figureCitation>
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), and the geographic distribution of
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<taxonomicName authorityName="Pocock" authorityYear="1895" class="Arachnida" family="Theraphosidae" genus="Phormingochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phormingochilus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Phormingochilus</emphasis>
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</taxonomicName>
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currently restricted to Borneo.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The generic name was named Phraya Tak (governor of Tak province), which is in honor of Taksin the Great, king of the Thonburi Kingdom, in commemoration of his early career.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Carapace longer than wide, low caput. Fovea deep, straight (males) or slightly procurved (females). Clypeus short, less than width of median ocular quadrangle in males and females. Eight eyes arranged on tubercle, anterior eye row slightly procurved and the posterior row straight. Outer cheliceral on lower surface from margin with five slightly curved pad of plumose setae on the retrolateral chelicerae. Maxillae longer than wide with>155 cuspules (male) or 149-183 (females), two horizontal rows of 10-11 stout thorn-like spines on the lower half of prolateral maxillae (below suture) and one horizontal row of six stout thorn-like spines on the upper half of prolateral maxillae (above suture). Spines of varying lengths, with the longest being at the top of the series; combined to form a stridulating organ. Labium wider than long, with 75 cuspules (male) or 125 (females). Sternum longer than wide, with two pairs of ovoid sigillae; Posterior sigilla is significantly remote from the edge, middle sigilla is close to the margin, and anterior sigilla is indistinguishable. Legs: formula 1423 (males);
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<normalizedToken originalValue="±">+/-</normalizedToken>
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Total lengths of legs I and IV = 0.48, 4123 (females)
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<normalizedToken originalValue="±">+/-</normalizedToken>
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Total lengths of legs I and IV = 2.41-3.33, RF = 101 (males) or 90.6-93 (females). Scopulae distinct, thickly set on tarsus; ventral surface not divided. Tibial spur capped with multitude of thin, short black spines, with no single megaspine on the inside of the tibial apophyses. Palpal bulb is ellipsoid and partly concave, embolus short compared to palpal bulb length (1:1), moderately curved, rounded apex, with single retrolateral keel. Spermathecae have twin seminal receptacles, rounded tombstone receptacles, fused in the basal region.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Tak province, Thailand</paragraph>
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</subSubSection>
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</treatment>
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</document> |