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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.707.13042" ID-GBIF-Dataset="77aa16c7-25ee-4130-a594-57ed6e914df1" ID-PMC="PMC5674146" ID-Pensoft-Pub="1313-2970-707-1" ID-PubMed="29118626" ID-ZBK="6A0BCD992749450C874B440D0C0A232C" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-707-1" ModsDocOrigin="ZooKeys 707" ModsDocTitle="Abyssal fauna of the UK-1 polymetallic nodule exploration area, Clarion-Clipperton Zone, central Pacific Ocean: Mollusca" checkinTime="1507668034441" checkinUser="pensoft" docAuthor="Wiklund, Helena, Taylor, John D., Dahlgren, Thomas G., Todt, Christiane, Ikebe, Chiho, Rabone, Muriel &amp; Glover, Adrian G." docDate="2017" docId="12D3DC37B39B5D35CC81BC44CBD55D57" docLanguage="en" docName="ZooKeys 707: 1-46" docOrigin="ZooKeys 707" docSource="http://dx.doi.org/10.3897/zookeys.707.13042" docTitle="Bathyspinula calcar Dall 1908" docType="treatment" docVersion="5" lastPageNumber="8" masterDocId="FFC7FFE8482EB03042654660FFF4EB7D" masterDocTitle="Abyssal fauna of the UK- 1 polymetallic nodule exploration area, Clarion-Clipperton Zone, central Pacific Ocean: Mollusca" masterLastPageNumber="46" masterPageNumber="1" pageNumber="7" updateTime="1668164910568" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Abyssal fauna of the UK- 1 polymetallic nodule exploration area, Clarion-Clipperton Zone, central Pacific Ocean: Mollusca</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wiklund, Helena</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Taylor, John D.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dahlgren, Thomas G.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Todt, Christiane</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ikebe, Chiho</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rabone, Muriel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Glover, Adrian G.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>707</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>46</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.707.13042</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.707.13042</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-707-1</mods:identifier>
<mods:identifier type="ZBK">6A0BCD992749450C874B440D0C0A232C</mods:identifier>
<mods:identifier type="ZooBank">6A0BCD992749450C874B440D0C0A232C</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="135490002" LSID="urn:lsid:plazi:treatment:12D3DC37B39B5D35CC81BC44CBD55D57" httpUri="http://treatment.plazi.org/id/12D3DC37B39B5D35CC81BC44CBD55D57" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection pageId="6" pageNumber="7" type="nomenclature">
<paragraph pageId="6" pageNumber="7">
<taxonomicName authority="Dall, 1908" authorityName="Dall" authorityYear="1908" class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Pholadomyoida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar (Dall, 1908)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph pageId="6" pageNumber="7">Material examined.</paragraph>
<paragraph pageId="6" pageNumber="7">NHM_146 NHMUK 20170044, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/61f15e3c-f070-48a1-b484-780b37f7feb6</paragraph>
<paragraph pageId="7" pageNumber="8">
<pageBreakToken pageId="7" pageNumber="8" start="start">NHM</pageBreakToken>
_149A NHMUK 20170045, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/c44da298-9b61-4d6d-a1cd-2d6c3bd70859
</paragraph>
<paragraph pageId="7" pageNumber="8">NHM_181 NHMUK 20170043, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/3ab74908-1a5d-465f-890c-49373a44906c</paragraph>
<paragraph pageId="7" pageNumber="8">NHM_284 NHMUK 20170046, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/ad2cb87b-1fce-415d-ab45-1619bbc4352b</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="description">
<paragraph pageId="7" pageNumber="8">Description.</paragraph>
<paragraph pageId="7" pageNumber="8">Shell sub-ovate, laterally compressed, with long, sharply pointed posterior rostrum. Periostracum shiny, medium brown. Posterior rostrum shorter, less defined in juveniles. Voucher specimen NHM_181 shell length 13.5 mm, width 7.6 mm (Fig. 6A).</paragraph>
<caption pageId="7" pageNumber="8">
<paragraph pageId="7" pageNumber="8">
Figure 6.
<taxonomicName class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar</taxonomicName>
(Dall, 1908) A Specimen NHM_181, Image of live specimen after recovery, length 13.5 mm
<normalizedToken originalValue="BD">B-D</normalizedToken>
Specimen NHM_149A confirmed juvenile
<taxonomicName lsidName="B. calcar" pageId="7" pageNumber="8" rank="species" species="calcar">B. calcar</taxonomicName>
using DNA evidence, total length of animal ~2mm. Scale bars: 5 mm (A); 1 mm (
<normalizedToken originalValue="BD">B-D</normalizedToken>
). Image attribution Glover, Taylor, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="genetic data">
<paragraph pageId="7" pageNumber="8">Genetic data.</paragraph>
<paragraph pageId="7" pageNumber="8">GenBank NHM_146 18S-157475, COI-MF157505; NHM_149A COI-MF157506; NHM_181 18S-MF157479, COI-MF157507; NHM_284 COI-MF157514.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="remarks">
<paragraph pageId="7" pageNumber="8">Remarks.</paragraph>
<paragraph pageId="7" pageNumber="8">
Widely distributed in the eastern Pacific at depths of 400-5000 m (see
<bibRefCitation author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott 2012</bibRefCitation>
). The holotype (USNM 110573) was collected 725 km west of Trujillo, Peru at 2370 fathoms (4334 m). Forms a unique monophyletic clade distinct from all other AB01 specimens. Genetic match in 18S to
<taxonomicName class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar</taxonomicName>
(GenBank KC993875) from the north eastern Pacific (
<bibRefCitation author="Sharma, PP" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="31" pageNumber="32" pagination="188 - 204" title="Into the deep: A phylogenetic approach to the bivalve subclass Protobranchia." url="https://doi.org/10.1016/j.ympev.2013.05.018" volume="69" year="2013">Sharma et al. 2013</bibRefCitation>
), but as the GenBank 18S sequence from
<taxonomicName lsidName="B. calcar" pageId="7" pageNumber="8" rank="species" species="calcar">B. calcar</taxonomicName>
was only 289 bp long and as that specimen lacked COI, it was not included in the analyses. Some very small juvenile specimens (Fig. 6
<normalizedToken originalValue="BD">B-D</normalizedToken>
) were recovered that superficially resemble
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella knudseni" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="knudseni">Ledella knudseni</taxonomicName>
sp. n. (Fig. 7) and may be easily confused. Genetic data confirmed these to be
<taxonomicName class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar</taxonomicName>
(Fig. 12). These may be distinguised from
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">Ledella</taxonomicName>
by the shiny and iridescent nature of the shell surface of
<taxonomicName lsidName="B. calcar" pageId="7" pageNumber="8" rank="species" species="calcar">B. calcar</taxonomicName>
, which is preserved in the juveniles.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="ecology">
<paragraph pageId="7" pageNumber="8">Ecology.</paragraph>
<paragraph pageId="7" pageNumber="8">Relatively large bivalve recovered from epibenthic sledge tow in polymetallic nodule province.</paragraph>
</subSubSection>
</treatment>
</document>