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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.122.33032" ID-GBIF-Dataset="f497907c-ee54-41cd-b666-7292c766b9c3" ID-PMC="PMC6548746" ID-Pensoft-Pub="1314-2003-122-29" ID-Pensoft-UUID="FFB7FFA5CF25AD00FFBDFFEBFFFCFFDD" ID-PubMed="31182907" ID-Zenodo-Dep="3238737" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2003-122-29" ModsDocOrigin="PhytoKeys 122" ModsDocTitle="Páramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in páramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia" checkinTime="1559110536903" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Soreng, Robert J., Bravo-Pedraza, William J., Cuta-Alarcon, Lia E., Giraldo-Canas, Diego, Aguilar-Cano, Jose &amp; Peterson, Paul M." docDate="2019" docId="0E5F140C488BC1F779F990B08B33C56E" docLanguage="en" docName="PhytoKeys 122: 29-78" docOrigin="PhytoKeys 122" docSource="http://dx.doi.org/10.3897/phytokeys.122.33032" docTitle="Calamagrostis rigida Trin. ex Steud., Nomencl. Bot. (ed. 2) 1: 251. 1840. Deyeuxia rigida Kunth, Nov. Gen. Sp. (quarto ed.) 1: 144. 1815 [1816]. Arundo rigida (Kunth) Poir., Encycl. 4: 705. 1816" docType="treatment" docVersion="6" id="FFB7FFA5CF25AD00FFBDFFEBFFFCFFDD" lastPageNumber="29" masterDocId="FFB7FFA5CF25AD00FFBDFFEBFFFCFFDD" masterDocTitle="Paramo Calamagrostis s. l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia" masterLastPageNumber="78" masterPageNumber="29" pageNumber="29" updateTime="1668139302904" updateUser="ExternalLinkService">
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<mods:title>Paramo Calamagrostis s. l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sylvester, Steven P.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Soreng, Robert J.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bravo-Pedraza, William J.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cuta-Alarcon, Lia E.</mods:namePart>
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<mods:namePart>Giraldo-Canas, Diego</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Aguilar-Cano, Jose</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Peterson, Paul M.</mods:namePart>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2019</mods:date>
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<mods:number>122</mods:number>
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<treatment ID-GBIF-Taxon="157294006" LSID="urn:lsid:plazi:treatment:0E5F140C488BC1F779F990B08B33C56E" httpUri="http://treatment.plazi.org/id/0E5F140C488BC1F779F990B08B33C56E" lastPageNumber="29" pageId="0" pageNumber="29">
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<paragraph pageId="0" pageNumber="29">
<taxonomicName LSID="9604CAD0-1055-5518-A429-E70A2F6B78EF" authority="(Kunth) Trin. ex Steud., Nomencl. Bot. (ed. 2) 1: 251. 1840. Deyeuxia rigida Kunth, Nov. Gen. Sp. (quarto ed.) 1: 144. 1815 [1816]. Arundo rigida (Kunth) Poir., Encycl. 4: 705. 1816." class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis rigida" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="rigida">Calamagrostis rigida (Kunth) Trin. ex Steud., Nomencl. Bot. (ed. 2) 1: 251. 1840. Deyeuxia rigida Kunth, Nov. Gen. Sp. (quarto ed.) 1: 144. 1815[1816]. Arundo rigida (Kunth) Poir., Encycl. 4: 705. 1816.</taxonomicName>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A general habit and a spikelet of Calamagrostis guamanensis Escalona, J. R. I. Wood 4064 (FMB), Calamagrostis heterophylla (Wedd.) Pilg., S. P. Sylvester 3158 (US), Calamagrostis pisinna Swallen, S. P. Sylvester 3107 (US), and Calamagrostis rigida (Kunth) Trin. ex Steud., S. P. Sylvester 3125 (US). Habit scale bar 5 cm; spikelet scale bar 1 mm." figureDoi="10.3897/phytokeys.122.33032.figure5" httpUri="https://binary.pensoft.net/fig/305286" pageId="0" pageNumber="29">Fig. 5 G, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29" type="reference_group">
<paragraph pageId="0" pageNumber="29">
<emphasis bold="true" pageId="0" pageNumber="29">Type.</emphasis>
ECUADOR. In planitie frigida Antisanae [inter speluncam, Machay de Antisana], et Chussulongo, 2200 hexap. (4023 m alt.)], Regni Quitensi [Pichincha], 1833, A.J.A. Bonpland 2271 [s.n.] (holotype: P (P00729827 [image!]); isotypes: BM (BM000938560 [image!]), GH (GH00023418 [image!]) fragm., LE (LE-TRIN-1804.01!) fragm. ex Herb. Humb., P (P00129585 [image!], P00740362 [image!], P026296 [image!]), US (US00406356!) fragm., W???).
</paragraph>
<paragraph pageId="0" pageNumber="29">
=
<taxonomicName class="Liliopsida" family="Poaceae" genus="Deyeuxia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deyeuxia gracilis" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="gracilis">Deyeuxia gracilis</taxonomicName>
Wedd., Bull. Soc. Bot. Fr. 22: 179. 1875.
<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis gracilis" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="gracilis">Calamagrostis gracilis</taxonomicName>
(Wedd.) Pilg., Bot. Jahrb. Syst. 42: 71. 1908.
<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis gracilis" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="gracilis">Calamagrostis gracilis</taxonomicName>
(Wedd.) Henrard, Mede. Rijks-Herb. 40: 61. 1921, nom. illeg. hom. Type: BOLIVIA. Province de Larecaja, Cordillera de Sorata, 1851 [1857], Weddell s.n. (holotype: P (P00729786 [image!]); isotypes: P (P030111 [image!]), S (S-R-1458 [image!]), US (US00479086!, US00479087!)).
</paragraph>
<paragraph pageId="0" pageNumber="29">
=
<taxonomicName class="Liliopsida" family="Poaceae" genus="Deyeuxia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deyeuxia sulcata" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="sulcata">Deyeuxia sulcata</taxonomicName>
Wedd., Bull. Soc. Bot. Fr. 22: 178-180. 1875. Type: BOLIVIA. Prov. Larecaja, viciniis Sorata [Andes de Sorata], Puerta del Inca, prope trincheras Chiliata, in Scopulosos, Reg. Alp., 3800 m alt., Mar. [Apr.] 1858 [1868], G. Mandon 1308 bis. (holotype P; isotypes: GH (GH00023431 [image!]), JE (JE00014185 [image!]) fragm., L (L0044085 [image!], L0044083 [image!]), NY (NY380547 [image!]), P (P030105 [image!]), S (S-R-1469 [image!]), US (US00149252!) fragm.).
</paragraph>
<paragraph pageId="0" pageNumber="29">
=
<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis antoniana" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="antoniana">Calamagrostis antoniana</taxonomicName>
Steud. ex Lechler, Berberid. Amer. Austr. 56. 1857, nom. nud.
</paragraph>
<paragraph pageId="0" pageNumber="29">
=
<taxonomicName authorityName="Griseb" authorityYear="1879" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis antoniana" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="antoniana">Agrostis antoniana</taxonomicName>
Griseb., Abh.
<normalizedToken originalValue="Königl">Koenigl</normalizedToken>
. Ges. Wiss.
<normalizedToken originalValue="Göttingen">Goettingen</normalizedToken>
24: 293. 1879.
<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis antoniana" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="antoniana">Calamagrostis antoniana</taxonomicName>
(Griseb.) Hack. ex
<normalizedToken originalValue="Dusén">Dusen</normalizedToken>
, Rep. Princeton Univ. Exp. Patagonia, Botany, Suppl. 8: 42. 1915.
<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis antoniana" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="antoniana">Calamagrostis antoniana</taxonomicName>
(Griseb.) Steud. ex Hitchc., Contr. U.S. Natl. Herb. 24(8): 378. 1927.
<taxonomicName authorityName="Parodi" authorityYear="1953" baseAuthorityName="Griseb." class="Liliopsida" family="Poaceae" genus="Deyeuxia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deyeuxia antoniana" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="antoniana">Deyeuxia antoniana</taxonomicName>
(Griseb.) Parodi, Revista Argent. Agron. 20: 14. 1953.
<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis antoniana" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="antoniana">Calamagrostis antoniana</taxonomicName>
(Griseb.) D.M. Moore, Fl. Tierra del Fuego 310. 1983, comb. illeg. hom. Type: ARGENTINA: Salta: Umgebung des Nevado del Castillo [Alrededores del Nevado del Castillo], 10-13000 ft [3048-3962 m alt.], 19-23 Mar. 1873, P.G. Lorentz &amp; G.H.E.W. Hieronymus 67 + 72 (lectotype, designated here: GOET (GOET006107 [image!]); isolectotypes: GOET (GOET006106 [image!], GOET006108 [image!]); P.G. Lorentz &amp; G.H.E.W. Hieronymus 67 isolectotypes: BAA (BAA00001327 [image!]) fragm., CORD (CORD00004685 [image!], CORD00004686 [image!]); P.G. Lorentz &amp; G.H.E.W. Hieronymus 72 isolectotypes: BAA (BAA00001326 [image!]) fragm., CORD (CORD00004687 [image!], CORD00004688 [image!], CORD00004689 [image!]), K [cited by
<bibRefCitation author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae Volumen 3 Tomo 2. IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="29" pagination="180 - 218" refId="B33" refString="Rugolo de Agrasar, ZE, 2012. Deyeuxia Clarion ex P. Beauv. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae Volumen 3 Tomo 2. IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 180 - 218" title="Deyeuxia Clarion ex P. Beauv." volumeTitle="Flora Argentina." year="2012">
<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
de Agrasar 2012
</bibRefCitation>
: 208, but not seen], US (US00406312!, US00406404!)). Other original material: BOLIVIA [PERU]. Viciniis Sorata, prope Milipaya, in scopulosis, reg. alpina, 3700 m alt., Mar.-May 1861, G. Mandon 1308 (GOET (GOET006230 [image!])). PERU. In graminosis pr. San Antonio rara, Jun. [1854], [W. Lechler s.n.], W. Lechler Pl. Peruv. 1800 (BAA (BAA00001648 [image!]) fragm., BR (BR0000006865757 [image!]), G (G00099542 [image!], G00099543 [image!]), GOET (GOET006109 [image!]), K (K000308438 [image!]), LE (LE00009360 [image!], LE00009361 [image!]), M?, P (P00729795 [image!], P00729844 [image!], P00740472 [image!]), S (S-R-7637 [image!]), TUB (TUB009259 [image!]), US (US00131135!), W (W1889-0241764 [image!])).
</paragraph>
<paragraph pageId="0" pageNumber="29">
=
<taxonomicName authorityName="Pilg" authorityYear="1908" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis sandiensis" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="sandiensis">Calamagrostis sandiensis</taxonomicName>
Pilg., Bot. Jahrb. Syst. 42: 68. 1908. Type: PERU. Prope Cuyocuyo, provincia Sandia, 3700-3800 m alt., 3 May 1902, Weberbauer 906 (lectotype, designated by
<bibRefCitation author="Vega, AS" journalOrPublisher="Botanica" pageId="0" pageNumber="29" pagination="31 - 35" refId="B44" refString="Vega, AS, Rugolo de Agrasar, ZE, 2013. Lectotypifications in taxa of the genera Calamagrostis, Deyeuxia, and Digitaria (Poaceae). Gayana. Botanica 70 (1): 31 - 35" title="Lectotypifications in taxa of the genera Calamagrostis, Deyeuxia, and Digitaria (Poaceae). Gayana." volume="70" year="2013">
Vega and
<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
de Agrasar (2013
</bibRefCitation>
: 28): MOL; isolectotypes: BAA (BAA00000768 [image!] fragm. ex B, S (S-R-834 [image!]), US (US00149279!) fragm.).
</paragraph>
<paragraph pageId="0" pageNumber="29">
=
<taxonomicName authorityName="Pilg" authorityYear="1925" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis gusindei" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="gusindei">Calamagrostis gusindei</taxonomicName>
Pilg. ex Skottsberg, Acta Horti Gothob. 2: 29. 1926.
<taxonomicName authorityName="Parodi" authorityYear="1953" baseAuthorityName="Pilg." class="Liliopsida" family="Poaceae" genus="Deyeuxia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deyeuxia gusindei" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="gusindei">Deyeuxia gusindei</taxonomicName>
(Pilg ex Skottsb.) Parodi, Revista Argent. Agron. 20: 14. 1953. Type: CHILE. Feuerland, Beagle [Beagle Kanal, zwischen (between) Steinen am Ufer; Tierra del Fuego, Remolino, Canal de Beagle], Mar. 1923, P. Gusinde 40 (holotype: BG; isotypes: BAA (BAA00000761 [image!]) fragm., W (W1941-0001573 [image!])).
</paragraph>
<paragraph pageId="0" pageNumber="29">
=
<taxonomicName authorityName="Hack ex Sodiro" authorityYear="1930" class="Liliopsida" family="Poaceae" genus="Deyeuxia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deyeuxia crassifolia" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="crassifolia">Deyeuxia crassifolia</taxonomicName>
Hack. ex Sodiro, Revista Col. Nac. Vicente Rocafuerte 12: 64, 73. 1930. Type: ECUADOR. Crece en los pajonales del Monte Pichincha, Sep. [Jun.] 1887 [1886], Sodiro 25/9 [s.n.] (holotype: Q; isotypes: QPSL, S (S-R-1457 [image!]), US (US00406342!), W (W1916-0038065 [image!], W1916-0038066 [image!])).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29" type="comments">
<paragraph pageId="0" pageNumber="29">Comments.</paragraph>
<paragraph pageId="0" pageNumber="29">
Previously considered to have its northernmost distribution in Ecuador (
<bibRefCitation author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae Volumen 3 Tomo 2. IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="29" pagination="180 - 218" refId="B33" refString="Rugolo de Agrasar, ZE, 2012. Deyeuxia Clarion ex P. Beauv. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae Volumen 3 Tomo 2. IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 180 - 218" title="Deyeuxia Clarion ex P. Beauv." volumeTitle="Flora Argentina." year="2012">
<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
de Agrasar 2012
</bibRefCitation>
), although
<bibRefCitation author="Luteyn, JL" journalOrPublisher="Memoirs of the New York Botanical Garden" pageId="0" pageNumber="29" refId="B20" refString="Luteyn, JL, 1999. Paramos, a checklist of plant diversity, geographical distribution, and botanical literature. Memoirs of the New York Botanical Garden 84: viii-xv, 1-278." title="Paramos, a checklist of plant diversity, geographical distribution, and botanical literature. Memoirs of the New York Botanical Garden 84: viii-xv, 1 - 278." year="1999">Luteyn (1999)</bibRefCitation>
mentions its presence in Costa Rica, this is the first record of
<taxonomicName class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. rigida" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="29">C. rigida</emphasis>
</taxonomicName>
for Colombia. It has been erroneously determined as
<taxonomicName authorityName="Trin ex Steud" authorityYear="1840" baseAuthorityName="Kunth" class="Liliopsida" family="Gramineae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis recta" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="recta">
<emphasis italics="true" pageId="0" pageNumber="29">Calamagrostis recta</emphasis>
</taxonomicName>
(Kunth) Trin. ex Steud. The latter differs from
<taxonomicName class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. rigida" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="29">C. rigida</emphasis>
</taxonomicName>
by the rachilla hairs reaching up to
<normalizedToken originalValue="¾">3/4</normalizedToken>
the length of the lemma (as opposed to usually reaching to slightly surpassing the lemma apex in
<taxonomicName class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. rigida" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="29">C. rigida</emphasis>
</taxonomicName>
), the awns being generally longer (6.2-7.5 mm as opposed to 4-6 mm in
<taxonomicName class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. rigida" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="29">C. rigida</emphasis>
</taxonomicName>
) and only slightly surpassing the glumes, the lemma apex being slightly bidentate (as opposed to bifid in
<taxonomicName class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. rigida" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="29">C. rigida</emphasis>
</taxonomicName>
) and the ligule being generally truncate and shorter, 1
<normalizedToken originalValue="5(">-5(-</normalizedToken>
6.6) mm long (as opposed to long acuminate, (3
<normalizedToken originalValue=")8">-)8-</normalizedToken>
12 mm long in
<taxonomicName class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. rigida" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="29">C. rigida</emphasis>
</taxonomicName>
), amongst other things.
</paragraph>
<paragraph pageId="0" pageNumber="29">
There has been discrepancy regarding the typification of
<taxonomicName authorityName="Griseb" authorityYear="1879" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis antoniana" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="antoniana">
<emphasis italics="true" pageId="0" pageNumber="29">Agrostis antoniana</emphasis>
</taxonomicName>
, with previous research (e.g.
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<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
de Agrasar 2012
</bibRefCitation>
) citing the specimen P.G. Lorentz and G.H.E.W. Hieronymus 72 housed at CORD herbarium from Argentina as holotype, while the protologue mentions this (although not explicitly giving collector and number) as well as three other collections: Spruce pl. ecuad. 5927 presumably from Ecuador, G. Mandon 1308 from Bolivia [Peru] and W. Lechler 1800 from Peru. A further specimen, P.G. Lorentz and G.H.E.W. Hieronymus 67, makes a total of at least five syntypes for this name, although the three GOET Lorentz and Hieronymus syntypes were annotated by Grisebach with &quot;67 + 72&quot;.
<bibRefCitation author="Hitchcock, AS" journalOrPublisher="Contributions from the United States National Herbarium" pageId="0" pageNumber="29" pagination="291 - 556" refId="B12" refString="Hitchcock, AS, 1927. The grasses of Ecuador, Peru, and Bolivia. Contributions from the United States National Herbarium 24 (8): 291 - 556" title="The grasses of Ecuador, Peru, and Bolivia." volume="24" year="1927">Hitchcock (1927)</bibRefCitation>
gave a partial lectotypification by dictating the type as Lechler 1800, but did not indicate the herbarium. The Lechler 1800 specimen was distributed with the following label annotated by Hohenacker: &quot;W. Lechler Pl. Peruvian. Ed. R. F. Hohenacker 1800
<taxonomicName class="Liliopsida" family="Gramineae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="genus">Calamagrostis</taxonomicName>
Antoniana Steud. Ipse [&quot;he himself said it&quot;]. In graminosis pr. San Antonio rara Jun. m.&quot;. As no annotations by Grisebach were found on any of the Lechler 1800, Mandon 1308 or Lorentz and Hieronymus 67 and 72 CORD specimens (Spruce pl. ecuad. 5927 specimens not found), with only Lechler 1800 specimens at P being verified by Steudel, we lectotypify to the best of the three GOET Lorentz and Hieronymus 67 + 72 syntypes annotated by Grisebach and consider syntypes Lorentz and Hieronymus 67 and Lorentz and Hieronymus 72 to be isolectotypes.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29" type="materials_examined">
<paragraph pageId="0" pageNumber="29">Specimens Examined.</paragraph>
<paragraph pageId="0" pageNumber="29">
COLOMBIA.
<emphasis bold="true" pageId="0" pageNumber="29">
<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
</emphasis>
: Mun. Chiscas,
<normalizedToken originalValue="páramo">paramo</normalizedToken>
de Chacaritas, asociado a rocas de 4 m de altura,
<geoCoordinate degrees="6" direction="north" minutes="37.3362" orientation="latitude" precision="1" value="6.62227">6°37.3362'N</geoCoordinate>
;
<geoCoordinate degrees="72" direction="west" minutes="23.424" orientation="longitude" precision="1" value="-72.3904">72°23.424'W</geoCoordinate>
, 4192 m alt., 4 Mar. 2018, S.P. Sylvester, R.J. Soreng, W. Bravo &amp; L.E. Cuta 3119 (COL, FMB, US, UPTC); Mun. Chiscas,
<normalizedToken originalValue="páramo">paramo</normalizedToken>
de Chacaritas, arribando a la morrena,
<geoCoordinate degrees="6" direction="north" minutes="37.0674" orientation="latitude" precision="1" value="6.61779">6°37.0674'N</geoCoordinate>
;
<geoCoordinate degrees="72" direction="west" minutes="23.3394" orientation="longitude" precision="1" value="-72.38899">72°23.3394'W</geoCoordinate>
, 4354 m alt., 4 Mar. 2018, S.P. Sylvester, R.J. Soreng, W. Bravo &amp; L.E. Cuta 3125 (COL, FMB, K, SI, UPTC, US); Mun. Chiscas,
<normalizedToken originalValue="páramo">paramo</normalizedToken>
de Chacaritas, arribando a la morrena,
<geoCoordinate degrees="6" direction="north" minutes="37.0674" orientation="latitude" precision="1" value="6.61779">6°37.0674'N</geoCoordinate>
;
<geoCoordinate degrees="72" direction="west" minutes="23.3394" orientation="longitude" precision="1" value="-72.38899">72°23.3394'W</geoCoordinate>
, 4354 m alt., 4 Mar. 2018, S.P. Sylvester, R.J. Soreng, W. Bravo &amp; L.E. Cuta 3126 (US). Dep. Santander:
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de la Angostura, Vereda El Mortino, Ubicada en borde de quebrada,
<geoCoordinate degrees="6" direction="north" minutes="57.5" orientation="latitude" precision="92" value="6.9583335">6°57.5'N</geoCoordinate>
;
<geoCoordinate degrees="72" direction="west" minutes="43.5" orientation="longitude" precision="92" value="-72.725">72°43.5'W</geoCoordinate>
, 3605 m alt., 17 Nov. 2007, M.C. Gomez 1 (US).
</paragraph>
</subSubSection>
</treatment>
</document>