480 lines
78 KiB
XML
480 lines
78 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E7830FFFBFF013A78ACB1EE5F" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Pseudocentrobolus Wesener & Enghoff & Sierwald 2009, gen. n." docType="treatment" docUuid="4DFE0C0E-5992-4BCC-A6B4-557EC7B6FF65" docUuidSource="ZooBank" docVersion="7" lastPageNumber="62" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="56" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791411" ID-GBIF-Taxon="164184224" ID-Zenodo-Dep="3791411" LSID="urn:lsid:zoobank.org:act:4DFE0C0E-5992-4BCC-A6B4-557EC7B6FF65" httpUri="http://treatment.plazi.org/id/1772122E7830FFFBFF013A78ACB1EE5F" lastPageId="61" lastPageNumber="62" pageId="55" pageNumber="56">
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<subSubSection box="[140,560,1292,1319]" pageId="55" pageNumber="56" type="nomenclature">
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<paragraph blockId="55.[140,983,1292,1355]" box="[140,560,1292,1319]" pageId="55" pageNumber="56">
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<heading box="[140,560,1292,1319]" fontSize="11" level="2" pageId="55" pageNumber="56" reason="3">
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<emphasis bold="true" box="[140,560,1292,1319]" pageId="55" pageNumber="56">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,467,1292,1319]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis bold="true" box="[140,359,1292,1319]" italics="true" pageId="55" pageNumber="56">Pseudocentrobolus</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[479,560,1293,1319]" pageId="55" pageNumber="56" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection box="[140,983,1328,1355]" pageId="55" pageNumber="56" type="description">
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<paragraph blockId="55.[140,983,1292,1355]" box="[140,983,1328,1355]" pageId="55" pageNumber="56">
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<uri box="[140,983,1328,1355]" pageId="55" pageNumber="56">
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urn:lsid:zoobank.org:act:
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<uuid box="[418,983,1328,1355]" pageId="55" pageNumber="56">4DFE0C0E-5992-4BCC-A6B4-557EC7B6FF65</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSection lastPageId="61" lastPageNumber="62" pageId="55" pageNumber="56" type="multiple">
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<subSubSection box="[140,640,1398,1425]" pageId="55" pageNumber="56" type="type_taxon">
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<paragraph blockId="55.[140,640,1398,1425]" box="[140,640,1398,1425]" pageId="55" pageNumber="56">
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<emphasis bold="true" box="[140,287,1398,1425]" pageId="55" pageNumber="56">
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<typeStatus box="[140,198,1399,1425]" pageId="55" pageNumber="56">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[302,571,1399,1425]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[302,571,1399,1425]" italics="true" pageId="55" pageNumber="56">Pseudocentrobolus aureus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[579,640,1399,1425]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="55" pageNumber="56" type="reference_group">
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<paragraph blockId="55.[140,419,1469,1531]" box="[140,419,1469,1496]" pageId="55" pageNumber="56">
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<emphasis bold="true" box="[140,413,1469,1495]" pageId="55" pageNumber="56">Other species included</emphasis>
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:
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</paragraph>
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<paragraph blockId="55.[140,419,1469,1531]" box="[140,371,1504,1531]" pageId="55" pageNumber="56">
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,302,1504,1530]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[140,302,1504,1530]" italics="true" pageId="55" pageNumber="56">P. vohibasiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[310,371,1505,1531]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="diagnosis">
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<paragraph blockId="55.[140,1108,1574,1672]" lastBlockId="56.[140,1108,167,1003]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
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<emphasis bold="true" box="[140,259,1574,1600]" inLineHeading="true" pageId="55" pageNumber="56" reason="1">Diagnosis</emphasis>
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: posterior gonopods are unique, distinct (
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<figureCitation box="[746,831,1574,1601]" captionStart="Figure 35" captionStartId="63.[140,212,1558,1580]" captionTargetBox="[157,1090,165,1538]" captionTargetId="figure@63.[157,1091,165,1539]" captionTargetPageId="63" captionText="Figure 35. Pseudocentrobolus aureus sp. n., male paratype, A–C left, D–E right posterior gonopod. A overview, anterior view B detail of disc, anterior view C membranous process, anterior view D overview, posterior view E detail of disc, posterior view F detail of membranous process, posterior view. Cx = coxite; mem-p = membranous process; St = sternite; T = telopodite." figureDoi="http://doi.org/10.5281/zenodo.3767318" httpUri="https://zenodo.org/record/3767318/files/figure.png" pageId="55" pageNumber="56">Figs 35</figureCitation>
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C–F): telopodite basally with a membranous process carrying sperm canal; telopodite apically with a distinct, unique rounded disc, which is covered with numerous spine-like structures (
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<figureCitation captionStart="Figure 35" captionStartId="63.[140,212,1558,1580]" captionTargetBox="[157,1090,165,1538]" captionTargetId="figure@63.[157,1091,165,1539]" captionTargetPageId="63" captionText="Figure 35. Pseudocentrobolus aureus sp. n., male paratype, A–C left, D–E right posterior gonopod. A overview, anterior view B detail of disc, anterior view C membranous process, anterior view D overview, posterior view E detail of disc, posterior view F detail of membranous process, posterior view. Cx = coxite; mem-p = membranous process; St = sternite; T = telopodite." figureDoi="http://doi.org/10.5281/zenodo.3767318" httpUri="https://zenodo.org/record/3767318/files/figure.png" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">Figs 35B, E</figureCitation>
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); coxite basally at sperm canal with an obligate finger-shaped process, which is not curved but straight like in
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[440,580,202,228]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[440,580,202,228]" italics="true" pageId="56" pageNumber="57">Granitobolus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[588,665,202,228]" pageId="56" pageNumber="57" rank="genus">gen. n.</taxonomicNameLabel>
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Natural coloration conspicuously contrasting, with golden, black or red pattern (
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<figureCitation box="[641,824,237,264]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="56" pageNumber="57">Figs 31A, C, D</figureCitation>
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, 36B). Gnathochilarium with a subdivided mentum and a single sclerotized ledge on each stipites (
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<figureCitation box="[955,1038,272,298]" captionStart="Figure 32" captionStartId="58.[140,212,1526,1548]" captionTargetBox="[190,1057,165,1506]" captionTargetId="figure@58.[190,1058,165,1507]" captionTargetPageId="58" captionText="Figure 32. Pseudocentrobolus aureus sp. n. male paratype, SEM. A antenna, apical view B antenna, overview C antennomere 5 with sensilla basiconica; D gnathochilarium, ventral view E gnathochilarium, dorsal view F mentum of gnathochilarium, ventral view G endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="56" pageNumber="57">Figs 32</figureCitation>
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D–F), like in
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[219,357,307,333]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis box="[219,357,307,333]" italics="true" pageId="56" pageNumber="57">Flagellobolus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[372,515,307,333]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis box="[372,515,307,333]" italics="true" pageId="56" pageNumber="57">Riotintobolus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[530,670,307,333]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[530,670,307,333]" italics="true" pageId="56" pageNumber="57">Granitobolus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[678,756,308,334]" pageId="56" pageNumber="57" rank="genus">gen. n.</taxonomicNameLabel>
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,
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[771,893,307,333]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[771,893,307,333]" italics="true" pageId="56" pageNumber="57">Caprobolus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[901,979,308,334]" pageId="56" pageNumber="57" rank="genus">gen. n.</taxonomicNameLabel>
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,
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis italics="true" pageId="56" pageNumber="57">Alluviobolus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[179,259,343,369]" pageId="56" pageNumber="57" rank="genus">gen. n.</taxonomicNameLabel>
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,
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[276,403,343,369]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[276,403,343,369]" italics="true" pageId="56" pageNumber="57">Ostinobolus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[413,496,343,369]" pageId="56" pageNumber="57" rank="genus">
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<emphasis bold="true" box="[413,489,343,369]" pageId="56" pageNumber="57">gen. n</emphasis>
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.
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</taxonomicNameLabel>
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Shares numerous characters only with species of the genus
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[211,351,378,404]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. n.">
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||
<emphasis box="[211,351,378,404]" italics="true" pageId="56" pageNumber="57">Granitobolus</emphasis>
|
||
</taxonomicName>
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<taxonomicNameLabel box="[357,436,378,404]" pageId="56" pageNumber="57" rank="genus">gen. n.</taxonomicNameLabel>
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: a stout preanal process, which prominently extends beyond anal valves (
|
||
<figureCitation box="[276,383,413,439]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="56" pageNumber="57">Figs 31C</figureCitation>
|
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, 36B), well-developed lips on the anal valves, thick moniliform body rings which overlap each widely to the metazonite (
|
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<figureCitation box="[780,884,448,474]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="56" pageNumber="57">Fig. 31C</figureCitation>
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), rectangular extensions (
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<figureCitation box="[210,344,483,510]" captionStart="Figure 33" captionStartId="60.[140,212,1558,1580]" captionTargetBox="[170,1078,339,1541]" captionTargetId="figure@60.[170,1079,339,1541]" captionTargetPageId="60" captionText="Figure 33. Pseudocentrobolus aureus sp. n. A–C male, D female paratype, SEM. A left mandible, mesal view B midbody ring, overview C midbody ring, detail of metazona D right vulva, lateral view. 3iT = 3-combed internal tooth; av = anterior valve; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; re = rectangular excavations." figureDoi="http://doi.org/10.5281/zenodo.3767314" httpUri="https://zenodo.org/record/3767314/files/figure.png" pageId="56" pageNumber="57">Figs 33B, C</figureCitation>
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) on the posterior margins of the rings and a greatly enlarged collum, which is ventrally longer than the body ring 2. The habitus and shape of the anterior gonopod (
|
||
<figureCitation box="[256,391,554,580]" captionStart="Figure 34" captionStartId="61.[140,212,641,663]" captionTargetBox="[140,1108,165,621]" captionTargetId="figure@61.[140,1108,165,621]" captionTargetPageId="61" captionText="Figure 34. Pseudocentrobolus aureus sp. n., male holotype, anterior gonopod A anterior view B posterior view. Cx = coxite; St = sternite; T = telopodite. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767316" httpUri="https://zenodo.org/record/3767316/files/figure.png" pageId="56" pageNumber="57">Figs 34A, B</figureCitation>
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) are also very similar to species of
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[780,920,554,580]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[780,920,554,580]" italics="true" pageId="56" pageNumber="57">Granitobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[927,1005,554,580]" pageId="56" pageNumber="57" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="56" pageNumber="57" type="distribution">
|
||
<paragraph blockId="56.[140,1108,167,1003]" pageId="56" pageNumber="57">
|
||
<emphasis bold="true" box="[185,486,589,615]" pageId="56" pageNumber="57">
|
||
<emphasis box="[185,336,589,615]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Distribution</emphasis>
|
||
and ecology
|
||
</emphasis>
|
||
: two species of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[663,856,589,615]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[663,856,589,615]" italics="true" pageId="56" pageNumber="57">Pseudocentrobolus</emphasis>
|
||
</taxonomicName>
|
||
are recorded from the dry spiny and subhumid forests in Southwestern
|
||
<collectingCountry box="[687,820,624,650]" name="Madagascar" pageId="56" pageNumber="57">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation box="[834,918,624,650]" captionStart="Figure 30" captionStartId="56.[140,210,1654,1676]" captionTargetBox="[129,1120,1092,1644]" captionTargetId="figure@56.[140,1108,1098,1642]" captionTargetPageId="56" captionText="Figure 30. Distribution of the genera Pseudocentrobolus gen. n., Granitobolus gen. n. and Caprobolus gen.n." figureDoi="http://doi.org/10.5281/zenodo.3820818" httpUri="https://zenodo.org/record/3820818/files/figure.png" pageId="56" pageNumber="57">Fig. 30</figureCitation>
|
||
). A third species, whose external characters fit very well into
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[615,808,659,685]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[615,808,659,685]" italics="true" pageId="56" pageNumber="57">Pseudocentrobolus</emphasis>
|
||
</taxonomicName>
|
||
but where no mature male could be collected, was discovered in the spiny forest of Andohahela in Southeastern
|
||
<collectingCountry box="[140,273,730,756]" name="Madagascar" pageId="56" pageNumber="57">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation box="[288,372,730,756]" captionStart="Figure 30" captionStartId="56.[140,210,1654,1676]" captionTargetBox="[129,1120,1092,1644]" captionTargetId="figure@56.[140,1108,1098,1642]" captionTargetPageId="56" captionText="Figure 30. Distribution of the genera Pseudocentrobolus gen. n., Granitobolus gen. n. and Caprobolus gen.n." figureDoi="http://doi.org/10.5281/zenodo.3820818" httpUri="https://zenodo.org/record/3820818/files/figure.png" pageId="56" pageNumber="57">Fig. 30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="61" lastPageNumber="62" pageId="56" pageNumber="57" type="description">
|
||
<paragraph blockId="56.[140,1108,167,1003]" pageId="56" pageNumber="57">
|
||
<emphasis bold="true" box="[185,335,765,791]" pageId="56" pageNumber="57">
|
||
<emphasis box="[185,330,765,791]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[342,406,765,791]" italics="true" pageId="56" pageNumber="57">Males</emphasis>
|
||
: length up to
|
||
<quantity box="[567,653,765,792]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="56" pageNumber="57" unit="mm" value="48.0">48 mm</quantity>
|
||
, diameter up to
|
||
<quantity box="[839,933,765,792]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="56" pageNumber="57" unit="mm" value="4.5">4.5 mm</quantity>
|
||
. 38–40 podous rings. Females: length up to
|
||
<quantity box="[465,551,800,827]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="56" pageNumber="57" unit="mm" value="48.0">48 mm</quantity>
|
||
, diameter up to
|
||
<quantity box="[740,835,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="56" pageNumber="57" unit="mm" value="5.1">5.1 mm</quantity>
|
||
. 39 or 40 podous rings. Apodous rings absent in all adult specimens.
|
||
</paragraph>
|
||
<paragraph blockId="56.[140,1108,167,1003]" pageId="56" pageNumber="57">
|
||
<emphasis box="[185,260,871,897]" italics="true" pageId="56" pageNumber="57">Colour</emphasis>
|
||
(after short time in alcohol): mesozona golden (
|
||
<figureCitation box="[805,906,871,897]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="56" pageNumber="57">Fig. 31A</figureCitation>
|
||
) or greenish (fad- ed to grey in some specimens), metazona blackish (
|
||
<figureCitation box="[739,842,906,932]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="56" pageNumber="57">Fig. 31A</figureCitation>
|
||
) or golden (
|
||
<figureCitation box="[989,1095,906,933]" captionStart="Figure 36" captionStartId="64.[140,212,1526,1548]" captionTargetBox="[140,1108,692,1507]" captionTargetId="figure@64.[140,1108,692,1109]" captionTargetPageId="64" captionText="Figure 36. Pseudocentrobolus vohibasiensis sp. n., male holotype. A head and legs 1–7, ventral view B telson C anterior gonopod, anterior view D posterior gonopods, posterior view E posterior gonopods, anterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; scl-p = sclerotized process; St = sternite; T = telopodite. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767320" httpUri="https://zenodo.org/record/3767320/files/figure.png" pageId="56" pageNumber="57">Fig. 36B</figureCitation>
|
||
). Head, antennae, gonopods, legs and at least posterior part of telson golden (
|
||
<figureCitation box="[997,1099,941,967]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="56" pageNumber="57">Fig. 31C</figureCitation>
|
||
) or reddish (
|
||
<figureCitation box="[271,375,976,1003]" captionStart="Figure 36" captionStartId="64.[140,212,1526,1548]" captionTargetBox="[140,1108,692,1507]" captionTargetId="figure@64.[140,1108,692,1109]" captionTargetPageId="64" captionText="Figure 36. Pseudocentrobolus vohibasiensis sp. n., male holotype. A head and legs 1–7, ventral view B telson C anterior gonopod, anterior view D posterior gonopods, posterior view E posterior gonopods, anterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; scl-p = sclerotized process; St = sternite; T = telopodite. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767320" httpUri="https://zenodo.org/record/3767320/files/figure.png" pageId="56" pageNumber="57">Fig. 36B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820818" ID-Zenodo-Dep="3820818" box="[140,1108,1654,1676]" httpUri="https://zenodo.org/record/3820818/files/figure.png" pageId="56" pageNumber="57" startId="56.[140,210,1654,1676]" targetBox="[129,1120,1092,1644]" targetPageId="56">
|
||
<paragraph blockId="56.[140,1108,1654,1676]" box="[140,1108,1654,1676]" pageId="56" pageNumber="57">
|
||
<emphasis bold="true" box="[140,248,1654,1676]" pageId="56" pageNumber="57">Figure 30.</emphasis>
|
||
Distribution of the genera
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[491,646,1654,1675]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[491,646,1654,1675]" italics="true" pageId="56" pageNumber="57">Pseudocentrobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[651,713,1655,1676]" pageId="56" pageNumber="57" rank="genus">gen. n.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[723,836,1654,1675]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[723,836,1654,1675]" italics="true" pageId="56" pageNumber="57">Granitobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[840,901,1655,1676]" pageId="56" pageNumber="57" rank="genus">gen. n.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[944,1042,1654,1675]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[944,1042,1654,1675]" italics="true" pageId="56" pageNumber="57">Caprobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1047,1108,1655,1676]" pageId="56" pageNumber="57" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="57.[140,1108,167,404]" pageId="57" pageNumber="58">
|
||
<emphasis box="[185,244,167,193]" italics="true" pageId="57" pageNumber="58">Head</emphasis>
|
||
: each eye with circa 38–42 ocelli arranged in 7 or 8 vertical rows (
|
||
<figureCitation box="[994,1102,167,193]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="57" pageNumber="58">Figs 31D</figureCitation>
|
||
, 36A). Labrum with standard three irregular teeth and one row of 10–12 stout marginal setae (
|
||
<figureCitation box="[209,312,237,264]" captionStart="Figure 36" captionStartId="64.[140,212,1526,1548]" captionTargetBox="[140,1108,692,1507]" captionTargetId="figure@64.[140,1108,692,1109]" captionTargetPageId="64" captionText="Figure 36. Pseudocentrobolus vohibasiensis sp. n., male holotype. A head and legs 1–7, ventral view B telson C anterior gonopod, anterior view D posterior gonopods, posterior view E posterior gonopods, anterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; scl-p = sclerotized process; St = sternite; T = telopodite. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767320" httpUri="https://zenodo.org/record/3767320/files/figure.png" pageId="57" pageNumber="58">Fig. 36A</figureCitation>
|
||
). Clypeus with two setiferous foveolae on each side. Antennae short or very short, reaching back to ring 2 or 4 (
|
||
<figureCitation box="[596,705,272,298]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="57" pageNumber="58">Figs 31D</figureCitation>
|
||
, 36A). Relative lengths of antennomeres: 1<2=3=4=5=6 (
|
||
<figureCitation box="[396,499,307,334]" captionStart="Figure 32" captionStartId="58.[140,212,1526,1548]" captionTargetBox="[190,1057,165,1506]" captionTargetId="figure@58.[190,1058,165,1507]" captionTargetPageId="58" captionText="Figure 32. Pseudocentrobolus aureus sp. n. male paratype, SEM. A antenna, apical view B antenna, overview C antennomere 5 with sensilla basiconica; D gnathochilarium, ventral view E gnathochilarium, dorsal view F mentum of gnathochilarium, ventral view G endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="57" pageNumber="58">Fig. 32B</figureCitation>
|
||
). Terminal antennomere with four large sensory cones located together inside a membranous area (
|
||
<figureCitation box="[640,744,343,369]" captionStart="Figure 32" captionStartId="58.[140,212,1526,1548]" captionTargetBox="[190,1057,165,1506]" captionTargetId="figure@58.[190,1058,165,1507]" captionTargetPageId="58" captionText="Figure 32. Pseudocentrobolus aureus sp. n. male paratype, SEM. A antenna, apical view B antenna, overview C antennomere 5 with sensilla basiconica; D gnathochilarium, ventral view E gnathochilarium, dorsal view F mentum of gnathochilarium, ventral view G endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="57" pageNumber="58">Fig. 32A</figureCitation>
|
||
). Antennomere 5 latero-apically with four rows, antennomere 6 with two rows (
|
||
<figureCitation box="[673,811,378,404]" captionStart="Figure 32" captionStartId="58.[140,212,1526,1548]" captionTargetBox="[190,1057,165,1506]" captionTargetId="figure@58.[190,1058,165,1507]" captionTargetPageId="58" captionText="Figure 32. Pseudocentrobolus aureus sp. n. male paratype, SEM. A antenna, apical view B antenna, overview C antennomere 5 with sensilla basiconica; D gnathochilarium, ventral view E gnathochilarium, dorsal view F mentum of gnathochilarium, ventral view G endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="57" pageNumber="58">Figs 32A, C</figureCitation>
|
||
) of sensilla basiconica.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820820" ID-Zenodo-Dep="3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="57" pageNumber="58" startId="57.[140,212,1494,1516]" targetBox="[198,1049,450,1475]" targetPageId="57">
|
||
<paragraph blockId="57.[140,1109,1494,1676]" pageId="57" pageNumber="58">
|
||
<emphasis bold="true" box="[140,253,1494,1516]" pageId="57" pageNumber="58">Figure 31.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[261,484,1494,1516]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[261,484,1494,1516]" italics="true" pageId="57" pageNumber="58">Pseudocentrobolus aureus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[492,545,1495,1516]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
|
||
, A–D, G, H male
|
||
<typeStatus box="[728,810,1495,1516]" pageId="57" pageNumber="58">paratype</typeStatus>
|
||
; E, F female
|
||
<typeStatus box="[938,1020,1495,1516]" pageId="57" pageNumber="58">paratype</typeStatus>
|
||
.
|
||
<emphasis bold="true" box="[1031,1050,1494,1516]" pageId="57" pageNumber="58">A</emphasis>
|
||
head, lateral view
|
||
<emphasis bold="true" box="[249,266,1526,1548]" pageId="57" pageNumber="58">B</emphasis>
|
||
legs 1–7, ventral view
|
||
<emphasis bold="true" box="[475,494,1526,1548]" pageId="57" pageNumber="58">C</emphasis>
|
||
telson
|
||
<emphasis bold="true" box="[559,578,1526,1548]" pageId="57" pageNumber="58">D</emphasis>
|
||
head, antenna and collum, lateral view
|
||
<emphasis bold="true" box="[944,959,1526,1548]" pageId="57" pageNumber="58">E</emphasis>
|
||
second leg with vulva
|
||
<emphasis bold="true" box="[195,209,1558,1580]" pageId="57" pageNumber="58">F</emphasis>
|
||
female midbody ring with legs
|
||
<emphasis bold="true" box="[505,525,1558,1580]" pageId="57" pageNumber="58">G</emphasis>
|
||
posterior gonopods, anterior view
|
||
<emphasis bold="true" box="[850,870,1558,1580]" pageId="57" pageNumber="58">H</emphasis>
|
||
posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars =
|
||
<quantity box="[254,313,1655,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="57" pageNumber="58" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767312" ID-Zenodo-Dep="3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="58" pageNumber="59" startId="58.[140,212,1526,1548]" targetBox="[190,1057,165,1506]" targetPageId="58">
|
||
<paragraph pageId="58" pageNumber="59">
|
||
<emphasis bold="true" box="[140,254,1526,1548]" pageId="58" pageNumber="59">Figure 32.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[264,488,1526,1548]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[264,488,1526,1548]" italics="true" pageId="58" pageNumber="59">Pseudocentrobolus aureus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[497,551,1527,1548]" pageId="58" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
|
||
male paratype, SEM.
|
||
<emphasis bold="true" box="[770,789,1526,1548]" pageId="58" pageNumber="59">A</emphasis>
|
||
antenna, apical view
|
||
<emphasis bold="true" box="[1001,1018,1526,1548]" pageId="58" pageNumber="59">B</emphasis>
|
||
antenna, overview
|
||
<emphasis bold="true" box="[227,246,1558,1580]" pageId="58" pageNumber="59">C</emphasis>
|
||
antennomere 5 with sensilla basiconica;
|
||
<emphasis bold="true" box="[621,640,1558,1580]" pageId="58" pageNumber="59">D</emphasis>
|
||
gnathochilarium, ventral view
|
||
<emphasis bold="true" box="[927,942,1558,1580]" pageId="58" pageNumber="59">E</emphasis>
|
||
gnathochilarium, dorsal view
|
||
<emphasis bold="true" box="[252,266,1590,1612]" pageId="58" pageNumber="59">F</emphasis>
|
||
mentum of gnathochilarium, ventral view
|
||
<emphasis bold="true" box="[672,692,1590,1612]" pageId="58" pageNumber="59">G</emphasis>
|
||
endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="59.[140,1108,167,1672]" pageId="59" pageNumber="60">
|
||
<emphasis box="[185,373,167,193]" italics="true" pageId="59" pageNumber="60">Gnathochilarium</emphasis>
|
||
unusual (
|
||
<figureCitation box="[489,598,167,193]" captionStart="Figure 32" captionStartId="58.[140,212,1526,1548]" captionTargetBox="[190,1057,165,1506]" captionTargetId="figure@58.[190,1058,165,1507]" captionTargetPageId="58" captionText="Figure 32. Pseudocentrobolus aureus sp. n. male paratype, SEM. A antenna, apical view B antenna, overview C antennomere 5 with sensilla basiconica; D gnathochilarium, ventral view E gnathochilarium, dorsal view F mentum of gnathochilarium, ventral view G endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="59" pageNumber="60">Fig. 32D</figureCitation>
|
||
). Lamellae linguales each with two standard setae located behind one another. Stipites each with three apical setae (
|
||
<figureCitation box="[983,1094,202,228]" captionStart="Figure 32" captionStartId="58.[140,212,1526,1548]" captionTargetBox="[190,1057,165,1506]" captionTargetId="figure@58.[190,1058,165,1507]" captionTargetPageId="58" captionText="Figure 32. Pseudocentrobolus aureus sp. n. male paratype, SEM. A antenna, apical view B antenna, overview C antennomere 5 with sensilla basiconica; D gnathochilarium, ventral view E gnathochilarium, dorsal view F mentum of gnathochilarium, ventral view G endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="59" pageNumber="60">Fig. 32D</figureCitation>
|
||
). Mentum basally subdivided by a well-developed suture (
|
||
<figureCitation box="[795,940,237,264]" captionStart="Figure 32" captionStartId="58.[140,212,1526,1548]" captionTargetBox="[190,1057,165,1506]" captionTargetId="figure@58.[190,1058,165,1507]" captionTargetPageId="58" captionText="Figure 32. Pseudocentrobolus aureus sp. n. male paratype, SEM. A antenna, apical view B antenna, overview C antennomere 5 with sensilla basiconica; D gnathochilarium, ventral view E gnathochilarium, dorsal view F mentum of gnathochilarium, ventral view G endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="59" pageNumber="60">Figs 32D, F</figureCitation>
|
||
). Stipites each towards mentum with a wide sclerotized ledge (
|
||
<figureCitation box="[691,800,272,298]" captionStart="Figure 32" captionStartId="58.[140,212,1526,1548]" captionTargetBox="[190,1057,165,1506]" captionTargetId="figure@58.[190,1058,165,1507]" captionTargetPageId="58" captionText="Figure 32. Pseudocentrobolus aureus sp. n. male paratype, SEM. A antenna, apical view B antenna, overview C antennomere 5 with sensilla basiconica; D gnathochilarium, ventral view E gnathochilarium, dorsal view F mentum of gnathochilarium, ventral view G endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="59" pageNumber="60">Fig. 32D</figureCitation>
|
||
). Palpi of gnathochilarium with numerous sensilla. Hypopharyngeal crest with a field of spine-like structures (
|
||
<figureCitation captionStart="Figure 32" captionStartId="58.[140,212,1526,1548]" captionTargetBox="[190,1057,165,1506]" captionTargetId="figure@58.[190,1058,165,1507]" captionTargetPageId="58" captionText="Figure 32. Pseudocentrobolus aureus sp. n. male paratype, SEM. A antenna, apical view B antenna, overview C antennomere 5 with sensilla basiconica; D gnathochilarium, ventral view E gnathochilarium, dorsal view F mentum of gnathochilarium, ventral view G endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="59" pageNumber="60">Fig. 32E</figureCitation>
|
||
). Central pads of endochilarium separated into two areas: a group of 8–10 sensilla located apically, and medial palps with numerous sensilla each, located closely towards hypopharyngeal crest (
|
||
<figureCitation box="[395,502,413,439]" captionStart="Figure 32" captionStartId="58.[140,212,1526,1548]" captionTargetBox="[190,1057,165,1506]" captionTargetId="figure@58.[190,1058,165,1507]" captionTargetPageId="58" captionText="Figure 32. Pseudocentrobolus aureus sp. n. male paratype, SEM. A antenna, apical view B antenna, overview C antennomere 5 with sensilla basiconica; D gnathochilarium, ventral view E gnathochilarium, dorsal view F mentum of gnathochilarium, ventral view G endochilarium, dorsal view, arrow points to sensory cones. cp = endochilarium central pads; Endo = endochilarium; L = ledge; LL = lamella lingualis; Me = mentum; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767312" httpUri="https://zenodo.org/record/3767312/files/figure.png" pageId="59" pageNumber="60">Fig. 32G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="59.[140,1108,167,1672]" pageId="59" pageNumber="60">
|
||
<emphasis box="[185,291,448,474]" italics="true" pageId="59" pageNumber="60">Mandible</emphasis>
|
||
: external tooth simple, rounded; mesal tooth with three cusps (
|
||
<figureCitation box="[993,1095,448,474]" captionStart="Figure 33" captionStartId="60.[140,212,1558,1580]" captionTargetBox="[170,1078,339,1541]" captionTargetId="figure@60.[170,1079,339,1541]" captionTargetPageId="60" captionText="Figure 33. Pseudocentrobolus aureus sp. n. A–C male, D female paratype, SEM. A left mandible, mesal view B midbody ring, overview C midbody ring, detail of metazona D right vulva, lateral view. 3iT = 3-combed internal tooth; av = anterior valve; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; re = rectangular excavations." figureDoi="http://doi.org/10.5281/zenodo.3767314" httpUri="https://zenodo.org/record/3767314/files/figure.png" pageId="59" pageNumber="60">Fig. 33A</figureCitation>
|
||
). Six pectinate lamellae. Mesal margin of pectinate area (intermediate area) with circa four rows of small, slender spines. Molar plate with numerous (10 or 11) transverse furrows (
|
||
<figureCitation box="[241,345,554,580]" captionStart="Figure 33" captionStartId="60.[140,212,1558,1580]" captionTargetBox="[170,1078,339,1541]" captionTargetId="figure@60.[170,1079,339,1541]" captionTargetPageId="60" captionText="Figure 33. Pseudocentrobolus aureus sp. n. A–C male, D female paratype, SEM. A left mandible, mesal view B midbody ring, overview C midbody ring, detail of metazona D right vulva, lateral view. 3iT = 3-combed internal tooth; av = anterior valve; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; re = rectangular excavations." figureDoi="http://doi.org/10.5281/zenodo.3767314" httpUri="https://zenodo.org/record/3767314/files/figure.png" pageId="59" pageNumber="60">Fig. 33A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="59.[140,1108,167,1672]" pageId="59" pageNumber="60">
|
||
<emphasis box="[185,274,589,615]" italics="true" pageId="59" pageNumber="60">Collum:</emphasis>
|
||
smooth, laterally longer than body ring 2 (
|
||
<figureCitation box="[752,860,589,615]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="59" pageNumber="60">Figs 31D</figureCitation>
|
||
, 36A). Lobe enlarged, laterally covering antenna and part of head (
|
||
<figureCitation box="[638,744,624,650]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="59" pageNumber="60">Fig. 31D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="59.[140,1108,167,1672]" pageId="59" pageNumber="60">
|
||
<emphasis box="[185,306,659,685]" italics="true" pageId="59" pageNumber="60">Body rings:</emphasis>
|
||
moniliform, overlapping one another up to anterior half of mesozona. Metazona of swollen appearance, width greater than those of pro- or mesozona. Transverse impressions covering mesozona (
|
||
<figureCitation box="[573,675,730,756]" captionStart="Figure 33" captionStartId="60.[140,212,1558,1580]" captionTargetBox="[170,1078,339,1541]" captionTargetId="figure@60.[170,1079,339,1541]" captionTargetPageId="60" captionText="Figure 33. Pseudocentrobolus aureus sp. n. A–C male, D female paratype, SEM. A left mandible, mesal view B midbody ring, overview C midbody ring, detail of metazona D right vulva, lateral view. 3iT = 3-combed internal tooth; av = anterior valve; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; re = rectangular excavations." figureDoi="http://doi.org/10.5281/zenodo.3767314" httpUri="https://zenodo.org/record/3767314/files/figure.png" pageId="59" pageNumber="60">Fig. 33B</figureCitation>
|
||
), metazona with irregular punctations (
|
||
<figureCitation box="[148,257,765,791]" captionStart="Figure 33" captionStartId="60.[140,212,1558,1580]" captionTargetBox="[170,1078,339,1541]" captionTargetId="figure@60.[170,1079,339,1541]" captionTargetPageId="60" captionText="Figure 33. Pseudocentrobolus aureus sp. n. A–C male, D female paratype, SEM. A left mandible, mesal view B midbody ring, overview C midbody ring, detail of metazona D right vulva, lateral view. 3iT = 3-combed internal tooth; av = anterior valve; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; re = rectangular excavations." figureDoi="http://doi.org/10.5281/zenodo.3767314" httpUri="https://zenodo.org/record/3767314/files/figure.png" pageId="59" pageNumber="60">Fig. 33C</figureCitation>
|
||
). Posterior margin of rings slightly excavated into rectangular extensions (
|
||
<figureCitation box="[148,253,800,826]" captionStart="Figure 33" captionStartId="60.[140,212,1558,1580]" captionTargetBox="[170,1078,339,1541]" captionTargetId="figure@60.[170,1079,339,1541]" captionTargetPageId="60" captionText="Figure 33. Pseudocentrobolus aureus sp. n. A–C male, D female paratype, SEM. A left mandible, mesal view B midbody ring, overview C midbody ring, detail of metazona D right vulva, lateral view. 3iT = 3-combed internal tooth; av = anterior valve; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; re = rectangular excavations." figureDoi="http://doi.org/10.5281/zenodo.3767314" httpUri="https://zenodo.org/record/3767314/files/figure.png" pageId="59" pageNumber="60">Fig. 33C</figureCitation>
|
||
). Ozopores starting at ring 6, touching suture between mesozona and metazona (
|
||
<figureCitation box="[208,312,835,862]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="59" pageNumber="60">Fig. 31C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="59.[140,1108,167,1672]" pageId="59" pageNumber="60">
|
||
<emphasis box="[185,251,871,897]" italics="true" pageId="59" pageNumber="60">Telson</emphasis>
|
||
: anal valves with well-developed lips and micropunctation but with neither grooves nor setae. Preanal process well-rounded or sharp, extending beyond anal valves (
|
||
<figureCitation box="[148,252,941,967]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="59" pageNumber="60">Fig. 31C</figureCitation>
|
||
). Subanal scale inconspicuous.
|
||
</paragraph>
|
||
<paragraph blockId="59.[140,1108,167,1672]" pageId="59" pageNumber="60">
|
||
<emphasis box="[185,232,977,1002]" italics="true" pageId="59" pageNumber="60">Legs</emphasis>
|
||
: coxae 1 and 2 elongated and fused with sternum, podomeres from prefemur to tarsus in both sexes each with 4–10 ventral/mesal setae (
|
||
<figureCitation box="[896,1003,1011,1038]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="59" pageNumber="60">Fig. 31E</figureCitation>
|
||
). Length of midbody legs circa 0.6 times body diameter in both sexes (
|
||
<figureCitation box="[873,978,1047,1073]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="59" pageNumber="60">Fig. 31F</figureCitation>
|
||
). Each podomere with an apical, ventral seta. Coxae 3 and beyond of rectangular shape (
|
||
<figureCitation captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="59" pageNumber="60">Fig. 31F</figureCitation>
|
||
). Tarsus with a stout dorso-apical setae and a pair of stout ventro-apical ones in males, in females with three pair of setae, apical pair stout, others successively much smaller.
|
||
</paragraph>
|
||
<paragraph blockId="59.[140,1108,167,1672]" pageId="59" pageNumber="60">
|
||
<emphasis box="[185,423,1223,1249]" italics="true" pageId="59" pageNumber="60">Male sexual characters</emphasis>
|
||
: tarsi 3 and beyond with a tarsal pad not protruding past base of claw. Coxae 3–7 with species-specific modifications.
|
||
</paragraph>
|
||
<paragraph blockId="59.[140,1108,167,1672]" pageId="59" pageNumber="60">
|
||
<emphasis box="[185,389,1293,1319]" italics="true" pageId="59" pageNumber="60">Anterior gonopods</emphasis>
|
||
: median sternal projection triangular, with a wide rectangular tip (
|
||
<figureCitation box="[225,334,1328,1355]" captionStart="Figure 35" captionStartId="63.[140,212,1558,1580]" captionTargetBox="[157,1090,165,1538]" captionTargetId="figure@63.[157,1091,165,1539]" captionTargetPageId="63" captionText="Figure 35. Pseudocentrobolus aureus sp. n., male paratype, A–C left, D–E right posterior gonopod. A overview, anterior view B detail of disc, anterior view C membranous process, anterior view D overview, posterior view E detail of disc, posterior view F detail of membranous process, posterior view. Cx = coxite; mem-p = membranous process; St = sternite; T = telopodite." figureDoi="http://doi.org/10.5281/zenodo.3767318" httpUri="https://zenodo.org/record/3767318/files/figure.png" pageId="59" pageNumber="60">Figs 35A</figureCitation>
|
||
,
|
||
<figureCitation box="[345,401,1328,1354]" captionStart="Figure 49" captionStartId="87.[139,211,1654,1676]" captionTargetBox="[141,1112,1220,1629]" captionTargetId="figure@87.[139,1107,1228,1634]" captionTargetPageId="87" captionText="Figure 49. Distribution map of the genus Ostinobolus gen. n.." figureDoi="http://doi.org/10.5281/zenodo.3767340" httpUri="https://zenodo.org/record/3767340/files/figure.png" pageId="59" pageNumber="60">49C</figureCitation>
|
||
). Sternite protruding almost as high as coxite and telopodite (
|
||
<figureCitation box="[148,260,1363,1390]" captionStart="Figure 35" captionStartId="63.[140,212,1558,1580]" captionTargetBox="[157,1090,165,1538]" captionTargetId="figure@63.[157,1091,165,1539]" captionTargetPageId="63" captionText="Figure 35. Pseudocentrobolus aureus sp. n., male paratype, A–C left, D–E right posterior gonopod. A overview, anterior view B detail of disc, anterior view C membranous process, anterior view D overview, posterior view E detail of disc, posterior view F detail of membranous process, posterior view. Cx = coxite; mem-p = membranous process; St = sternite; T = telopodite." figureDoi="http://doi.org/10.5281/zenodo.3767318" httpUri="https://zenodo.org/record/3767318/files/figure.png" pageId="59" pageNumber="60">Fig. 35A</figureCitation>
|
||
). Coxite wide, smoothly rounded, laterally curving around telopodite, on posterior side visible laterally as a small, triangular sclerite (
|
||
<figureCitation box="[914,1025,1398,1425]" captionStart="Figure 35" captionStartId="63.[140,212,1558,1580]" captionTargetBox="[157,1090,165,1538]" captionTargetId="figure@63.[157,1091,165,1539]" captionTargetPageId="63" captionText="Figure 35. Pseudocentrobolus aureus sp. n., male paratype, A–C left, D–E right posterior gonopod. A overview, anterior view B detail of disc, anterior view C membranous process, anterior view D overview, posterior view E detail of disc, posterior view F detail of membranous process, posterior view. Cx = coxite; mem-p = membranous process; St = sternite; T = telopodite." figureDoi="http://doi.org/10.5281/zenodo.3767318" httpUri="https://zenodo.org/record/3767318/files/figure.png" pageId="59" pageNumber="60">Fig. 35B</figureCitation>
|
||
). Coxite anteriorly almost entirely covering telopodite (
|
||
<figureCitation box="[754,868,1434,1461]" captionStart="Figure 35" captionStartId="63.[140,212,1558,1580]" captionTargetBox="[157,1090,165,1538]" captionTargetId="figure@63.[157,1091,165,1539]" captionTargetPageId="63" captionText="Figure 35. Pseudocentrobolus aureus sp. n., male paratype, A–C left, D–E right posterior gonopod. A overview, anterior view B detail of disc, anterior view C membranous process, anterior view D overview, posterior view E detail of disc, posterior view F detail of membranous process, posterior view. Cx = coxite; mem-p = membranous process; St = sternite; T = telopodite." figureDoi="http://doi.org/10.5281/zenodo.3767318" httpUri="https://zenodo.org/record/3767318/files/figure.png" pageId="59" pageNumber="60">Fig. 35A</figureCitation>
|
||
). Telopodite with a short, well-rounded process. Process curved laterally, not or only slightly projecting above telopodite margin (
|
||
<figureCitation box="[502,613,1504,1531]" captionStart="Figure 35" captionStartId="63.[140,212,1558,1580]" captionTargetBox="[157,1090,165,1538]" captionTargetId="figure@63.[157,1091,165,1539]" captionTargetPageId="63" captionText="Figure 35. Pseudocentrobolus aureus sp. n., male paratype, A–C left, D–E right posterior gonopod. A overview, anterior view B detail of disc, anterior view C membranous process, anterior view D overview, posterior view E detail of disc, posterior view F detail of membranous process, posterior view. Cx = coxite; mem-p = membranous process; St = sternite; T = telopodite." figureDoi="http://doi.org/10.5281/zenodo.3767318" httpUri="https://zenodo.org/record/3767318/files/figure.png" pageId="59" pageNumber="60">Fig. 35B</figureCitation>
|
||
). Mesal margin elevated, laterally with a sharp ridge (
|
||
<figureCitation box="[288,396,1539,1566]" captionStart="Figure 35" captionStartId="63.[140,212,1558,1580]" captionTargetBox="[157,1090,165,1538]" captionTargetId="figure@63.[157,1091,165,1539]" captionTargetPageId="63" captionText="Figure 35. Pseudocentrobolus aureus sp. n., male paratype, A–C left, D–E right posterior gonopod. A overview, anterior view B detail of disc, anterior view C membranous process, anterior view D overview, posterior view E detail of disc, posterior view F detail of membranous process, posterior view. Cx = coxite; mem-p = membranous process; St = sternite; T = telopodite." figureDoi="http://doi.org/10.5281/zenodo.3767318" httpUri="https://zenodo.org/record/3767318/files/figure.png" pageId="59" pageNumber="60">Fig. 35B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="59.[140,1108,167,1672]" lastBlockId="61.[140,1108,765,1038]" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60">
|
||
<emphasis box="[185,463,1575,1601]" italics="true" pageId="59" pageNumber="60">Posterior gonopods in situ</emphasis>
|
||
almost completely covered by anterior gonopod. Suture separating coxite and telopodite well-visible. Sternite sclerotized and well-visible (
|
||
<figureCitation captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="59" pageNumber="60">Figs 31H</figureCitation>
|
||
, 34D). Finger-shaped process of coxite (
|
||
<figureCitation box="[644,747,1645,1671]" captionStart="Figure 34" captionStartId="61.[140,212,641,663]" captionTargetBox="[140,1108,165,621]" captionTargetId="figure@61.[140,1108,165,621]" captionTargetPageId="61" captionText="Figure 34. Pseudocentrobolus aureus sp. n., male holotype, anterior gonopod A anterior view B posterior view. Cx = coxite; St = sternite; T = telopodite. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767316" httpUri="https://zenodo.org/record/3767316/files/figure.png" pageId="59" pageNumber="60">Fig. 34D</figureCitation>
|
||
) not curved apically, but coaxial to coxite. Coxite mesally with a single groove, basally wide, protruding into a short stem towards telopodite. Telopodite as long and as wide as coxite (
|
||
<figureCitation box="[918,1027,202,228]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="60" pageNumber="61">Fig. 31G</figureCitation>
|
||
). Telopodites arranged face-to-face with one another (
|
||
<figureCitation box="[679,825,237,264]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="60" pageNumber="61">Figs 31G, H</figureCitation>
|
||
). A rounded disc present on apical part of telopodite (
|
||
<figureCitation box="[456,600,272,299]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="60" pageNumber="61">Figs 31G, H</figureCitation>
|
||
), disc covered with numerous tiny sclerotized teeth (
|
||
<figureCitation box="[214,356,765,792]" captionStart="Figure 34" captionStartId="61.[140,212,641,663]" captionTargetBox="[140,1108,165,621]" captionTargetId="figure@61.[140,1108,165,621]" captionTargetPageId="61" captionText="Figure 34. Pseudocentrobolus aureus sp. n., male holotype, anterior gonopod A anterior view B posterior view. Cx = coxite; St = sternite; T = telopodite. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767316" httpUri="https://zenodo.org/record/3767316/files/figure.png" pageId="61" pageNumber="62">Figs 34B, E</figureCitation>
|
||
), basally with a membranous process, into which sperm canal is discharging (
|
||
<figureCitation box="[284,459,800,827]" captionStart="Figure 34" captionStartId="61.[140,212,641,663]" captionTargetBox="[140,1108,165,621]" captionTargetId="figure@61.[140,1108,165,621]" captionTargetPageId="61" captionText="Figure 34. Pseudocentrobolus aureus sp. n., male holotype, anterior gonopod A anterior view B posterior view. Cx = coxite; St = sternite; T = telopodite. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767316" httpUri="https://zenodo.org/record/3767316/files/figure.png" pageId="61" pageNumber="62">Figs 34A, D, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767314" ID-Zenodo-Dep="3767314" httpUri="https://zenodo.org/record/3767314/files/figure.png" pageId="60" pageNumber="61" startId="60.[140,212,1558,1580]" targetBox="[170,1078,339,1541]" targetPageId="60">
|
||
<paragraph pageId="60" pageNumber="61">
|
||
<emphasis bold="true" box="[140,252,1558,1580]" pageId="60" pageNumber="61">Figure 33.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[258,480,1558,1580]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[258,480,1558,1580]" italics="true" pageId="60" pageNumber="61">Pseudocentrobolus aureus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[486,537,1559,1580]" pageId="60" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
|
||
A–C male, D female paratype, SEM.
|
||
<emphasis bold="true" box="[894,913,1558,1580]" pageId="60" pageNumber="61">A</emphasis>
|
||
left mandible, mesal view
|
||
<emphasis bold="true" box="[190,207,1590,1612]" pageId="60" pageNumber="61">B</emphasis>
|
||
midbody ring, overview
|
||
<emphasis bold="true" box="[445,464,1590,1612]" pageId="60" pageNumber="61">C</emphasis>
|
||
midbody ring, detail of metazona
|
||
<emphasis bold="true" box="[795,814,1590,1612]" pageId="60" pageNumber="61">D</emphasis>
|
||
right vulva, lateral view. 3iT = 3-combed internal tooth; av = anterior valve; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; re = rectangular excavations.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767316" ID-Zenodo-Dep="3767316" httpUri="https://zenodo.org/record/3767316/files/figure.png" pageId="61" pageNumber="62" startId="61.[140,212,641,663]" targetBox="[140,1108,165,621]" targetPageId="61">
|
||
<paragraph blockId="61.[140,1107,641,695]" pageId="61" pageNumber="62">
|
||
<emphasis bold="true" box="[140,250,641,663]" pageId="61" pageNumber="62">Figure 34.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[256,477,641,663]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[256,477,641,663]" italics="true" pageId="61" pageNumber="62">Pseudocentrobolus aureus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[482,533,641,662]" pageId="61" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male holotype, anterior gonopod
|
||
<emphasis bold="true" box="[852,871,641,663]" pageId="61" pageNumber="62">A</emphasis>
|
||
anterior view
|
||
<emphasis bold="true" box="[1003,1020,641,663]" pageId="61" pageNumber="62">B</emphasis>
|
||
posterior view. Cx = coxite; St = sternite; T = telopodite. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="61.[140,1108,765,1038]" pageId="61" pageNumber="62">
|
||
<emphasis box="[185,454,835,861]" italics="true" pageId="61" pageNumber="62">Female sexual characters:</emphasis>
|
||
vulva simple, bivalve-like, with a small, poorly sclerotized operculum at base (
|
||
<figureCitation box="[362,469,871,897]" captionStart="Figure 33" captionStartId="60.[140,212,1558,1580]" captionTargetBox="[170,1078,339,1541]" captionTargetId="figure@60.[170,1079,339,1541]" captionTargetPageId="60" captionText="Figure 33. Pseudocentrobolus aureus sp. n. A–C male, D female paratype, SEM. A left mandible, mesal view B midbody ring, overview C midbody ring, detail of metazona D right vulva, lateral view. 3iT = 3-combed internal tooth; av = anterior valve; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; re = rectangular excavations." figureDoi="http://doi.org/10.5281/zenodo.3767314" httpUri="https://zenodo.org/record/3767314/files/figure.png" pageId="61" pageNumber="62">Fig. 33D</figureCitation>
|
||
). Anterior valve apically overlapping posterior one, both valves smooth, lacking sensory cones. Towards opening on each valve basally with two or three rows of setae (
|
||
<figureCitation box="[394,501,941,967]" captionStart="Figure 33" captionStartId="60.[140,212,1558,1580]" captionTargetBox="[170,1078,339,1541]" captionTargetId="figure@60.[170,1079,339,1541]" captionTargetPageId="60" captionText="Figure 33. Pseudocentrobolus aureus sp. n. A–C male, D female paratype, SEM. A left mandible, mesal view B midbody ring, overview C midbody ring, detail of metazona D right vulva, lateral view. 3iT = 3-combed internal tooth; av = anterior valve; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; re = rectangular excavations." figureDoi="http://doi.org/10.5281/zenodo.3767314" httpUri="https://zenodo.org/record/3767314/files/figure.png" pageId="61" pageNumber="62">Fig. 33D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="61" pageNumber="62" type="etymology">
|
||
<paragraph blockId="61.[140,1108,765,1038]" pageId="61" pageNumber="62">
|
||
<emphasis bold="true" box="[185,315,976,1002]" inLineHeading="true" pageId="61" pageNumber="62" reason="1">Etymology</emphasis>
|
||
:
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[331,524,976,1002]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[331,524,976,1002]" italics="true" pageId="61" pageNumber="62">Pseudocentrobolus</emphasis>
|
||
</taxonomicName>
|
||
, masculine, refers to the superficial resemblance of species of this genus to some species of the South African genus
|
||
<taxonomicName authorityName="Cook" authorityYear="1897" box="[864,993,1011,1037]" class="Diplopoda" family="Pachybolidae" genus="Centrobolus" kingdom="Animalia" order="Spirobolida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[864,993,1011,1037]" italics="true" pageId="61" pageNumber="62">Centrobolus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |