256 lines
21 KiB
XML
256 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.110.25846" ID-GBIF-Dataset="44b94249-6f5a-4a6c-8039-c441517a1144" ID-PMC="PMC6224366" ID-Pensoft-Pub="1314-2003-110-23" ID-Pensoft-UUID="FFD8FFA5FFABFC45FF8EFE1DD80CFF9C" ID-PubMed="30416344" ID-Zenodo-Dep="1481316" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-2003-110-23" ModsDocOrigin="PhytoKeys " ModsDocTitle="Begoniayenyeniae (Begoniaceae), a new species from Endau Rompin National Park, Johor, Malaysia" checkinTime="1553125356695" checkinUser="pensoft" docAuthor="Tan, Joanne Pei-Chih, Tam, Sheh May & Kiew, Ruth" docDate="2018" docId="121134B5711D5CA51B54E60AC7FD543D" docLanguage="en" docName="PhytoKeys 110: 23-37" docOrigin="PhytoKeys 110" docSource="http://dx.doi.org/10.3897/phytokeys.110.25846" docTitle="Begonia yenyeniae J. P. C. Tan 2018, sp. nov." docType="treatment" docVersion="6" id="FFD8FFA5FFABFC45FF8EFE1DD80CFF9C" lastPageNumber="23" masterDocId="FFD8FFA5FFABFC45FF8EFE1DD80CFF9C" masterDocTitle="Begonia yenyeniae (Begoniaceae), a new species from Endau Rompin National Park, Johor, Malaysia" masterLastPageNumber="37" masterPageNumber="23" pageNumber="23" updateTime="1668139057705" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Begonia yenyeniae (Begoniaceae), a new species from Endau Rompin National Park, Johor, Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tan, Joanne Pei-Chih</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tam, Sheh May</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kiew, Ruth</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>110</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>23</mods:start>
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<mods:end>37</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.110.25846</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.110.25846</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-110-23</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFD8FFA5FFABFC45FF8EFE1DD80CFF9C</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1481316</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="154472609" LSID="urn:lsid:plazi:treatment:121134B5711D5CA51B54E60AC7FD543D" httpUri="http://treatment.plazi.org/id/121134B5711D5CA51B54E60AC7FD543D" lastPageNumber="23" pageId="0" pageNumber="23">
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<subSubSection pageId="0" pageNumber="23" type="nomenclature">
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<paragraph pageId="0" pageNumber="23">
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<taxonomicName LSID="450A219E-0351-5E80-A25A-633315960329" authority="J. P. C. Tan" authorityName="J. P. C. Tan" authorityYear="2018" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia yenyeniae" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="yenyeniae" status="sp. nov.">Begonia yenyeniae J.P.C.Tan</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="23">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Begonia yenyeniae J. P. C. Tan, sp. nov. A Side and front view female flower B Back and front view of male flower C Stamen mass D Anthers E Habitat: moss-covered rocky slope by waterfall F Young fruit with stigma still attached G Transverse section of fruit H Seeds I Mature leaf J Upper leaf surface (moderately bullate) K Veins completely prominent on lower leaf surface L Petiole M Young blade N Stipules O-Q Upper, lower and side view of leaf margin R A pair of bracts and bracteole at peduncle and rachis; hairs scarcely on ventral surface of outer tepals. (Photographs by E Y. Y. Sam, D P. T. Ong)" figureDoi="10.3897/phytokeys.110.25846.figure2" httpUri="https://binary.pensoft.net/fig/239606" pageId="0" pageNumber="23">Figure 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="23" type="section">
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<paragraph pageId="0" pageNumber="23">Section.</paragraph>
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<paragraph pageId="0" pageNumber="23">
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<taxonomicName class="Magnoliopsida" family="Begoniaceae" genus="Jackia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Jackia" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="23">Jackia</emphasis>
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</taxonomicName>
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M.Hughes
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="23" type="diagnosis">
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<paragraph pageId="0" pageNumber="23">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="23">
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Similar to
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<taxonomicName class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia rajah" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="rajah">
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<emphasis italics="true" pageId="0" pageNumber="23">Begonia rajah</emphasis>
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</taxonomicName>
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Ridl. (1894:213) in its handsome leaves, striking brownish-pink to brownish-red with greenish-yellow veins in young leaves becoming bronzy variegation at maturity; in its creeping growth habit, number of tepals in male (4 tepals) and female flowers (3), palmate leaf venation and many-flowered cymes, but several notable characters distinguish the new species, including its orbicular-reniform leaf blades (vs. subrotund with an abruptly acute apex in
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<taxonomicName class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. rajah" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="rajah">
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<emphasis italics="true" pageId="0" pageNumber="23">B. rajah</emphasis>
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</taxonomicName>
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), smaller stipules, 9-12
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<normalizedToken originalValue="×">x</normalizedToken>
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3-4 mm, three times longer than wide (vs. 15-20
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<normalizedToken originalValue="×">x</normalizedToken>
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10-12 mm, less than twice as long as wide), smaller ovate or obovate bracts 2-3
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<normalizedToken originalValue="×">x</normalizedToken>
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1.5-2 mm (vs. bracts bowl-shaped, wide ovate, 5-8
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<normalizedToken originalValue="×">x</normalizedToken>
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7.5-8 mm), margin shallowly crenate (vs. margin angular), bullate leaf surface (vs. conspicuously pronounced bullate). Furthermore, the leaf colour of cultivated
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<taxonomicName class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. rajah" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="rajah">
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<emphasis italics="true" pageId="0" pageNumber="23">B. rajah</emphasis>
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</taxonomicName>
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is more vivid and, in contrast, its tepals, stipules and bracts are also a deeper shade of pink, compared to those of
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<taxonomicName class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. yenyeniae" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="yenyeniae">
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<emphasis italics="true" pageId="0" pageNumber="23">B. yenyeniae</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="23">
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It is also similar to
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<taxonomicName class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. reginula" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="reginula">
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<emphasis italics="true" pageId="0" pageNumber="23">B. reginula</emphasis>
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</taxonomicName>
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<bibRefCitation author="Kiew, R" journalOrPublisher="Nature Malaysiana" pageId="0" pageNumber="23" refId="B10" refString="Kiew, R, 2005. Begonias of Peninsular Malaysia. Natural History Publications (Borneo) Sdn. Bhd., Kota Kinabalu, Malaysia, 1-308." title="Begonias of Peninsular Malaysia. Natural History Publications (Borneo) Sdn. Bhd., Kota Kinabalu, Malaysia, 1 - 308." year="2005">Kiew (2005</bibRefCitation>
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: 218) in its habit, leaf colour, less pronounced bullate blade, palmate venation, bracts, ovary with 3 equal wings of similar shape, but
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<taxonomicName class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. yenyeniae" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="yenyeniae">
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<emphasis italics="true" pageId="0" pageNumber="23">B. yenyeniae</emphasis>
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</taxonomicName>
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is different in its relatively smaller and narrower tepals 5-7
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<normalizedToken originalValue="×">x</normalizedToken>
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6 mm (vs. 6-10
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<normalizedToken originalValue="×">x</normalizedToken>
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9-11 mm in
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<taxonomicName class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. reginula" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="reginula">
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<emphasis italics="true" pageId="0" pageNumber="23">B. reginula</emphasis>
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</taxonomicName>
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), 3 in male or 4 tepals in female flowers (vs. 2 tepals in both male and female flowers) and rounded base (vs. subcordate), stipules with prominently keeled (vs. keel absent), and apex rounded (vs. apex attenuate).
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</paragraph>
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<paragraph pageId="0" pageNumber="23">
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Although molecular data indicate a close relationship to
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<taxonomicName class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. foxworthyi" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="foxworthyi">
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<emphasis italics="true" pageId="0" pageNumber="23">B. foxworthyi</emphasis>
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</taxonomicName>
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((1925: 311), the latter species is morphologically distinct in its conspicuously oblique leaf with an acute to acuminate apex (vs. orbicular-reniform with a rounded apex), its entire margin (vs. crenate) and plain green, non-bullate leaf surface (vs. purplish-green to brownish-purple and bullate) and its male flowers with 2 tepals (vs. 4 tepals). In addition, it usually grows on limestone substrates (vs. confined to granite substrates.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="23" type="materials_examined">
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<paragraph pageId="0" pageNumber="23">Type.</paragraph>
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<paragraph pageId="0" pageNumber="23">Peninsular Malaysia. Johor, Mersing, Endau-Rompin National Park, Sungai Selai, 14 April 2018, Kiew FRI 81950 (holotype E!; iso-: K!, SAR!, SING!).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="23" type="description">
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<paragraph pageId="0" pageNumber="23">Description.</paragraph>
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<paragraph pageId="0" pageNumber="23">
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Lithophytic herb with rhizomatous stem. Indumentum of soft hairs, usually cream- coloured, sparse on stipules (mostly on the keel and 0.5-1.5 mm long), petioles more densely hairy towards blade (2-3 mm), leaf margin (hairs ca. 1 mm) and veins beneath (ca. 1 mm) but sparse (ca. 0.5 mm) or absent on ventral surface of outer tepals and on peduncle hairs hardly visible to the naked eye.
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<emphasis bold="true" pageId="0" pageNumber="23">Stems</emphasis>
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creeping, apex usually slightly erect, branched, 5-9 mm thick, light yellowish-green; stipules 3 per node, two triangular, abruptly narrowed to an attenuate apex terminating in a hair, keeled from base to apex, margin entire and translucent, 9-12
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<normalizedToken originalValue="×">x</normalizedToken>
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3-4 mm, pale yellowish-green, one narrowly lanceolate, ca. 3 mm long, persistent.
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<emphasis bold="true" pageId="0" pageNumber="23">Leaves</emphasis>
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tufted, alternate, 2-3 mm apart; petioles terete, 2.5-3.5 mm across, up to 12.5 cm long, pale brownish-pink to darker pink towards blade; blades thinly succulent, glabrous, orbicular-reniform, asymmetric, 8-11.5
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<normalizedToken originalValue="×">x</normalizedToken>
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9.5-13.5 cm, scarcely angular, margin somewhat crenate with acute teeth bent abruptly downwards between teeth, ciliate, basal lobes cordate, overlapping when mature, moderately raised between veins, light purplish-green to dull brownish-purple, young blades brownish-pink to brownish-red, paler beneath; veins palmate, slightly prominent towards the base but impressed where branched towards the margin, prominent beneath, lateral veins ca. 2-3 pairs, greenish-yellow when young and whitish-green when mature.
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<emphasis bold="true" pageId="0" pageNumber="23">Inflorescences</emphasis>
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axillary, more or less erect, 10-23 cm long, brownish-red, peduncles 8-18.5 cm long, two main branches 2-3 cm long, pedicels 6-9 mm; bracts in pairs at node of peduncle, glabrous, elliptic-ovate or obovate, margin towards apex laciniate, 2-3
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<normalizedToken originalValue="×">x</normalizedToken>
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1.5-2 mm, light yellowish-green sometimes with a faint tinge of pink, persistent.
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<emphasis bold="true" pageId="0" pageNumber="23">Male flowers</emphasis>
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with 4 tepals, margin entire, 1-1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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1.1-1.4 cm; outer 2 tepals rotund, concave at centre, 5-7
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<normalizedToken originalValue="×">x</normalizedToken>
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6 mm, pale greenish-white or light pink or pale pinkish-white, inner 2 tepals obovate, apex rounded or sometimes retuse and impressed along centre, ca. 5
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<normalizedToken originalValue="×">x</normalizedToken>
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3 mm, white; stamens numerous, torus ca. 1 mm long, stamen mass globose, symmetric, ca. 2.5 mm across; anthers obovoid-oblong, tip emarginate, ca. 0.5 mm long, dehiscing through 2 longitudinal slits.
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<emphasis bold="true" pageId="0" pageNumber="23">Female flowers</emphasis>
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with a light yellowish-green ovary sometimes with faint light pink tinge, ca. 5.5
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<normalizedToken originalValue="×">x</normalizedToken>
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9 mm, locules 3, wings 3 equal, placentation axile, 1 placenta per locule, each placenta usually with 2 minute branches at the base; usually tepals 3, outer 2 tepals, rotund, concave at centre, ca. 4
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<normalizedToken originalValue="×">x</normalizedToken>
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4.5 mm, white with faint green tinge; styles and stigmas 3, 1.5-2 mm, style light greenish-yellow, stigma pale yellow, papillose, spiral band.
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<emphasis bold="true" pageId="0" pageNumber="23">Capsules</emphasis>
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ca. 6
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<normalizedToken originalValue="×">x</normalizedToken>
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12 mm, locules 3, splitting longitudinally between locules, wing 3 equal, ca. 4 mm wide, surface glabrous, styles and stigmas persisting after tepals have fallen, dangling on a fine, thread-like pedicel ca. 5.5 mm long.
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<emphasis bold="true" pageId="0" pageNumber="23">Seeds</emphasis>
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numerous, barrel-shaped, 0.3-0.34
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<normalizedToken originalValue="×">x</normalizedToken>
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0.2-0.22 mm, collar cells slightly more than half the seed length, surface sculptured.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="23" type="distribution">
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<paragraph pageId="0" pageNumber="23">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="23">Endemic in Peninsular Malaysia, Johor, Mersing District, Endau-Rompin National Park, Sungai Selai. It is apparently a rare species as it is known only from the type locality in Endau Rompin National Park.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="23" type="etymology">
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<paragraph pageId="0" pageNumber="23">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="23">
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Named after Dr Sam Yen-Yen, Malaysian botanist, specialist in
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<taxonomicName class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="family">Zingiberaceae</taxonomicName>
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who first discovered the species and recognised its potential as an ornamental plant.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="23" type="conservation status">
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<paragraph pageId="0" pageNumber="23">Conservation status.</paragraph>
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<paragraph pageId="0" pageNumber="23">Critically Endangered CR B2ab(iii, v), D1. Although the type locality is within a Totally Protected Area, it is known from only one population about 1.5 km2 and its habitat is threatened by ecotourism activity and illegal collecting. The area of observed population covers about 1.5 km2 (P.T. Ong pers. comm.).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="23" type="ecology">
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<paragraph pageId="0" pageNumber="23">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="23">In primary lowland mixed dipterocarp forest, growing on moss-covered rocks, rarely epiphytic, near a waterfall in deep shade.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="23" type="notes">
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<paragraph pageId="0" pageNumber="23">Note.</paragraph>
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<paragraph pageId="0" pageNumber="23">
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The ovary of species in section
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<taxonomicName class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="section" section="Jackia">Jackia</taxonomicName>
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have three locules, each with an unbranched placenta, but in this species two vestigial branches are present near the base of the placenta (Figure
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Begonia yenyeniae J. P. C. Tan, sp. nov. A Side and front view female flower B Back and front view of male flower C Stamen mass D Anthers E Habitat: moss-covered rocky slope by waterfall F Young fruit with stigma still attached G Transverse section of fruit H Seeds I Mature leaf J Upper leaf surface (moderately bullate) K Veins completely prominent on lower leaf surface L Petiole M Young blade N Stipules O-Q Upper, lower and side view of leaf margin R A pair of bracts and bracteole at peduncle and rachis; hairs scarcely on ventral surface of outer tepals. (Photographs by E Y. Y. Sam, D P. T. Ong)" figureDoi="10.3897/phytokeys.110.25846.figure2" httpUri="https://binary.pensoft.net/fig/239606" pageId="0" pageNumber="23">2G</figureCitation>
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). This is also seen, but is less pronounced, in
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<taxonomicName class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. rajah" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="rajah">
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<emphasis italics="true" pageId="0" pageNumber="23">B. rajah</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<caption ID-Zenodo-Dep="1481326" doi="10.3897/phytokeys.110.25846.figure2" httpUri="https://binary.pensoft.net/fig/239606" pageId="0" pageNumber="23" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="23">
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<emphasis bold="true" pageId="0" pageNumber="23">Figure 2.</emphasis>
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<taxonomicName authorityName="J. P. C. Tan" authorityYear="2018" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia yenyeniae" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="yenyeniae">
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<emphasis italics="true" pageId="0" pageNumber="23">Begonia yenyeniae</emphasis>
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</taxonomicName>
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J.P.C.Tan, sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="23">A</emphasis>
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Side and front view female flower
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<emphasis bold="true" pageId="0" pageNumber="23">B</emphasis>
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Back and front view of male flower
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<emphasis bold="true" pageId="0" pageNumber="23">C</emphasis>
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Stamen mass
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<emphasis bold="true" pageId="0" pageNumber="23">D</emphasis>
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Anthers
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<emphasis bold="true" pageId="0" pageNumber="23">E</emphasis>
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Habitat: moss-covered rocky slope by waterfall
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<emphasis bold="true" pageId="0" pageNumber="23">F</emphasis>
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Young fruit with stigma still attached
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<emphasis bold="true" pageId="0" pageNumber="23">G</emphasis>
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Transverse section of fruit
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<emphasis bold="true" pageId="0" pageNumber="23">H</emphasis>
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Seeds
|
||
<emphasis bold="true" pageId="0" pageNumber="23">I</emphasis>
|
||
Mature leaf
|
||
<emphasis bold="true" pageId="0" pageNumber="23">J</emphasis>
|
||
Upper leaf surface (moderately bullate)
|
||
<emphasis bold="true" pageId="0" pageNumber="23">K</emphasis>
|
||
Veins completely prominent on lower leaf surface
|
||
<emphasis bold="true" pageId="0" pageNumber="23">L</emphasis>
|
||
Petiole
|
||
<emphasis bold="true" pageId="0" pageNumber="23">M</emphasis>
|
||
Young blade
|
||
<emphasis bold="true" pageId="0" pageNumber="23">N</emphasis>
|
||
Stipules
|
||
<emphasis bold="true" pageId="0" pageNumber="23">
|
||
<normalizedToken originalValue="O–Q">O-Q</normalizedToken>
|
||
</emphasis>
|
||
Upper, lower and side view of leaf margin
|
||
<emphasis bold="true" pageId="0" pageNumber="23">R</emphasis>
|
||
A pair of bracts and bracteole at peduncle and rachis; hairs scarcely on ventral surface of outer tepals. (Photographs by
|
||
<emphasis bold="true" pageId="0" pageNumber="23">E</emphasis>
|
||
Y.Y. Sam,
|
||
<emphasis bold="true" pageId="0" pageNumber="23">D</emphasis>
|
||
P.T. Ong)
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="23" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="23">Other specimens examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="23">
|
||
<emphasis bold="true" pageId="0" pageNumber="23">Peninsular Malaysia.</emphasis>
|
||
Johor: Mersing, Endau-Rompin National Park, Sungai Selai, 15 Aug 2002, Sam et al. FRI 47082 (KEP!), 24 July 2012, Hairul et al. FRI 78570 (KEP!, SING!).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |