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<document id="53581F4F4C7D4D7F01AB8652306E8472" ID-CLB-Dataset="24105" ID-DOI="10.5852/ejt.2019.585" ID-GBIF-Dataset="dca608e8-fccf-4c1c-b8df-ef0c28e1d518" ID-Zenodo-Dep="3660259" ID-ZooBank="181B6FBA-ED75-4BB4-84C4-FB512B794749" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581251975682" checkinUser="plazi" docAuthor="Ebersole, Jun A., Cicimurri, David J. &amp; Stringer, Gary L." docDate="2019" docId="18174D41FFC8FFE0FE2E9E8B4E950A89" docLanguage="en" docName="ejt-585_ebersole_cicimurri_stinger.pdf.imf" docOrigin="European Journal of Taxonomy 585" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhizoprionodon ganntourensis" docType="treatment" docVersion="19" lastPageNumber="80" masterDocId="E42E3539FF85FFAFFFC69F344D4B0F37" masterDocTitle="Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths" masterLastPageNumber="274" masterPageNumber="1" pageNumber="78" updateTime="1698783570407" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A50D67E1BE90F4534EBEF3941EBC38FB">Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths</mods:title>
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<treatment id="18174D41FFC8FFE0FE2E9E8B4E950A89" ID-DOI="http://doi.org/10.5281/zenodo.3664585" ID-GBIF-Taxon="162073057" ID-Zenodo-Dep="3664585" LSID="urn:lsid:plazi:treatment:18174D41FFC8FFE0FE2E9E8B4E950A89" httpUri="http://treatment.plazi.org/id/18174D41FFC8FFE0FE2E9E8B4E950A89" lastPageId="79" lastPageNumber="80" pageId="77" pageNumber="78">
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<taxonomicName id="57BE87D4FFC8FFE2FE2E9E8B49000EED" authority="(Arambourg, 1952)" baseAuthorityName="Arambourg" baseAuthorityYear="1952" box="[488,1099,447,474]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="ganntourensis">
<emphasis id="A2CA2045FFC8FFE2FE2E9E8B4E2B0EED" bold="true" box="[488,864,447,474]" italics="true" pageId="77" pageNumber="78">Rhizoprionodon ganntourensis</emphasis>
(
<bibRefCitation id="F42F81A6FFC8FFE2FCA99E8B49080EED" author="Arambourg C." box="[879,1091,447,474]" pageId="77" pageNumber="78" pagination="1 - 372" refId="ref136183" refString="Arambourg C. 1952. Les vertebres fossiles des gisements de phosphates (Maroc-Algerie-Tunisie). Notes et Memoires du Service geologique du Maroc 92: 1 - 372." type="journal article" year="1952">Arambourg, 1952</bibRefCitation>
)
</taxonomicName>
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</paragraph>
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<paragraph id="9001FC57FFC8FFE2FD369ED04E080EC9" blockId="77.[488,1099,447,510]" box="[752,835,484,510]" pageId="77" pageNumber="78">
<figureCitation id="0885E0D2FFC8FFE2FD369ED04E080EC9" box="[752,835,484,510]" captionStart="Fig" captionStartId="78.[189,232,1472,1498]" captionTargetBox="[202,1385,264,1423]" captionTargetId="figure@78.[189,1398,240,1456]" captionTargetPageId="78" captionText="Fig. 28. Rhizoprionodon ganntourensis (Arambourg, 1952), teeth. AC. MSC 2173.4, lower lateroposterior tooth, basal Gosport Sand. A. Labial view. B. Lingual view. C. Mesial view. DF. MSC 37674.3, lower anterior tooth, basal Lisbon Formation. D. Labial view. E. Lingual view. F. Mesial view. GI. MSC 2173.5, lower lateral tooth, basal Gosport Sand. G. Labial view. H. Lingual view. I. Mesial view. JL. MSC 37907.1, lower lateral tooth, lower Tallahatta Formation. J. Labial view. K. Lingual view. L. Mesial view. MO. MSC 2173.6, upper anterolateral tooth, basal Gosport Sand. M. Labial view. N. Lingual view. O. Mesial view. PR. MSC 38504.7, lower lateroposterior tooth, basal Gosport Sand. P. Labial view. Q. Lingual view. R. Mesial view. SU. MSC 35753.3, upper anterolateral tooth, lower Tallahatta Formation. S. Labial view. T. Lingual view. U. Mesial view. VX. MSC 37648.2, lower lateral tooth, basal Gosport Sand. V. Labial view. W. Lingual view. X. Mesial view. YAA. MSC 37098, lower lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. Y. Labial view. Z. Lingual view. AA. Mesial view. BBDD. MSC 37674.1, lower lateroposterior tooth, basal Lisbon Formation. BB. Labial view. CC. Lingual view. DD. Mesial view. EEGG. MSC 37648.1, lower lateral tooth, basal Gosport Sand. EE. Labial view. FF. Lingual view. GG. Mesial view. HHJJ. MSC 37674.4, lower lateral tooth, basal Lisbon Formation. HH. Labial view. II. Lingual view. JJ. Mesial view. Scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660313" httpUri="https://zenodo.org/record/3660313/files/figure.png" pageId="77" pageNumber="78">Fig. 28</figureCitation>
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<taxonomicName id="57BE87D4FFC8FFE2FF7B9D124FB80D77" authority="Arambourg, 1952: 164" authorityName="Arambourg" authorityPageNumber="164" authorityYear="1952" box="[189,755,550,577]" class="Elasmobranchii" family="Carcharhinidae" genus="Scoliodon" kingdom="Animalia" order="Carcharhiniformes" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="ganntourensis">
<emphasis id="A2CA2045FFC8FFE2FF7B9D124C940D77" box="[189,479,550,576]" italics="true" pageId="77" pageNumber="78">Scoliodon ganntourensis</emphasis>
<treatmentCitation id="111FDA46FFC8FFE2FE229D124FB80D77" author="Arambourg C." box="[484,755,550,577]" page="164" pageId="77" pageNumber="78" year="1952">
<bibRefCitation id="F42F81A6FFC8FFE2FE229D124FB80D77" author="Arambourg C." box="[484,755,550,577]" pageId="77" pageNumber="78" pagination="1 - 372" refId="ref136183" refString="Arambourg C. 1952. Les vertebres fossiles des gisements de phosphates (Maroc-Algerie-Tunisie). Notes et Memoires du Service geologique du Maroc 92: 1 - 372." type="journal article" year="1952">Arambourg, 1952: 164</bibRefCitation>
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, pl. 24, figs 4963; text-fig. 33.
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</paragraph>
<paragraph id="9001FC57FFC8FFE2FF7B9D5D4E390DB4" blockId="77.[189,993,616,752]" box="[189,882,616,643]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="B0AEDB79FFC8FFE2FF7B9D5D4E390DB4" box="[189,882,616,643]" pageId="77" pageNumber="78">
<taxonomicName id="57BE87D4FFC8FFE2FF7B9D5D4CE80DB4" authorityName="Whitley" authorityYear="1929" box="[189,419,617,643]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="sp.">
<emphasis id="A2CA2045FFC8FFE2FF7B9D5D4C310DB4" box="[189,378,617,643]" italics="true" pageId="77" pageNumber="78">Rhizoprionodon</emphasis>
sp.
</taxonomicName>
<treatmentCitation id="111FDA46FFC8FFE2FE069D5C4E5F0DB5" author="Thurmond J. T. &amp; Jones D. E." box="[448,788,616,643]" page="68" pageId="77" pageNumber="78" year="1981">
<bibRefCitation id="F42F81A6FFC8FFE2FE069D5C4E5F0DB5" author="Thurmond J. T. &amp; Jones D. E." box="[448,788,616,643]" pageId="77" pageNumber="78" refId="ref152244" refString="Thurmond J. T. &amp; Jones D. E. 1981. Fossil Vertebrates of Alabama. University of Alabama Press, Tuscaloosa." type="book" year="1981">Thurmond &amp; Jones 1981: 68</bibRefCitation>
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, fig. 33.
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</paragraph>
<paragraph id="9001FC57FFC8FFE2FF7B9DB94EAA0D90" blockId="77.[189,993,616,752]" box="[189,993,653,679]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="B0AEDB79FFC8FFE2FF7B9DB94EAA0D90" box="[189,993,653,679]" pageId="77" pageNumber="78">
<taxonomicName id="57BE87D4FFC8FFE2FF7B9DB94E960D90" authority="- Cappetta &amp; Traverse 1988: 361" authorityName="Cappetta &amp; Traverse" authorityPageNumber="361" authorityYear="1988" box="[189,989,653,679]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="ganntourensis">
<emphasis id="A2CA2045FFC8FFE2FF7B9DB94C310D90" box="[189,378,653,679]" italics="true" pageId="77" pageNumber="78">Rhizoprionodon</emphasis>
cf.
<emphasis id="A2CA2045FFC8FFE2FE619DBA4F070D90" box="[423,588,654,679]" italics="true" pageId="77" pageNumber="78">ganntourensis</emphasis>
<treatmentCitation id="111FDA46FFC8FFE2FDAF9DB94E960D90" author="Cappetta H. &amp; Traverse M." box="[617,989,653,679]" page="361" pageId="77" pageNumber="78" year="1988">
<bibRefCitation id="F42F81A6FFC8FFE2FDAF9DB94E960D90" author="Cappetta H. &amp; Traverse M." box="[617,989,653,679]" pageId="77" pageNumber="78" pagination="359 - 365" refId="ref138153" refString="Cappetta H. &amp; Traverse M. 1988. Une riche faune de selaciens dans le bassin a phosphate de Kpogame- Hahotoe (Eocene moyen du Togo), Note preliminaire et precisions sur la structure et l'age du gisement. Geobios 21 (3): 359 - 365. https: // doi. org / 10.1016 / S 0016 - 6995 (88) 80058 - 5" type="journal article" year="1988">Cappetta &amp; Traverse 1988: 361</bibRefCitation>
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.
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</paragraph>
<paragraph id="9001FC57FFC8FFE2FF7B9D854FB70DFC" blockId="77.[189,993,616,752]" box="[189,764,689,715]" pageId="77" pageNumber="78">
<taxonomicName id="57BE87D4FFC8FFE2FF7B9D854CE80DFC" authorityName="Whitley" authorityYear="1929" box="[189,419,689,715]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="speyeri">
<emphasis id="A2CA2045FFC8FFE2FF7B9D854C310DFC" box="[189,378,689,715]" italics="true" pageId="77" pageNumber="78">Rhizoprionodon</emphasis>
sp.
</taxonomicName>
<bibRefCitation id="F42F81A6FFC8FFE2FE069D854FEE0DFC" author="Clayton A. A. &amp; Ciampaglio, C. N. &amp; Cicimurri, D. J." box="[448,677,689,715]" pageId="77" pageNumber="78" pagination="60 - 73" refId="ref139906" refString="Clayton A. A., Ciampaglio, C. N. &amp; Cicimurri, D. J. 2013. An inquiry into the stratigraphic occurrence of a Claibornian (Eocene) vertebrate fauna from Covington County, Alabama. Bulletin Alabama Museum of Natural History 31 (2): 60 - 73." type="journal article" year="2013">
Clayton
<emphasis id="A2CA2045FFC8FFE2FDE29D864F140DFC" box="[548,607,689,715]" italics="true" pageId="77" pageNumber="78">et al.</emphasis>
2013
</bibRefCitation>
: fig. 2i.
</paragraph>
<paragraph id="9001FC57FFC8FFE2FF7B9DE14E2B0DD8" blockId="77.[189,993,616,752]" box="[189,864,725,752]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="B0AEDB79FFC8FFE2FF7B9DE14E2B0DD8" box="[189,864,725,752]" pageId="77" pageNumber="78">
<taxonomicName id="57BE87D4FFC8FFE2FF7B9DE14C0C0DC7" authorityName="Rafinesque" authorityYear="1810" box="[189,327,725,752]" class="Elasmobranchii" family="Sphyrnidae" genus="Sphyrna" kingdom="Animalia" order="Carcharhiniformes" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="speyeri">
<emphasis id="A2CA2045FFC8FFE2FF7B9DE14C540DD8" box="[189,287,725,751]" italics="true" pageId="77" pageNumber="78">Sphyrna</emphasis>
sp.
</taxonomicName>
<treatmentCitation id="111FDA46FFC8FFE2FEA39DE14FD60DD8" author="Cappetta H. &amp; Case G. R." box="[357,669,725,751]" page="61" pageId="77" pageNumber="78" year="2016">
<bibRefCitation id="F42F81A6FFC8FFE2FEA39DE14FD60DD8" author="Cappetta H. &amp; Case G. R." box="[357,669,725,751]" pageId="77" pageNumber="78" pagination="43 - 103" refId="ref137973" refString="Cappetta H. &amp; Case G. R. 2016. A selachian fauna from the middle Eocene (Lutetian, Lisbon Formation) of Andalusia, Covington County, Alabama, USA. Palaeontographica Abteilung A 307 (1 - 6): 43 - 103." type="journal article" year="2016">Cappetta &amp; Case 2016: 61</bibRefCitation>
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, pl. 10, figs 24.
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<paragraph id="9001FC57FFC8FFE2FF7B9C234CE10C06" blockId="77.[189,426,791,817]" box="[189,426,791,817]" pageId="77" pageNumber="78">
<heading id="CB494B3BFFC8FFE2FF7B9C234CE10C06" bold="true" box="[189,426,791,817]" fontSize="11" level="3" pageId="77" pageNumber="78" reason="3">
<emphasis id="A2CA2045FFC8FFE2FF7B9C234CE10C06" bold="true" box="[189,426,791,817]" pageId="77" pageNumber="78">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FFC8FFE2FF7B9C734CB80B68" blockId="77.[189,1399,839,1119]" pageId="77" pageNumber="78">
<materialsCitation id="20D6F60AFFC8FFE2FF7B9C734CA40B68" ID-GBIF-Occurrence="2564277926" collectionCode="ALMNH, MMNS, MSC, SC" country="United States of America" location="Claiborne Group" pageId="77" pageNumber="78" specimenCode="PVPV 2005.6, MSC 188.324, MSC 2173.1, 7, MSC 2379.1, 5, MSC 35753.1, MSC 37264.1, MSC 37648.1, 3, MSC 37674.1, 4, MSC 37904.1, MSC 37905.1, MSC 37906.1, 2, MSC 37907.1, MSC 38199.1, MSC 38486.1, 6, MSC 38504.1, 10, MSC 38544.1, 11, WSU 5017, WSU 5037" specimenCount="73" stateProvince="Alabama">
<collectingCountry id="E8A9BCC7FFC8FFE2FF7B9C734F230C56" box="[189,616,839,865]" name="United States of America" pageId="77" pageNumber="78">UNITED STATES OF AMERICA</collectingCountry>
<emphasis id="A2CA2045FFC8FFE2FD539C734E4C0C56" bold="true" box="[661,775,839,865]" pageId="77" pageNumber="78">
<collectingRegion id="527A32B5FFC8FFE2FD539C734E4C0C56" box="[661,775,839,865]" country="United States of America" name="Alabama" pageId="77" pageNumber="78">Alabama</collectingRegion>
</emphasis>
• 156 isolated teeth;
<location id="9561AA8CFFC8FFE2FBE19C7349B20C56" LSID="urn:lsid:plazi:treatment:18174D41FFC8FFE0FE2E9E8B4E950A89:9561AA8CFFC8FFE2FBE19C7349B20C56" box="[1063,1273,839,865]" country="United States of America" name="Claiborne Group" pageId="77" pageNumber="78" stateProvince="Alabama">Claiborne Group</location>
;
<collectionCode id="F6AF6492FFC8FFE2FACD9C73483C0C56" box="[1291,1399,839,865]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="77" pageNumber="78" type="Museum">ALMNH</collectionCode>
PV1993.2.426 (
<specimenCount id="86B837DEFFC8FFE2FEBF9C5F4F6B0CB1" box="[377,544,875,902]" pageId="77" pageNumber="78" type="generic">24 specimens</specimenCount>
),
<collectionCode id="F6AF6492FFC8FFE2FDF39C5F4FEA0CB2" box="[565,673,875,901]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="77" pageNumber="78" type="Museum">ALMNH</collectionCode>
PV1993.2.442 (
<specimenCount id="86B837DEFFC8FFE2FCA09C5F49460CB1" box="[870,1037,875,902]" pageId="77" pageNumber="78" type="generic">17 specimens</specimenCount>
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.
</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFFC8FFE2FF7B9BB249660AFA" pageId="77" pageNumber="78" type="description">
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<heading id="CB494B3BFFC8FFE2FF7B9BB24C050B97" bold="true" box="[189,334,1158,1184]" fontSize="11" level="3" pageId="77" pageNumber="78" reason="3">
<emphasis id="A2CA2045FFC8FFE2FF7B9BB24C050B97" bold="true" box="[189,334,1158,1184]" pageId="77" pageNumber="78">Description</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FFC8FFE2FF7B9B8249660AFA" blockId="77.[189,1399,1206,1485]" pageId="77" pageNumber="78">
Teeth small, most not exceeding
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in mesiodistal width. Crown wide, with broad-based and distally inclined triangular cusp; apex of cusp not extending past the distal edge of the crown. Mesial cutting edge elongate, continuous, smooth. Distal cutting edge very short, oblique, straight to weakly convex, smooth. Conspicuous distal heel forms acute angle with cusp, separated by distal notch. Cutting edge of distal heel convex to angular, smooth. Labial crown face is flat; lingual face is convex; both crown faces smooth. Root shallow with rounded root lobes. Root lobes extend mesially and distally beyond the base of the root. Tooth base is flat to slightly concave in mesial and distal views. A deep nutritive groove present on lingual root face that often forms a basal notch.
</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFFC8FFE0FF7B9AC14E950A89" lastPageId="79" lastPageNumber="80" pageId="77" pageNumber="78" type="discussion">
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<emphasis id="A2CA2045FFC8FFE2FF7B9AC14C640938" bold="true" box="[189,303,1525,1551]" pageId="77" pageNumber="78">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FFC8FFE2FF7B991149A909C3" blockId="77.[189,1399,1573,1780]" pageId="77" pageNumber="78">
Our
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<emphasis id="A2CA2045FFC8FFE2FF3499114CE40908" box="[242,431,1573,1599]" italics="true" pageId="77" pageNumber="78">Rhizoprionodon</emphasis>
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sample exhibits monognathic, dignathic and gynandric heterodonty. Monognathic heterodonty is expressed as a more erect crown on upper and lower anterior teeth when compared to lateral teeth, and lateral teeth are wider than tall with a more inclined main cusp. With regard to dignathic heterodonty, the cusp of upper teeth is taller and more broadly triangular than on lower teeth. Also, lower anterolateral and lateral teeth have a more elongated mesial edge. Male teeth have a mesiodistally thinner crown than those on females, and the distal edge of the crown is often less convex.
</paragraph>
<paragraph id="9001FC57FFC8FFE0FF7B9828482A0E7E" blockId="77.[189,1399,1820,2028]" lastBlockId="79.[189,1398,267,330]" lastPageId="79" lastPageNumber="80" pageId="77" pageNumber="78">
<bibRefCitation id="F42F81A6FFC8FFE2FF7B98284F520801" author="Thurmond J. T. &amp; Jones D. E." box="[189,537,1820,1846]" pageId="77" pageNumber="78" refId="ref152244" refString="Thurmond J. T. &amp; Jones D. E. 1981. Fossil Vertebrates of Alabama. University of Alabama Press, Tuscaloosa." type="book" year="1981">Thurmond &amp; Jones (1981: 68</bibRefCitation>
, fig. 33) first reported Eocene representatives of
<taxonomicName id="57BE87D4FFC8FFE2FB96982848460801" authorityName="Whitley" authorityYear="1929" box="[1104,1293,1820,1846]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="77" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFC8FFE2FB96982848460801" box="[1104,1293,1820,1846]" italics="true" pageId="77" pageNumber="78">Rhizoprionodon</emphasis>
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in North America when they referred teeth to this taxon that were derived from the basal Gosport Sand at site ACl-
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Clarke County, AL. The presence of this genus within the Claiborne Group of
<collectingRegion id="527A32B5FFC8FFE2FB109850480B0849" box="[1238,1344,1892,1918]" country="United States of America" name="Alabama" pageId="77" pageNumber="78">Alabama</collectingRegion>
was confirmed as part of this study by comparing the teeth in our sample to the dentitions of extant
<taxonomicName id="57BE87D4FFC8FFE2FAD698BC483C0895" box="[1296,1399,1928,1954]" class="Elasmobranchii" family="Carcharhinidae" genus="Loxodon" kingdom="Animalia" order="Carcharhiniformes" pageId="77" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFC8FFE2FAD698BC483C0895" box="[1296,1399,1928,1954]" italics="true" pageId="77" pageNumber="78">Loxodon</emphasis>
</taxonomicName>
(one species),
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<emphasis id="A2CA2045FFC8FFE2FEA098994F6808F0" box="[358,547,1965,1991]" italics="true" pageId="77" pageNumber="78">Rhizoprionodon</emphasis>
</taxonomicName>
(eight species), and
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<emphasis id="A2CA2045FFC8FFE2FCD398994EC108F0" box="[789,906,1965,1991]" italics="true" pageId="77" pageNumber="78">Scoliodon</emphasis>
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(one species) housed at MSC and SC and as illustrated by
<bibRefCitation id="F42F81A6FFC8FFE2FE4698E54F7408DB" author="Springer V. G." box="[384,575,2001,2028]" pageId="77" pageNumber="78" pagination="559 - 632" refId="ref150890" refString="Springer V. G. 1964. A revision of the carcharhinid shark genera Scoliodon, Loxodon, and Rhizoprionodon. Proceedings of the United States National Museum 115 (3493): 559 - 632." type="journal article" year="1964">Springer (1964)</bibRefCitation>
. Of these taxa, the teeth in our sample most closely resemble those of extant species of
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<emphasis id="A2CA2045FFCAFFE0FE4E9E3F4F0E0E12" box="[392,581,267,293]" italics="true" pageId="79" pageNumber="80">Rhizoprionodon</emphasis>
</taxonomicName>
with respect to the shape of the mesial and distal edges of the crown, the mesiodistal thickness of the crown base, overall crown height, and the extent of distal inclination.
</paragraph>
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<emphasis id="A2CA2045FFCBFFE1FF7B9AF44C6D0AED" bold="true" box="[189,294,1472,1498]" pageId="78" pageNumber="79">Fig. 28.</emphasis>
<taxonomicName id="57BE87D4FFCBFFE1FEFF9AF44EFD0AEC" authority="(Arambourg, 1952)" authorityName="Arambourg" authorityYear="1952" baseAuthorityName="Arambourg" baseAuthorityYear="1952" box="[313,950,1472,1499]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="78" pageNumber="79" phylum="Chordata" rank="species" species="ganntourensis">
<emphasis id="A2CA2045FFCBFFE1FEFF9AF44FE40AED" box="[313,687,1472,1498]" italics="true" pageId="78" pageNumber="79">Rhizoprionodon ganntourensis</emphasis>
(
<bibRefCitation id="F42F81A6FFCBFFE1FD0D9AF44EE60AEC" author="Arambourg C." box="[715,941,1472,1499]" pageId="78" pageNumber="79" pagination="1 - 372" refId="ref136183" refString="Arambourg C. 1952. Les vertebres fossiles des gisements de phosphates (Maroc-Algerie-Tunisie). Notes et Memoires du Service geologique du Maroc 92: 1 - 372." type="journal article" year="1952">Arambourg, 1952</bibRefCitation>
)
</taxonomicName>
, teeth.
<emphasis id="A2CA2045FFCBFFE1FBE49AF449100AED" bold="true" box="[1058,1115,1472,1498]" pageId="78" pageNumber="79">AC</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FBB09AF449FF0AED" box="[1142,1204,1472,1498]" pageId="78" pageNumber="79">MSC</collectionCode>
2173.4, lower lateroposterior tooth, basal Gosport Sand.
<emphasis id="A2CA2045FFCBFFE1FD6B9AD74F890ACA" bold="true" box="[685,706,1507,1533]" pageId="78" pageNumber="79">A</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FCAF9AD74E360ACA" bold="true" box="[873,893,1507,1533]" pageId="78" pageNumber="79">B</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FBF59AD749030ACA" bold="true" box="[1075,1096,1507,1533]" pageId="78" pageNumber="79">C</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FB329AD748610ACA" bold="true" box="[1268,1322,1507,1533]" pageId="78" pageNumber="79">DF</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FAFE9AD7483D0ACA" box="[1336,1398,1507,1533]" pageId="78" pageNumber="79">MSC</collectionCode>
37674.3, lower anterior tooth, basal Lisbon Formation.
<emphasis id="A2CA2045FFCBFFE1FC8799324E1D0917" bold="true" box="[833,854,1542,1568]" pageId="78" pageNumber="79">D</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FC3C993249450917" bold="true" box="[1018,1038,1542,1568]" pageId="78" pageNumber="79">E</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FB049932499F0917" bold="true" box="[1218,1236,1542,1568]" pageId="78" pageNumber="79">F</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FF7B991D4DA50974" bold="true" box="[189,238,1577,1603]" pageId="78" pageNumber="79">GI</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FF3B991D4C700974" box="[253,315,1577,1603]" pageId="78" pageNumber="79">MSC</collectionCode>
2173.5, lower lateral tooth, basal Gosport Sand.
<emphasis id="A2CA2045FFCBFFE1FCB8991D4EDE0974" bold="true" box="[894,917,1577,1603]" pageId="78" pageNumber="79">G</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FBFB991E491F0973" bold="true" box="[1085,1108,1578,1604]" pageId="78" pageNumber="79">H</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FACA991E485C0973" bold="true" box="[1292,1303,1578,1604]" pageId="78" pageNumber="79">I</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FEC299794C7E0950" bold="true" box="[260,309,1613,1639]" pageId="78" pageNumber="79">JL</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FE8099794CCF0950" box="[326,388,1613,1639]" pageId="78" pageNumber="79">MSC</collectionCode>
37907.1, lower lateral tooth, lower Tallahatta Formation.
<emphasis id="A2CA2045FFCBFFE1FBFF997949030950" bold="true" box="[1081,1096,1613,1639]" pageId="78" pageNumber="79">J</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FB32997948400950" bold="true" box="[1268,1291,1613,1639]" pageId="78" pageNumber="79">K</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FEC099444C5109BD" bold="true" box="[262,282,1648,1674]" pageId="78" pageNumber="79">L</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FE0899444F4409BD" bold="true" box="[462,527,1648,1674]" pageId="78" pageNumber="79">MO</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FDE699444F1509BD" box="[544,606,1648,1674]" pageId="78" pageNumber="79">MSC</collectionCode>
2173.6, upper anterolateral tooth, basal Gosport Sand.
<emphasis id="A2CA2045FFCBFFE1FB389944485109BD" bold="true" box="[1278,1306,1648,1674]" pageId="78" pageNumber="79">M</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FEC599A74C53099A" bold="true" box="[259,280,1683,1709]" pageId="78" pageNumber="79">N</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FE1699A74CAC099A" bold="true" box="[464,487,1683,1709]" pageId="78" pageNumber="79">O</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FD5599A74F82099A" bold="true" box="[659,713,1683,1709]" pageId="78" pageNumber="79">PR</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FD1E99A74E5D099A" box="[728,790,1683,1709]" pageId="78" pageNumber="79">MSC</collectionCode>
38504.7, lower lateroposterior tooth, basal Gosport Sand.
<emphasis id="A2CA2045FFCBFFE1FEC199824C5209E7" bold="true" box="[263,281,1718,1744]" pageId="78" pageNumber="79">P</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FE0499824C9209E7" bold="true" box="[450,473,1718,1744]" pageId="78" pageNumber="79">Q</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FD5499824FEC09E7" bold="true" box="[658,679,1718,1744]" pageId="78" pageNumber="79">R</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FC9399824EC209E7" bold="true" box="[853,905,1718,1744]" pageId="78" pageNumber="79">SU</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FC5F99824E9C09E7" box="[921,983,1718,1744]" pageId="78" pageNumber="79">MSC</collectionCode>
35753.3, upper anterolateral tooth, lower Tallahatta Formation.
<emphasis id="A2CA2045FFCBFFE1FDC099ED4F5D09C4" bold="true" box="[518,534,1753,1779]" pageId="78" pageNumber="79">S</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FD7F99EE4F8609C3" bold="true" box="[697,717,1754,1780]" pageId="78" pageNumber="79">T</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FC4699EE4EDE09C3" bold="true" box="[896,917,1754,1780]" pageId="78" pageNumber="79">U</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FBFB99EE493D09C3" bold="true" box="[1085,1142,1754,1780]" pageId="78" pageNumber="79">VX</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FB4599ED498A09C4" box="[1155,1217,1753,1779]" pageId="78" pageNumber="79">MSC</collectionCode>
37648.2, lower lateral tooth, basal Gosport Sand.
<emphasis id="A2CA2045FFCBFFE1FD8299C94F120820" bold="true" box="[580,601,1789,1815]" pageId="78" pageNumber="79">V</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FD3D99C94E530820" bold="true" box="[763,792,1789,1815]" pageId="78" pageNumber="79">W</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FC0D99C94EAB0820" bold="true" box="[971,992,1789,1815]" pageId="78" pageNumber="79">X</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FB4199C8499E0821" bold="true" box="[1159,1237,1788,1814]" pageId="78" pageNumber="79">YAA</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FB2499C9486B0820" box="[1250,1312,1789,1815]" pageId="78" pageNumber="79">MSC</collectionCode>
37098, lower lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery.
<emphasis id="A2CA2045FFCBFFE1FB0398144991080D" bold="true" box="[1221,1242,1824,1850]" pageId="78" pageNumber="79">Y</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FF7B98774D9A086A" bold="true" box="[189,209,1859,1885]" pageId="78" pageNumber="79">Z</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FE4898774CF3086A" bold="true" box="[398,440,1859,1885]" pageId="78" pageNumber="79">AA</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FDAC98774F81086A" bold="true" box="[618,714,1859,1885]" pageId="78" pageNumber="79">BBDD</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FD1C98774E53086A" box="[730,792,1859,1885]" pageId="78" pageNumber="79">MSC</collectionCode>
37674.1, lower lateroposterior tooth, basal Lisbon Formation.
<emphasis id="A2CA2045FFCBFFE1FE8398524C2708B7" bold="true" box="[325,364,1894,1920]" pageId="78" pageNumber="79">BB</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FDD698524F7108B7" bold="true" box="[528,570,1894,1920]" pageId="78" pageNumber="79">CC</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FD2898524E5308B7" bold="true" box="[750,792,1894,1920]" pageId="78" pageNumber="79">DD</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FC079852496F08B7" bold="true" box="[961,1060,1894,1920]" pageId="78" pageNumber="79">EEGG</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FBF59852493A08B7" box="[1075,1137,1894,1920]" pageId="78" pageNumber="79">MSC</collectionCode>
37648.1, lower lateral tooth, basal Gosport Sand.
<emphasis id="A2CA2045FFCBFFE1FE3298BE4F500893" bold="true" box="[500,539,1930,1956]" pageId="78" pageNumber="79">EE</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FD7A98BE4FAB0893" bold="true" box="[700,736,1930,1956]" pageId="78" pageNumber="79">FF</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FC5498BD4E8B0894" bold="true" box="[914,960,1929,1955]" pageId="78" pageNumber="79">GG</emphasis>
. Mesial view.
<emphasis id="A2CA2045FFCBFFE1FBA098BE498B0893" bold="true" box="[1126,1216,1930,1956]" pageId="78" pageNumber="79">HHJJ</emphasis>
.
<collectionCode id="F6AF6492FFCBFFE1FB0A98BD48410894" box="[1228,1290,1929,1955]" pageId="78" pageNumber="79">MSC</collectionCode>
37674.4, lower lateral tooth, basal Lisbon Formation.
<emphasis id="A2CA2045FFCBFFE1FD0F98994FBC08F0" bold="true" box="[713,759,1965,1991]" pageId="78" pageNumber="79">HH</emphasis>
. Labial view.
<emphasis id="A2CA2045FFCBFFE1FC5998994EFD08F0" bold="true" box="[927,950,1965,1991]" pageId="78" pageNumber="79">II</emphasis>
. Lingual view.
<emphasis id="A2CA2045FFCBFFE1FBAB989949C108F0" bold="true" box="[1133,1162,1965,1991]" pageId="78" pageNumber="79">JJ</emphasis>
. Mesial view. Scale bars = 2.5 mm.
</paragraph>
</caption>
<paragraph id="9001FC57FFCAFFE0FF7B9E46482C0C2F" blockId="79.[189,1399,370,792]" pageId="79" pageNumber="80">
Examination of the dentitions of Recent female
<taxonomicName id="57BE87D4FFCAFFE0FC859E4648730EBB" authority="(Ruppell, 1837)" baseAuthorityName="Ruppell" baseAuthorityYear="1837" box="[835,1336,370,396]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="acutus">
<emphasis id="A2CA2045FFCAFFE0FC859E46492B0EBC" box="[835,1120,370,396]" italics="true" pageId="79" pageNumber="80">Rhizoprionodon acutus</emphasis>
(Rüppell, 1837)
</taxonomicName>
and
<taxonomicName id="57BE87D4FFCAFFE0FF7B9EA24E4B0E87" authority="(Richardson, 1836)" baseAuthorityName="Richardson" baseAuthorityYear="1836" box="[189,768,405,432]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="terraenovae">
<emphasis id="A2CA2045FFCAFFE0FF7B9EA24F590E98" box="[189,530,406,432]" italics="true" pageId="79" pageNumber="80">Rhizoprionodon terraenovae</emphasis>
(Richardson, 1836)
</taxonomicName>
specimens in the SC collection revealed differences between the distal heels between the two species. The finely serrated distal heel of
<taxonomicName id="57BE87D4FFCAFFE0FB5C9E8F48420EE3" baseAuthorityName="Ruppell" baseAuthorityYear="1837" box="[1178,1289,443,468]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="acutus">
<emphasis id="A2CA2045FFCAFFE0FB5C9E8F48420EE3" box="[1178,1289,443,468]" italics="true" pageId="79" pageNumber="80">R. acutus</emphasis>
</taxonomicName>
teeth are broadly convex, but a single small denticle often provides some angularity, most evident on upper anterior teeth. Teeth in the
<taxonomicName id="57BE87D4FFCAFFE0FDD39D374F810D2B" baseAuthorityName="Richardson" baseAuthorityYear="1836" box="[533,714,515,540]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="terraenovae">
<emphasis id="A2CA2045FFCAFFE0FDD39D374F810D2B" box="[533,714,515,540]" italics="true" pageId="79" pageNumber="80">R. terraenovae</emphasis>
</taxonomicName>
dentition exhibit a similarly serrated distal heel, but a distinctively large denticle results in a more angular appearance than
<taxonomicName id="57BE87D4FFCAFFE0FBD59D1349CE0D77" baseAuthorityName="Ruppell" baseAuthorityYear="1837" box="[1043,1157,551,576]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="acutus">
<emphasis id="A2CA2045FFCAFFE0FBD59D1349CE0D77" box="[1043,1157,551,576]" italics="true" pageId="79" pageNumber="80">R. acutus</emphasis>
</taxonomicName>
, particularly on the upper teeth. We did not observe any significant variation within the Claiborne sample that could indicate multiple species, and we refer all of the material to one taxon. Our Claiborne Group teeth are similar in age and overall morphology to the
<typeStatus id="4F0542F5FFCAFFE0FD999DA64FD90D9B" box="[607,658,658,684]" pageId="79" pageNumber="80">type</typeStatus>
specimens of
<taxonomicName id="57BE87D4FFCAFFE0FCF89DA649E20D9B" baseAuthorityName="Arambourg" baseAuthorityYear="1952" box="[830,1193,658,684]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="ganntourensis">
<emphasis id="A2CA2045FFCAFFE0FCF89DA649E20D9B" box="[830,1193,658,684]" italics="true" pageId="79" pageNumber="80">Rhizoprionodon ganntourensis</emphasis>
</taxonomicName>
as described and figured by
<bibRefCitation id="F42F81A6FFCAFFE0FEF99D824F530DE6" author="Arambourg C." box="[319,536,694,721]" pageId="79" pageNumber="80" pagination="1 - 372" refId="ref136183" refString="Arambourg C. 1952. Les vertebres fossiles des gisements de phosphates (Maroc-Algerie-Tunisie). Notes et Memoires du Service geologique du Maroc 92: 1 - 372." type="journal article" year="1952">Arambourg (1952</bibRefCitation>
: pl. 26, figs 4963), and both samples have teeth with a distal heel that varies in shape from rounded to angular. Due to these similarities, the Claiborne material is assigned to
<taxonomicName id="57BE87D4FFCAFFE0FF7B9DCB4CC80C2F" baseAuthorityName="Arambourg" baseAuthorityYear="1952" box="[189,387,767,792]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="ganntourensis">
<emphasis id="A2CA2045FFCAFFE0FF7B9DCB4CC80C2F" box="[189,387,767,792]" italics="true" pageId="79" pageNumber="80">R. ganntourensis</emphasis>
</taxonomicName>
, currently the only recognized Eocene species within the genus (see
<bibRefCitation id="F42F81A6FFCAFFE0FB6D9DCA48110C2F" author="Cappetta H." box="[1195,1370,766,792]" pageId="79" pageNumber="80" refId="ref137948" refString="Cappetta H. 2012. Chondrichthyes (Mesozoic and Cenozoic Elasmobranchii, Teeth). Handbook of Palaeoichthyology. Verlag Friedrich Pfeil, Munchen." type="book" year="2012">Cappetta 2012</bibRefCitation>
).
</paragraph>
<paragraph id="9001FC57FFCAFFE0FF7B9C744F4E0B60" blockId="79.[189,1399,832,1111]" pageId="79" pageNumber="80">
The dentition of extant
<taxonomicName id="57BE87D4FFCAFFE0FE0D9C744E8C0C6D" authority="Muller &amp; Henle, 1838" authorityName="Muller &amp; Henle" authorityYear="1838" box="[459,967,832,858]" class="Elasmobranchii" family="Carcharhinidae" genus="Scoliodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="laticaudus">
<emphasis id="A2CA2045FFCAFFE0FE0D9C744F8B0C6D" box="[459,704,832,858]" italics="true" pageId="79" pageNumber="80">Scoliodon laticaudus</emphasis>
Müller &amp; Henle, 1838
</taxonomicName>
exhibits gynandric heterodonty, with tooth crowns on female lateral and posterolateral teeth being much more distally inclined than those in the male dentitions (which instead have a thin crown with an upturned apex and an elongated mesial shoulder). Only two species of extant
<taxonomicName id="57BE87D4FFCAFFE0FDB89C984E700CF1" authorityName="Whitley" authorityYear="1929" box="[638,827,940,966]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFCAFFE0FDB89C984E700CF1" box="[638,827,940,966]" italics="true" pageId="79" pageNumber="80">Rhizoprionodon</emphasis>
</taxonomicName>
possess gynandric heterodonty (
<bibRefCitation id="F42F81A6FFCAFFE0FB7A9C9848220CF0" author="Springer V. G." box="[1212,1385,940,967]" pageId="79" pageNumber="80" pagination="559 - 632" refId="ref150890" refString="Springer V. G. 1964. A revision of the carcharhinid shark genera Scoliodon, Loxodon, and Rhizoprionodon. Proceedings of the United States National Museum 115 (3493): 559 - 632." type="journal article" year="1964">Springer 1964</bibRefCitation>
), with male anterolateral teeth having a thinner cusp base than on female teeth. A similar pattern can be observed within our
<taxonomicName id="57BE87D4FFCAFFE0FE7E9CC04F3E0B39" authorityName="Whitley" authorityYear="1929" box="[440,629,1012,1038]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFCAFFE0FE7E9CC04F3E0B39" box="[440,629,1012,1038]" italics="true" pageId="79" pageNumber="80">Rhizoprionodon</emphasis>
</taxonomicName>
sample. Male anterolateral teeth of
<taxonomicName id="57BE87D4FFCAFFE0FBE99CC049A70B39" authorityName="Whitley" authorityYear="1929" box="[1071,1260,1012,1038]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFCAFFE0FBE99CC049A70B39" box="[1071,1260,1012,1038]" italics="true" pageId="79" pageNumber="80">Rhizoprionodon</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="57BE87D4FFCAFFE0FE8E9B2C4CF60B05" authorityName="Muller &amp; Henle" authorityYear="1838" box="[328,445,1048,1074]" class="Elasmobranchii" family="Carcharhinidae" genus="Scoliodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFCAFFE0FE8E9B2C4CF60B05" box="[328,445,1048,1074]" italics="true" pageId="79" pageNumber="80">Scoliodon</emphasis>
</taxonomicName>
(see
<bibRefCitation id="F42F81A6FFCAFFE0FDC79B2C4FA40B05" author="Cappetta H." box="[513,751,1048,1074]" pageId="79" pageNumber="80" refId="ref137948" refString="Cappetta H. 2012. Chondrichthyes (Mesozoic and Cenozoic Elasmobranchii, Teeth). Handbook of Palaeoichthyology. Verlag Friedrich Pfeil, Munchen." type="book" year="2012">Cappetta 2012: 300</bibRefCitation>
, fig. 283), but can be differentiated by their lack of a lingually twisted cusp apex.
</paragraph>
<paragraph id="9001FC57FFCAFFE0FF7B9B4A4E330BD6" blockId="79.[189,1399,1150,1249]" pageId="79" pageNumber="80">
<bibRefCitation id="F42F81A6FFCAFFE0FF7B9B4A4C910BAE" author="Cappetta H. &amp; Case G. R." box="[189,474,1150,1177]" pageId="79" pageNumber="80" pagination="43 - 103" refId="ref137973" refString="Cappetta H. &amp; Case G. R. 2016. A selachian fauna from the middle Eocene (Lutetian, Lisbon Formation) of Andalusia, Covington County, Alabama, USA. Palaeontographica Abteilung A 307 (1 - 6): 43 - 103." type="journal article" year="2016">Cappetta &amp; Case (2016)</bibRefCitation>
did not identify
<taxonomicName id="57BE87D4FFCAFFE0FD669B4A4E160BAF" authorityName="Whitley" authorityYear="1929" box="[672,861,1150,1176]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFCAFFE0FD669B4A4E160BAF" box="[672,861,1150,1176]" italics="true" pageId="79" pageNumber="80">Rhizoprionodon</emphasis>
</taxonomicName>
in their sample of teeth from site ACov-
<quantity id="574651B2FFCAFFE0FAF99B4A483C0BAF" box="[1343,1399,1150,1176]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="79" pageNumber="80" unit="in" value="11.0">11 in</quantity>
Covington County, AL. However, the authors figured three specimens (pl. 10, figs 24) they referred to
<taxonomicName id="57BE87D4FFCAFFE0FF7B9BF24C540BD7" authorityName="Rafinesque" authorityYear="1810" box="[189,287,1222,1248]" class="Elasmobranchii" family="Sphyrnidae" genus="Sphyrna" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFCAFFE0FF7B9BF24C540BD7" box="[189,287,1222,1248]" italics="true" pageId="79" pageNumber="80">Sphyrna</emphasis>
</taxonomicName>
sp. that, in our opinion, belong to
<taxonomicName id="57BE87D4FFCAFFE0FD729BF24E3A0BD7" authorityName="Whitley" authorityYear="1929" box="[692,881,1222,1248]" class="Elasmobranchii" family="Carcharhinidae" genus="Rhizoprionodon" kingdom="Animalia" order="Carcharhiniformes" pageId="79" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FFCAFFE0FD729BF24E3A0BD7" box="[692,881,1222,1248]" italics="true" pageId="79" pageNumber="80">Rhizoprionodon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9001FC57FFCAFFE0FF7B9A3C4E5F0A15" blockId="79.[189,788,1288,1315]" box="[189,788,1288,1315]" pageId="79" pageNumber="80">
<heading id="CB494B3BFFCAFFE0FF7B9A3C4E5F0A15" bold="true" box="[189,788,1288,1315]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="A2CA2045FFCAFFE0FF7B9A3C4E5F0A15" bold="true" box="[189,788,1288,1315]" pageId="79" pageNumber="80">
Stratigraphic and geographic range in
<collectingRegion id="527A32B5FFCAFFE0FD649A3C4E5F0A15" box="[674,788,1288,1314]" country="United States of America" name="Alabama" pageId="79" pageNumber="80">Alabama</collectingRegion>
</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FFCAFFE0FF7B9A0C4E950A89" blockId="79.[189,1399,1336,1471]" pageId="79" pageNumber="80">
The specimens in our sample were collected from the lower Tallahatta Formation at site ADl-1, the Tallahatta Formation at site AMo-8, the contact of the Tallahatta and
<collectingRegion id="527A32B5FFCAFFE0FC359A68490F0A41" box="[1011,1092,1372,1398]" country="Portugal" name="Lisboa" pageId="79" pageNumber="80">Lisbon</collectingRegion>
Formations and the basal
<collectingRegion id="527A32B5FFCAFFE0FF7B9AB44C450AAD" box="[189,270,1408,1434]" country="Portugal" name="Lisboa" pageId="79" pageNumber="80">Lisbon</collectingRegion>
Formation at site ACov-11, the basal Gosport Sand at site ACl-4, and the Gosport Sand and site ACh-21. Upper Ypresian to middle Bartonian, zones NP14 to NP17.
</paragraph>
</subSubSection>
</treatment>
</document>