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<document id="E02F8966E01DBFC56896B4CE15C7A007" ID-CLB-Dataset="22345" ID-DOI="10.5852/ejt.2015.161" ID-GBIF-Dataset="8d8f2463-08b8-425f-ae32-139eb62f3056" ID-ISSN="2118-9773" ID-Zenodo-Dep="3805619" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588784778089" checkinUser="carolina" docAuthor="Reboleira, Ana Sofia P. S., Gonçalves, Fernando, Oromí, Pedro &amp; Taiti, Stefano" docDate="2015" docId="2A668781C410FFC8F2C8FB8BFD345AC9" docLanguage="en" docName="ejt-2015-161.pdf.imf" docOrigin="European Journal of Taxonomy 161" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Troglonethes arrabidaensis Reboleira &amp; Gonçalves &amp; Oromí &amp; Taiti 2015, sp. nov." docType="treatment" docUuid="78A12550-F7CA-4892-9E90-7F85446423D3" docUuidSource="ZooBank" docVersion="9" lastPageNumber="28" masterDocId="D65FFFF9C408FFD3F312FFD4FF875873" masterDocTitle="The cavernicolous Oniscidea (Crustacea: Isopoda) of Portugal" masterLastPageNumber="61" masterPageNumber="1" pageNumber="25" updateTime="1698838751992" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="84F6DF2273463359A16D261F1D6608CA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1E0D5BE06347408D296CAD4F01FE6770">
<mods:title id="A8BEA4DA8D5CE1988150A99C92F90C8D">The cavernicolous Oniscidea (Crustacea: Isopoda) of Portugal</mods:title>
</mods:titleInfo>
<mods:name id="6C0BEAB38B64C1B65F9A85158B595BA9" type="personal">
<mods:role id="3DCEF341F47DB2D8628C300155D5A8A5">
<mods:roleTerm id="3665B03B83EBED1C13160006E5EBB9BC">Author</mods:roleTerm>
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<mods:namePart id="C4FD74C573EA6CD33FF7DD6AC997D3AF">Reboleira, Ana Sofia P. S.</mods:namePart>
<mods:nameIdentifier id="C8F73B809391B806C549773703610542" type="LSID">338DE845-4839-4EF5-B684-587C021F076C</mods:nameIdentifier>
<mods:affiliation id="8E5D4EDA9A31CE8B61F9DB24ABD34528">Departamento de Biologia &amp; CESAM, Universidade de Aveiro, Campus Universitário de Santiago, 3810 - 193 Aveiro, Portugal. &amp; Departamento de Biología Animal, Facultad de Biología, Universidad de La Laguna, 38206 La Laguna, Tenerife, Islas Canarias, Spain. &amp; Zoological Museum, Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, 2100 Copenhagen Ø, Denmark. &amp; urn: lsid: zoobank. org: author: 338 DE 845 - 4839 - 4 EF 5 - B 684 - 587 C 021 F 076 C &amp; Corresponding author: sreboleira @ ua. pt</mods:affiliation>
<mods:nameIdentifier id="AC6B4DB95DD88A1831E1BA7770A85C01" type="email">sreboleira@ua.pt</mods:nameIdentifier>
</mods:name>
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<mods:role id="61975D6185E0F8086D6EA2630EC80EBE">
<mods:roleTerm id="F7812993A7D75FDE096E9BE6EF48962B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F6B15095F134D136271989F8F6F0573A">Gonçalves, Fernando</mods:namePart>
<mods:nameIdentifier id="D1403EB687A9FB9398EC5ACEDA1EEAF6" type="LSID">4B8A2E04-3543-465F-9949-B7E1745DA931</mods:nameIdentifier>
<mods:affiliation id="935BC7B78F890126FA6498B1BEBDBC46">Departamento de Biologia &amp; CESAM, Universidade de Aveiro, Campus Universitário de Santiago, 3810 - 193 Aveiro, Portugal. &amp; Email: fjmg @ ua. pt &amp; urn: lsid: zoobank. org: author: 4 B 8 A 2 E 04 - 3543 - 465 F- 9949 - B 7 E 1745 DA 931</mods:affiliation>
<mods:nameIdentifier id="107B50EC17F3CA6FEFE50039AC6BED29" type="email">fjmg@ua.pt</mods:nameIdentifier>
</mods:name>
<mods:name id="587D914FCC6905E0FE08062191A8F50B" type="personal">
<mods:role id="6710F3C356CC152D76608C114D8671EE">
<mods:roleTerm id="30F720360FDFFC2FFCDFC018D4BB5C69">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4C0E705677C33F71965926CED80B0CBA">Oromí, Pedro</mods:namePart>
<mods:nameIdentifier id="70241D02F30A82D005E91C4349664581" type="LSID">00F2ACC4-308B-4BEE-BBA4-DE6AC8905449</mods:nameIdentifier>
<mods:affiliation id="F802C1BC7459A2CB7C26AC3CBC31C6C3">Departamento de Biología Animal, Facultad de Biología, Universidad de La Laguna, 38206 La Laguna, Tenerife, Islas Canarias, Spain. &amp; Email: poromi @ ull. es &amp; urn: lsid: zoobank. org: author: 00 F 2 ACC 4 - 308 B- 4 BEE-BBA 4 - DE 6 AC 8905449</mods:affiliation>
<mods:nameIdentifier id="5B396BB50DB2F57440627E5DE496C261" type="email">poromi@ull.es</mods:nameIdentifier>
</mods:name>
<mods:name id="529ACA3DDFE55A0A097CB74A68D9E91E" type="personal">
<mods:role id="FA673EB581C1C0B730E91BD7CEFDADD4">
<mods:roleTerm id="CE9C7A46C3F2DE04C80177D3904DCA85">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CEE71001A9BFCFEEDCC0460943850956">Taiti, Stefano</mods:namePart>
<mods:nameIdentifier id="7132E2248A0994B8E874DB785DB3D1B7" type="LSID">62E97059-6AE5-4984-9ABB-7FB6F7358BD6</mods:nameIdentifier>
<mods:affiliation id="4AC899AE8F3755DF781CE3ED4108002C">Istituto per lo Studio degli Ecosistemi, Consiglio Nazionale delle Ricerche, Via Madonna del Piano 10, 50019 Sesto Fiorentino (Firenze), Italy. &amp; Email: stefano. taiti @ ise. cnr. it &amp; urn: lsid: zoobank. org: author: 62 E 97059 - 6 AE 5 - 4984 - 9 ABB- 7 FB 6 F 7358 BD 6</mods:affiliation>
<mods:nameIdentifier id="8186678137FBB3D69B67D27530391D8A" type="email">stefano.taiti@ise.cnr.it</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="279A26AB0E18715E8C8F0455442A20A0">text</mods:typeOfResource>
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<mods:title id="32254F9982C5EF89A9B431955852D672">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="9790A0E15A8BA771C8A570203F9BA105">
<mods:date id="8FFB32AFFD4960472E0DEF03B89EA4DE">2015</mods:date>
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<mods:number id="2032A958A4B47E910B68482721B8E227">2015-12-11</mods:number>
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<mods:number id="6FC7E04EB7FC414971BD8D7473E3297F">161</mods:number>
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<mods:start id="125781B9D7714D7056627354361C8CDD">1</mods:start>
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<mods:classification id="980921030E32BC67ABD6172645312080">journal article</mods:classification>
<mods:identifier id="24276A1EE2ED7BE0990CADE42637E3BD" type="CLB-Dataset">22345</mods:identifier>
<mods:identifier id="13CA927A560C3D9A5B17682CB087B41A" type="DOI">10.5852/ejt.2015.161</mods:identifier>
<mods:identifier id="1E7D3EAE65826CB971B7CFC71866F8AB" type="GBIF-Dataset">8d8f2463-08b8-425f-ae32-139eb62f3056</mods:identifier>
<mods:identifier id="EEE4A7B21AD2348F8B9CFD8F99996A9E" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="D328156872001396C3A5BFE46D86EA06" type="Zenodo-Dep">3805619</mods:identifier>
<mods:identifier id="06E209AC822AB45406CEA2582B2621B3" type="ZooBank">2297E4A3-D279-4D0A-923C-D5E0D5DCB3C0</mods:identifier>
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<treatment id="2A668781C410FFC8F2C8FB8BFD345AC9" ID-DOI="http://doi.org/10.5281/zenodo.3815799" ID-GBIF-Taxon="163970029" ID-Zenodo-Dep="3815799" LSID="urn:lsid:zoobank.org:act:78A12550-F7CA-4892-9E90-7F85446423D3" httpUri="http://treatment.plazi.org/id/2A668781C410FFC8F2C8FB8BFD345AC9" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25">
<subSubSection id="EAD5651CC410FFCBF2C8FB8BFBDD5C09" box="[474,1114,1119,1146]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="A2703697C410FFCBF2C8FB8BFBDD5C09" blockId="24.[377,1210,1119,1216]" box="[474,1114,1119,1146]" pageId="24" pageNumber="25">
<heading id="F93881FBC410FFCBF2C8FB8BFBDD5C09" box="[474,1114,1119,1146]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="65CF4D14C410FFCBF2C8FB8BFC7F5C09" ID-CoL="598ZR" authority="Reboleira &amp; Taiti" authorityName="Reboleira &amp; Gonçalves &amp; Oromí &amp; Taiti" authorityYear="2015" box="[474,1016,1119,1146]" class="Malacostraca" family="Trichoniscidae" genus="Troglonethes" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="arrabidaensis" status="sp. nov.">
<emphasis id="90BBEA85C410FFCBF2C8FB8BFCA25C09" bold="true" box="[474,805,1119,1146]" italics="true" pageId="24" pageNumber="25">Troglonethes arrabidaensis</emphasis>
Reboleira &amp; Taiti
</taxonomicName>
<taxonomicNameLabel id="8B8857FEC410FFCBF0EDFBB4FBDD5C09" box="[1023,1114,1120,1146]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EAD5651CC410FFCBF26BFB57FC135CB3" pageId="24" pageNumber="25" type="description">
<paragraph id="A2703697C410FFCBF26BFB57FB3D5CED" blockId="24.[377,1210,1119,1216]" box="[377,1210,1155,1182]" pageId="24" pageNumber="25">
<uri id="D65E3A95C410FFCBF26BFB57FB3D5CED" box="[377,1210,1155,1182]" pageId="24" pageNumber="25">
urn:lsid:zoobank.org:act:
<uuid id="D6690C42C410FFCBF185FB57FB3D5CED" box="[663,1210,1155,1182]" pageId="24" pageNumber="25">78A12550-F7CA-4892-9E90-7F85446423D3</uuid>
</uri>
</paragraph>
<paragraph id="A2703697C410FFCBF18DFB72FC135CB3" blockId="24.[377,1210,1119,1216]" box="[671,916,1190,1216]" pageId="24" pageNumber="25">
<figureCitation id="3AF42A12C410FFCBF18DFB72FD725CB3" box="[671,757,1190,1216]" captionStart="Fig" captionStartId="25.[189,232,1873,1899]" captionTargetBox="[189,1405,449,1839]" captionTargetId="figure@25.[180,1411,438,1847]" captionTargetPageId="25" captionText="Fig. 16. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida. — Paratype, ♀. A. Adult specimen in dorsal view. B. Dorsal scale-seta. C. Cephalon in frontal view. D. Cephalon in dorsal view. E. Pleonites 4, 5, telson and left uropod. — Paratype, ♂. F. Antennula. G. Antenna." figureDoi="http://doi.org/10.5281/zenodo.3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="24" pageNumber="25">Figs 16</figureCitation>
A–G, 17A–D
</paragraph>
</subSubSection>
<subSubSection id="EAD5651CC410FFCBF3AFFB3CFBCC5D04" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="A2703697C410FFCBF3AFFB3CFEB15D71" blockId="24.[189,310,1256,1282]" box="[189,310,1256,1282]" pageId="24" pageNumber="25">
<heading id="F93881FBC410FFCBF3AFFB3CFEB15D71" bold="true" box="[189,310,1256,1282]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="90BBEA85C410FFCBF3AFFB3CFEB15D71" bold="true" box="[189,310,1256,1282]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A2703697C410FFCBF3AFFAC2FBCC5D04" blockId="24.[189,1399,1302,1399]" pageId="24" pageNumber="25">
A species of
<taxonomicName id="65CF4D14C410FFCBF24AFAC2FE775D43" authorityName="Cruz" authorityYear="1989" box="[344,496,1302,1328]" class="Malacostraca" family="Trichoniscidae" genus="Troglonethes" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="90BBEA85C410FFCBF24AFAC2FE775D43" box="[344,496,1302,1328]" italics="true" pageId="24" pageNumber="25">Troglonethes</emphasis>
</taxonomicName>
characterised by the antenna having three flagellar articles, the male pereopod 7 carpus enlarged in the basal part, the pleopod 1 exopod triangular, as wide as long, and the male pleopod 2 endopod with the distal article bearing an apical hook-like process.
</paragraph>
</subSubSection>
<subSubSection id="EAD5651CC410FFCBF3AFFA4AFBAC5D94" box="[189,1067,1438,1511]" pageId="24" pageNumber="25" type="etymology">
<paragraph id="A2703697C410FFCBF3AFFA4AFEC35DCB" blockId="24.[189,324,1438,1464]" box="[189,324,1438,1464]" pageId="24" pageNumber="25">
<heading id="F93881FBC410FFCBF3AFFA4AFEC35DCB" bold="true" box="[189,324,1438,1464]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="90BBEA85C410FFCBF3AFFA4AFEC35DCB" bold="true" box="[189,324,1438,1464]" pageId="24" pageNumber="25">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="A2703697C410FFCBF3AFFA19FBAC5D94" blockId="24.[189,1067,1485,1511]" box="[189,1067,1485,1511]" pageId="24" pageNumber="25">
The new species is named after the
<typeStatus id="7D748835C410FFCBF14DFA19FD155D94" box="[607,658,1485,1511]" pageId="24" pageNumber="25">type</typeStatus>
locality, the Arrábida karst massif.
</paragraph>
</subSubSection>
<subSubSection id="EAD5651CC410FFCBF3AFF9DAFBCB5EAC" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="A2703697C410FFCBF3AFF9DAFE2D5E5B" blockId="24.[189,426,1550,1576]" box="[189,426,1550,1576]" pageId="24" pageNumber="25">
<heading id="F93881FBC410FFCBF3AFF9DAFE2D5E5B" bold="true" box="[189,426,1550,1576]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="90BBEA85C410FFCBF3AFF9DAFE2D5E5B" bold="true" box="[189,426,1550,1576]" pageId="24" pageNumber="25">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="12A73CCAC410FFCBF3F8F9E9FBAB5E09" ID-GBIF-Occurrence="2610630304" collectingDate="2013-08-16" collectionCode="MZUF" country="Portugal" location="Gruta do Frade" municipality="Arrabida Massif" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="A2703697C410FFCBF3F8F9E9FEDB5E24" blockId="24.[189,1068,1597,1658]" box="[234,348,1597,1623]" pageId="24" pageNumber="25">
<typeStatus id="7D748835C410FFCBF3F8F9E9FEDB5E24" box="[234,348,1597,1623]" pageId="24" pageNumber="25" type="holotype">
<emphasis id="90BBEA85C410FFCBF3F8F9E9FEDB5E24" bold="true" box="[234,348,1597,1623]" pageId="24" pageNumber="25">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="A2703697C410FFCBF3AFF9B4FBAB5E09" blockId="24.[189,1068,1597,1658]" box="[189,1068,1632,1658]" pageId="24" pageNumber="25">
<collectingCountry id="DAD87607C410FFCBF3AFF9B4FEDA5E09" box="[189,349,1632,1658]" name="Portugal" pageId="24" pageNumber="25">PORTUGAL</collectingCountry>
:
<specimenCount id="B4C9FD1EC410FFCBF275F9B4FEF85E09" box="[359,383,1632,1658]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
,
<location id="A710604CC410FFCBF29EF9B4FDC65E09" LSID="urn:lsid:plazi:treatment:2A668781C410FFC8F2C8FB8BFD345AC9:A710604CC410FFCBF29EF9B4FDC65E09" box="[396,577,1632,1658]" country="Portugal" municipality="Arrabida Massif" name="Gruta do Frade" pageId="24" pageNumber="25">Gruta do Frade</location>
,
<collectingMunicipality id="4214ACEDC410FFCBF158F9B4FC8A5E09" box="[586,781,1632,1658]" pageId="24" pageNumber="25">Arrábida Massif</collectingMunicipality>
,
<collectingDate id="C635E9BFC410FFCBF00BF9B4FC3F5E09" box="[793,952,1632,1658]" pageId="24" pageNumber="25" value="2013-08-16">
<date id="D6711057C410FFCBF00BF9B4FC3F5E09" box="[793,952,1632,1658]" pageId="24" pageNumber="25" value="2013-08-16">16 Aug. 2013</date>
</collectingDate>
(
<collectionCode id="C4DEAE52C410FFCBF0D5F9B4FB985E09" box="[967,1055,1632,1658]" collectionName="Italy, Firenze, Museo Zoologico &quot;La Specola&quot;" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="24" pageNumber="25">MZUF</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="A2703697C410FFCBF3F8F975FEE05EC8" blockId="24.[189,1100,1697,1759]" box="[234,359,1697,1723]" pageId="24" pageNumber="25">
<typeStatus id="7D748835C410FFCBF3F8F975FEE05EC8" box="[234,359,1697,1723]" pageId="24" pageNumber="25" type="paratype">
<emphasis id="90BBEA85C410FFCBF3F8F975FEE05EC8" bold="true" box="[234,359,1697,1723]" pageId="24" pageNumber="25">Paratypes</emphasis>
</typeStatus>
</paragraph>
<paragraph id="A2703697C410FFCBF3AFF911FBCB5EAC" blockId="24.[189,1100,1697,1759]" box="[189,1100,1732,1759]" pageId="24" pageNumber="25">
<collectingCountry id="DAD87607C410FFCBF3AFF911FEDA5EAC" box="[189,349,1733,1759]" name="Portugal" pageId="24" pageNumber="25">PORTUGAL</collectingCountry>
:
<specimenCount id="B4C9FD1EC410FFCBF275F911FE2C5EAC" box="[359,427,1733,1759]" pageId="24" pageNumber="25" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="B4C9FD1EC410FFCBF2AAF911FD8E5EAC" box="[440,521,1733,1759]" pageId="24" pageNumber="25" type="female">13 ♀♀</specimenCount>
, same data as
<typeStatus id="7D748835C410FFCBF1A1F910FC9D5EAD" box="[691,794,1732,1758]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
(
<collectionCode id="C4DEAE52C410FFCBF03BF911FCFA5EAC" box="[809,893,1733,1759]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="24" pageNumber="25">MZUF</collectionCode>
,
<collectionCode id="C4DEAE52C410FFCBF09AF911FC2B5EAC" box="[904,940,1733,1759]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14050" lsid="urn:lsid:biocol.org:col:14050" name="Sichuan Institute of Natural Resources" pageId="24" pageNumber="25" type="Herbarium">SR</collectionCode>
and
<collectionCode id="C4DEAE52C410FFCBF0F7F911FBB85EAC" box="[997,1087,1733,1759]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="24" pageNumber="25">ZMUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="EAD5651CC410FFC9F3AFF8D2FD445A2D" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="description">
<paragraph id="A2703697C410FFCBF3AFF8D2FEC95F53" blockId="24.[189,334,1798,1824]" box="[189,334,1798,1824]" pageId="24" pageNumber="25">
<heading id="F93881FBC410FFCBF3AFF8D2FEC95F53" bold="true" box="[189,334,1798,1824]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="90BBEA85C410FFCBF3AFF8D2FEC95F53" bold="true" box="[189,334,1798,1824]" pageId="24" pageNumber="25">Description</emphasis>
</heading>
</paragraph>
<paragraph id="A2703697C410FFCAF3AFF8E0FAF159FC" blockId="24.[189,1399,1844,2012]" lastBlockId="25.[189,1399,267,399]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
Maximum size:
<specimenCount id="B4C9FD1EC410FFCBF26EF8E1FE155F3C" box="[380,402,1845,1871]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
and
<specimenCount id="B4C9FD1EC410FFCBF2D8F8E1FE675F3C" box="[458,480,1845,1871]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
2.7×
<quantity id="65379B72C410FFCBF136F8E1FD065F3C" box="[548,641,1845,1871]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="24" pageNumber="25" unit="mm" value="0.8">0.8 mm</quantity>
. Body colourless and elongated (
<figureCitation id="3AF42A12C410FFCBF717F8E0FBF55F3C" box="[1029,1138,1844,1871]" captionStart="Fig" captionStartId="25.[189,232,1873,1899]" captionTargetBox="[189,1405,449,1839]" captionTargetId="figure@25.[180,1411,438,1847]" captionTargetPageId="25" captionText="Fig. 16. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida. — Paratype, ♀. A. Adult specimen in dorsal view. B. Dorsal scale-seta. C. Cephalon in frontal view. D. Cephalon in dorsal view. E. Pleonites 4, 5, telson and left uropod. — Paratype, ♂. F. Antennula. G. Antenna." figureDoi="http://doi.org/10.5281/zenodo.3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="24" pageNumber="25">Fig. 16A</figureCitation>
). Dorsum of cephalon and pereonites with two rows and one row of large granules, respectively; each granule with a scale-seta (
<figureCitation id="3AF42A12C410FFCBF3D4F8AFFEB45FE5" box="[198,307,1915,1942]" captionStart="Fig" captionStartId="25.[189,232,1873,1899]" captionTargetBox="[189,1405,449,1839]" captionTargetId="figure@25.[180,1411,438,1847]" captionTargetPageId="25" captionText="Fig. 16. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida. — Paratype, ♀. A. Adult specimen in dorsal view. B. Dorsal scale-seta. C. Cephalon in frontal view. D. Cephalon in dorsal view. E. Pleonites 4, 5, telson and left uropod. — Paratype, ♂. F. Antennula. G. Antenna." figureDoi="http://doi.org/10.5281/zenodo.3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="24" pageNumber="25">Fig. 16B</figureCitation>
); pleon and telson smooth. Cephalon (
<figureCitation id="3AF42A12C410FFCBF017F8AFFCDD5FE5" box="[773,858,1915,1942]" captionStart="Fig" captionStartId="25.[189,232,1873,1899]" captionTargetBox="[189,1405,449,1839]" captionTargetId="figure@25.[180,1411,438,1847]" captionTargetPageId="25" captionText="Fig. 16. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida. — Paratype, ♀. A. Adult specimen in dorsal view. B. Dorsal scale-seta. C. Cephalon in frontal view. D. Cephalon in dorsal view. E. Pleonites 4, 5, telson and left uropod. — Paratype, ♂. F. Antennula. G. Antenna." figureDoi="http://doi.org/10.5281/zenodo.3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="24" pageNumber="25">Fig. 16</figureCitation>
C–D) with short, quadrangular frontal lateral lobes. Eyes absent. Pereonites 1–3 with rounded posterior corners; pereonites 4–7 with epimera pointing backwards (
<figureCitation id="3AF42A12C410FFCBF25CF815FE385FAF" box="[334,447,1985,2012]" captionStart="Fig" captionStartId="25.[189,232,1873,1899]" captionTargetBox="[189,1405,449,1839]" captionTargetId="figure@25.[180,1411,438,1847]" captionTargetPageId="25" captionText="Fig. 16. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida. — Paratype, ♀. A. Adult specimen in dorsal view. B. Dorsal scale-seta. C. Cephalon in frontal view. D. Cephalon in dorsal view. E. Pleonites 4, 5, telson and left uropod. — Paratype, ♂. F. Antennula. G. Antenna." figureDoi="http://doi.org/10.5281/zenodo.3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="24" pageNumber="25">Fig. 16A</figureCitation>
). Pleon (
<figureCitation id="3AF42A12C410FFCBF13CF815FD475FAF" box="[558,704,1985,2012]" captionStart="Fig" captionStartId="25.[189,232,1873,1899]" captionTargetBox="[189,1405,449,1839]" captionTargetId="figure@25.[180,1411,438,1847]" captionTargetPageId="25" captionText="Fig. 16. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida. — Paratype, ♀. A. Adult specimen in dorsal view. B. Dorsal scale-seta. C. Cephalon in frontal view. D. Cephalon in dorsal view. E. Pleonites 4, 5, telson and left uropod. — Paratype, ♂. F. Antennula. G. Antenna." figureDoi="http://doi.org/10.5281/zenodo.3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="24" pageNumber="25">Fig. 16A, E</figureCitation>
) slightly narrower than pereon; pleonites 3–5 with small epimera and very short posterior points. Telson (
<figureCitation id="3AF42A12C411FFCAF1EBFEDFFCE65955" box="[761,865,267,294]" captionStart="Fig" captionStartId="25.[189,232,1873,1899]" captionTargetBox="[189,1405,449,1839]" captionTargetId="figure@25.[180,1411,438,1847]" captionTargetPageId="25" captionText="Fig. 16. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida. — Paratype, ♀. A. Adult specimen in dorsal view. B. Dorsal scale-seta. C. Cephalon in frontal view. D. Cephalon in dorsal view. E. Pleonites 4, 5, telson and left uropod. — Paratype, ♂. F. Antennula. G. Antenna." figureDoi="http://doi.org/10.5281/zenodo.3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="25" pageNumber="26">Fig. 16E</figureCitation>
) about twice as wide as long, with distal part trapezoidal, with concave sides and broadly rounded apex. Antennula (
<figureCitation id="3AF42A12C411FFCAF0FDFEFAFBD4593A" box="[1007,1107,302,329]" captionStart="Fig" captionStartId="25.[189,232,1873,1899]" captionTargetBox="[189,1405,449,1839]" captionTargetId="figure@25.[180,1411,438,1847]" captionTargetPageId="25" captionText="Fig. 16. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida. — Paratype, ♀. A. Adult specimen in dorsal view. B. Dorsal scale-seta. C. Cephalon in frontal view. D. Cephalon in dorsal view. E. Pleonites 4, 5, telson and left uropod. — Paratype, ♂. F. Antennula. G. Antenna." figureDoi="http://doi.org/10.5281/zenodo.3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="25" pageNumber="26">Fig. 16F</figureCitation>
) with three articles; third article with three long aesthetascs at apex. Antenna (
<figureCitation id="3AF42A12C411FFCAF03BFE86FC13591E" box="[809,916,338,365]" captionStart="Fig" captionStartId="25.[189,232,1873,1899]" captionTargetBox="[189,1405,449,1839]" captionTargetId="figure@25.[180,1411,438,1847]" captionTargetPageId="25" captionText="Fig. 16. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida. — Paratype, ♀. A. Adult specimen in dorsal view. B. Dorsal scale-seta. C. Cephalon in frontal view. D. Cephalon in dorsal view. E. Pleonites 4, 5, telson and left uropod. — Paratype, ♂. F. Antennula. G. Antenna." figureDoi="http://doi.org/10.5281/zenodo.3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="25" pageNumber="26">Fig. 16G</figureCitation>
) with fourth and fifth article of peduncle with distinct tubercles bearing scale-setae; fifth article slightly shorter than flagellum; flagellum with
</paragraph>
<caption id="F6B0661FC411FFCAF3AFF885FECB5FA6" ID-DOI="http://doi.org/10.5281/zenodo.3805653" ID-Zenodo-Dep="3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,232,1873,1899]" targetBox="[189,1405,449,1839]" targetPageId="25">
<paragraph id="A2703697C411FFCAF3AFF885FECB5FA6" blockId="25.[189,1399,1873,2006]" pageId="25" pageNumber="26">
<emphasis id="90BBEA85C411FFCAF3AFF885FE995F1F" bold="true" box="[189,286,1873,1900]" pageId="25" pageNumber="26">Fig. 16.</emphasis>
<taxonomicName id="65CF4D14C411FFCAF238F886FCD45F1F" authority="Reboleira &amp; Taiti" authorityName="Reboleira &amp; Gonçalves &amp; Oromí &amp; Taiti" authorityYear="2015" box="[298,851,1874,1900]" class="Malacostraca" family="Trichoniscidae" genus="Troglonethes" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="arrabidaensis" status="sp. nov.">
<emphasis id="90BBEA85C411FFCAF238F886FDF65F1F" box="[298,625,1874,1900]" italics="true" pageId="25" pageNumber="26">Troglonethes arrabidaensis</emphasis>
Reboleira &amp; Taiti
</taxonomicName>
<taxonomicNameLabel id="8B8857FEC411FFCAF072F886FC385F1F" box="[864,959,1874,1900]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
from Gruta do Frade, Arrábida. — Paratype, ♀.
<emphasis id="90BBEA85C411FFCAF272F8A1FEF25FFC" bold="true" box="[352,373,1909,1935]" pageId="25" pageNumber="26">A</emphasis>
. Adult specimen in dorsal view.
<emphasis id="90BBEA85C411FFCAF003F8A1FCA25FFC" bold="true" box="[785,805,1909,1935]" pageId="25" pageNumber="26">B</emphasis>
. Dorsal scale-seta.
<emphasis id="90BBEA85C411FFCAF705F8A1FBAB5FFC" bold="true" box="[1047,1068,1909,1935]" pageId="25" pageNumber="26">C</emphasis>
. Cephalon in frontal view.
<emphasis id="90BBEA85C411FFCAF3AFF84CFF555FC1" bold="true" box="[189,210,1944,1970]" pageId="25" pageNumber="26">D</emphasis>
. Cephalon in dorsal view.
<emphasis id="90BBEA85C411FFCAF10BF84CFDAA5FC1" bold="true" box="[537,557,1944,1970]" pageId="25" pageNumber="26">E</emphasis>
. Pleonites 4, 5, telson and left uropod. — Paratype, ♂.
<emphasis id="90BBEA85C411FFCAF7C1F84CFB625FC1" bold="true" box="[1235,1253,1944,1970]" pageId="25" pageNumber="26">F</emphasis>
. Antennula.
<emphasis id="90BBEA85C411FFCAF3AFF86FFF535FA6" bold="true" box="[189,212,1979,2005]" pageId="25" pageNumber="26">G</emphasis>
. Antenna.
</paragraph>
</caption>
<paragraph id="A2703697C412FFC9F3AFFEDFFAB1593B" blockId="26.[189,1399,267,329]" pageId="26" pageNumber="27">
three articles, with two long aesthetascs on second article. Buccal pieces as in previous species. Uropod (
<figureCitation id="3AF42A12C412FFC9F3D4FEFAFEAA593A" box="[198,301,302,329]" captionStart="Fig" captionStartId="25.[189,232,1873,1899]" captionTargetBox="[189,1405,449,1839]" captionTargetId="figure@25.[180,1411,438,1847]" captionTargetPageId="25" captionText="Fig. 16. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida. — Paratype, ♀. A. Adult specimen in dorsal view. B. Dorsal scale-seta. C. Cephalon in frontal view. D. Cephalon in dorsal view. E. Pleonites 4, 5, telson and left uropod. — Paratype, ♂. F. Antennula. G. Antenna." figureDoi="http://doi.org/10.5281/zenodo.3805653" httpUri="https://zenodo.org/record/3805653/files/figure.png" pageId="26" pageNumber="27">Fig. 16E</figureCitation>
) with exopod slightly longer than endopod, exopod and endopod inserted at same level.
</paragraph>
<paragraph id="A2703697C412FFC9F3AFFEA4FD445A2D" blockId="26.[189,1399,368,606]" pageId="26" pageNumber="27">
MALE. Pereopod 1 (
<figureCitation id="3AF42A12C412FFC9F2BEFEA4FD9D59F9" box="[428,538,368,394]" captionStart="Fig" captionStartId="26.[189,232,1909,1935]" captionTargetBox="[194,1417,657,1877]" captionTargetId="figure@26.[163,1422,652,1889]" captionTargetPageId="26" captionText="Fig. 17. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida, paratype, ♂. A. Pereopod 1. B. Pereopod 7. C. Genital papilla and pleopod 1. D. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.3805655" httpUri="https://zenodo.org/record/3805655/files/figure.png" pageId="26" pageNumber="27">Fig. 17A</figureCitation>
) with no particular modifications. Pereopod 7 (
<figureCitation id="3AF42A12C412FFC9F74EFEA4FB4059F9" box="[1116,1223,368,394]" captionStart="Fig" captionStartId="26.[189,232,1909,1935]" captionTargetBox="[194,1417,657,1877]" captionTargetId="figure@26.[163,1422,652,1889]" captionTargetPageId="26" captionText="Fig. 17. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida, paratype, ♂. A. Pereopod 1. B. Pereopod 7. C. Genital papilla and pleopod 1. D. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.3805655" httpUri="https://zenodo.org/record/3805655/files/figure.png" pageId="26" pageNumber="27">Fig. 17B</figureCitation>
) ischium with straight sternal margin, carpus enlarged in basal part. Genital papilla (
<figureCitation id="3AF42A12C412FFC9F0FCFE47FBD059DE" box="[1006,1111,403,429]" captionStart="Fig" captionStartId="26.[189,232,1909,1935]" captionTargetBox="[194,1417,657,1877]" captionTargetId="figure@26.[163,1422,652,1889]" captionTargetPageId="26" captionText="Fig. 17. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida, paratype, ♂. A. Pereopod 1. B. Pereopod 7. C. Genital papilla and pleopod 1. D. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.3805655" httpUri="https://zenodo.org/record/3805655/files/figure.png" pageId="26" pageNumber="27">Fig. 17C</figureCitation>
) fusiform, with rounded apex, much longer than pleopod 1. Pleopod 1 (
<figureCitation id="3AF42A12C412FFC9F1C4FE62FCBA59A3" box="[726,829,438,464]" captionStart="Fig" captionStartId="26.[189,232,1909,1935]" captionTargetBox="[194,1417,657,1877]" captionTargetId="figure@26.[163,1422,652,1889]" captionTargetPageId="26" captionText="Fig. 17. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida, paratype, ♂. A. Pereopod 1. B. Pereopod 7. C. Genital papilla and pleopod 1. D. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.3805655" httpUri="https://zenodo.org/record/3805655/files/figure.png" pageId="26" pageNumber="27">Fig. 17C</figureCitation>
) exopod triangular, as wide as long, with broadly rounded apex; endopod with basal article elongated, shorter than exopod, with sinuous outer margin, distal article flagelliform. Pleopod 2 (
<figureCitation id="3AF42A12C412FFC9F166FE29FD585A64" box="[628,735,509,535]" captionStart="Fig" captionStartId="26.[189,232,1909,1935]" captionTargetBox="[194,1417,657,1877]" captionTargetId="figure@26.[163,1422,652,1889]" captionTargetPageId="26" captionText="Fig. 17. Troglonethes arrabidaensis Reboleira &amp; Taiti sp. nov. from Gruta do Frade, Arrábida, paratype, ♂. A. Pereopod 1. B. Pereopod 7. C. Genital papilla and pleopod 1. D. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.3805655" httpUri="https://zenodo.org/record/3805655/files/figure.png" pageId="26" pageNumber="27">Fig. 17D</figureCitation>
) exopod triangular, with broadly rounded apex with few short setae; endopod biarticulated, basal article more than twice as long as distal one, with parallel sides, distal article with hook-like process at apex.
</paragraph>
</subSubSection>
<caption id="F6B0661FC412FFC9F3AFF8A1FB395FC0" ID-DOI="http://doi.org/10.5281/zenodo.3805655" ID-Zenodo-Dep="3805655" httpUri="https://zenodo.org/record/3805655/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1909,1935]" targetBox="[194,1417,657,1877]" targetPageId="26">
<paragraph id="A2703697C412FFC9F3AFF8A1FB395FC0" blockId="26.[189,1398,1909,1971]" pageId="26" pageNumber="27">
<emphasis id="90BBEA85C412FFC9F3AFF8A1FE9B5FFC" bold="true" box="[189,284,1909,1935]" pageId="26" pageNumber="27">Fig. 17.</emphasis>
<taxonomicName id="65CF4D14C412FFC9F234F8A2FCC15FFC" authority="Reboleira &amp; Taiti" authorityName="Reboleira &amp; Gonçalves &amp; Oromí &amp; Taiti" authorityYear="2015" box="[294,838,1909,1936]" class="Malacostraca" family="Trichoniscidae" genus="Troglonethes" kingdom="Animalia" order="Isopoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="arrabidaensis" status="sp. nov.">
<emphasis id="90BBEA85C412FFC9F234F8A2FDEC5FE3" box="[294,619,1910,1936]" italics="true" pageId="26" pageNumber="27">Troglonethes arrabidaensis</emphasis>
Reboleira &amp; Taiti
</taxonomicName>
<taxonomicNameLabel id="8B8857FEC412FFC9F042F8A2FC295FE3" box="[848,942,1910,1936]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
from Gruta do Frade, Arrábida, paratype, ♂.
<emphasis id="90BBEA85C412FFC9F231F84DFEBF5FC0" bold="true" box="[291,312,1945,1971]" pageId="26" pageNumber="27">A</emphasis>
. Pereopod 1.
<emphasis id="90BBEA85C412FFC9F2C8F84DFE695FC0" bold="true" box="[474,494,1945,1971]" pageId="26" pageNumber="27">B</emphasis>
. Pereopod 7.
<emphasis id="90BBEA85C412FFC9F182F84DFD225FC0" bold="true" box="[656,677,1945,1971]" pageId="26" pageNumber="27">C</emphasis>
. Genital papilla and pleopod 1.
<emphasis id="90BBEA85C412FFC9F70FF84DFBB55FC0" bold="true" box="[1053,1074,1945,1971]" pageId="26" pageNumber="27">D</emphasis>
. Pleopod 2.
</paragraph>
</caption>
<subSubSection id="EAD5651CC413FFC8F3AFFEDFFC0E5A70" pageId="27" pageNumber="28" type="discussion">
<paragraph id="A2703697C413FFC8F3AFFEDFFEA85956" blockId="27.[189,303,267,293]" box="[189,303,267,293]" pageId="27" pageNumber="28">
<heading id="F93881FBC413FFC8F3AFFEDFFEA85956" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="90BBEA85C413FFC8F3AFFEDFFEA85956" bold="true" box="[189,303,267,293]" pageId="27" pageNumber="28">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="A2703697C413FFC8F3AFFEEDFC0E5A70" blockId="27.[189,1399,313,516]" pageId="27" pageNumber="28">
The new species is very similar to
<taxonomicName id="65CF4D14C413FFC8F140FEEDFCCB5920" authorityName="Cruz" authorityYear="1991" box="[594,844,313,339]" class="Malacostraca" family="Trichoniscidae" genus="Troglonethes" kingdom="Animalia" order="Isopoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="aurouxi">
<emphasis id="90BBEA85C413FFC8F140FEEDFCCB5920" box="[594,844,313,339]" italics="true" pageId="27" pageNumber="28">Troglonethes aurouxi</emphasis>
</taxonomicName>
in the dorsal granulation and shape of the male pleopod 2. It differs in having the antennal flagellum with three, instead of four, articles, and a wider male pleopod 1 exopod. It is readily distinguishable from
<taxonomicName id="65CF4D14C413FFC8F079FE55FB7359E9" authority="Reboleira &amp; Taiti" authorityName="Reboleira &amp; Gonçalves &amp; Oromí &amp; Taiti" authorityYear="2015" box="[875,1268,384,410]" class="Malacostraca" family="Trichoniscidae" genus="Troglonethes" kingdom="Animalia" order="Isopoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="olissipoensis" status="sp. nov.">
<emphasis id="90BBEA85C413FFC8F079FE55FB9859E9" box="[875,1055,384,410]" italics="true" pageId="27" pageNumber="28">T. olissipoensis</emphasis>
Reboleira &amp; Taiti
</taxonomicName>
<taxonomicNameLabel id="8B8857FEC413FFC8F7EFFE54FADF59E9" box="[1277,1368,384,410]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
in the larger and less numerous dorsal granules, the enlarged basal part of the male pereopod 7 carpus, the male pleopod 1 endopod with the basal article shorter than the exopod, and the distal article of the male pleopod 2 endopod lacking the hook-like process at the base.
</paragraph>
</subSubSection>
<subSubSection id="EAD5651CC413FFC8F3AFFDFFFD345AC9" pageId="27" pageNumber="28" type="biology_ecology">
<paragraph id="A2703697C413FFC8F3AFFDFFFE015A36" blockId="27.[189,390,555,581]" box="[189,390,555,581]" pageId="27" pageNumber="28">
<heading id="F93881FBC413FFC8F3AFFDFFFE015A36" bold="true" box="[189,390,555,581]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="90BBEA85C413FFC8F3AFFDFFFE015A36" bold="true" box="[189,390,555,581]" pageId="27" pageNumber="28">Ecological notes</emphasis>
</heading>
</paragraph>
<paragraph id="A2703697C413FFC8F3AFFD8DFD345AC9" blockId="27.[189,1399,601,698]" pageId="27" pageNumber="28">This species was only found in the Gruta do Frade in the Arrábida Massif. The cave entrance is by the seashore and there are several anchialine lakes inside, influenced by the sea tides. It is the first record of troglobiotic terrestrial isopods in Arrábida.</paragraph>
</subSubSection>
</treatment>
</document>
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