157 lines
19 KiB
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157 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.4.60525" ID-GBIF-Dataset="217bea89-5bd1-4ae7-bfce-fb945a933fc4" ID-GBIF-Taxon="176088092" ID-Pensoft-Pub="2535-0730-2-119" ID-Pensoft-UUID="6301B37BBB7D51A185FB132ED24E4E81" ID-ZooBank="8BD62DBF438C46508C4037C837FD573F" LSID="urn:lsid:zoobank.org:act:97DC4390-2AC3-49A8-A693-E942FA9AA6B5" ModsDocID="2535-0730-2-119" checkinTime="1608157809646" checkinUser="pensoft" docAuthor="Ingrisch, Sigfrid" docDate="2020" docId="0E05572C4EF8500F89E49E54BE90B8DA" docLanguage="en" docName="EvolutSyst 4(2): 119-132" docOrigin="Evolutionary Systematics 4 (2)" docPubDate="2020-12-15" docSource="http://dx.doi.org/10.3897/evolsyst.4.60525" docTitle="Depressacca kinabalu Ingrisch 2020, sp. nov." docType="treatment" docUuid="97DC4390-2AC3-49A8-A693-E942FA9AA6B5" docUuidSource="ZooBank" docVersion="3" id="6301B37BBB7D51A185FB132ED24E4E81" lastPageNumber="119" masterDocId="6301B37BBB7D51A185FB132ED24E4E81" masterDocTitle="New subgenera and species of Agraeciini (Orthoptera, Tettigoniidae, Conocephalinae) from South Asia found in historical insect collections" masterLastPageNumber="132" masterPageNumber="119" pageId="0" pageNumber="119" updateTime="1643451831480" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New subgenera and species of Agraeciini (Orthoptera, Tettigoniidae, Conocephalinae) from South Asia found in historical insect collections</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ingrisch, Sigfrid</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8624-0472</mods:nameIdentifier>
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<mods:affiliation>Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier type="email">s.ingrisch@leibniz-zfmk.de</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-12-15</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>119</mods:start>
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<mods:end>132</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/evolsyst.4.60525</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.4.60525</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2535-0730-2-119</mods:identifier>
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<mods:identifier type="ZooBank">8BD62DBF438C46508C4037C837FD573F</mods:identifier>
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<mods:identifier type="Pensoft-UUID">6301B37BBB7D51A185FB132ED24E4E81</mods:identifier>
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</mods:mods>
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<subSection pageId="0" pageNumber="119" type="systematics">
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<treatment ID-GBIF-Taxon="176088092" LSID="urn:lsid:zoobank.org:act:97DC4390-2AC3-49A8-A693-E942FA9AA6B5" httpUri="http://treatment.plazi.org/id/0E05572C4EF8500F89E49E54BE90B8DA" lastPageNumber="119" pageId="0" pageNumber="119">
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<subSubSection pageId="0" pageNumber="119" type="nomenclature">
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<paragraph pageId="0" pageNumber="119">
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<taxonomicName LSID="http://zoobank.org/97DC4390-2AC3-49A8-A693-E942FA9AA6B5" authority="Ingrisch, 2020" authorityName="Ingrisch" authorityYear="2020" class="Insecta" family="Tettigoniidae" genus="Depressacca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Depressacca kinabalu" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="kinabalu" status="sp. nov.">Depressacca kinabalu</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="119">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">Fig. 7A-J</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="holotype">
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<paragraph pageId="0" pageNumber="119">Holotype</paragraph>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">(male).</emphasis>
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East Malaysia: Sabah, Mt. Kinabalu, Mesilau (
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<geoCoordinate degrees="6" direction="north" minutes="10" orientation="latitude" precision="925" value="6.1666665">6°10'N</geoCoordinate>
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,
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<geoCoordinate degrees="116" direction="east" minutes="39" orientation="longitude" precision="925" value="116.65">116°39'E</geoCoordinate>
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), at light, 4.ii.1964, coll. J. Smart, Royal Soc. Exped. - depository: BMNH (Lo033S001; B.M.1964-250). The type is unique.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="etymology">
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<paragraph pageId="0" pageNumber="119">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="119">The name of the new species refers to the type locality; noun in apposition.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="diagnosis">
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<paragraph pageId="0" pageNumber="119">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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So far only one
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<taxonomicName authorityName="Ingrisch" authorityYear="1998" class="Insecta" family="Tettigoniidae" genus="Depressacca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Depressacca" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="119">Depressacca</emphasis>
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</taxonomicName>
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species has been known. The new species differs from
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<taxonomicName genus="D." lsidName="D. globosa" pageId="0" pageNumber="119" rank="species" species="globosa">
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<emphasis italics="true" pageId="0" pageNumber="119">D. globosa</emphasis>
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</taxonomicName>
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Ingrisch, 1998 by the male tenth abdominal tergite that does not have projections but a slightly concave apex, by the male subgenital plate that does not have a medial process and that has only minute styli, by different shapes of the titillators, by the cerci that have the internal tooth inserted subapical, not apical, and by the fastigium verticis that is provided with a distinct tubercle at base.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="description">
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<paragraph pageId="0" pageNumber="119">Description.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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Fastigium verticis upcurved, with a tubercle at base; apex obtuse; separated by a shallow furrow from fastigium frontis (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7B</figureCitation>
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). Eyes projecting laterad (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7A</figureCitation>
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). Frons little rugose (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7D</figureCitation>
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). Pronotum strongly rugose, depressed, wide; anterior margin truncate in middle; posterior margin little concave; transverse furrows indistinct, widely interrupted in middle; paranota at anterior margin with a spine; ventro-posterior margin sinuate, finely crenulated; auditory swelling greatly swollen but indistinctly outlined; humeral sinus weakly indicated (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7A, C</figureCitation>
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). Prosternum mute; meso- and metasternum much wider than long, without lobes. Tegmen little surpassing hind knees, nearly covering abdomen; narrowing from base to rounded apex (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7E</figureCitation>
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). Femora rugose, rather short; spines on anterior side distinctly longer than on posterior side. Fore tibia with spines on ventro-internal side much longer than on external side; spines of hind tibia on dorso-internal side more than twice as long as on the other three margins. Femora with the following number of spines on ventral margins: profemur 3 small external, 4 large internal; mesofemur 5 large external, 2 small internal; postfemur 6-7 mostly large external, no internal. Knee lobes of fore femur obtuse on both sides; of mid femur obtuse on external, acute on internal side; of hind femur spinose on both sides.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Figure 7.</emphasis>
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<taxonomicName authorityName="Ingrisch" authorityYear="2020" class="Insecta" family="Tettigoniidae" genus="Depressacca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Depressacca kinabalu" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="kinabalu">
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<emphasis italics="true" pageId="0" pageNumber="119">Depressacca kinabalu</emphasis>
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</taxonomicName>
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sp. nov. male holotype:
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<emphasis bold="true" pageId="0" pageNumber="119">A</emphasis>
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habitus dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="119">B</emphasis>
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fastigium verticis lateral;
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<emphasis bold="true" pageId="0" pageNumber="119">C</emphasis>
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head, pronotum and base of tegmen lateral;
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<emphasis bold="true" pageId="0" pageNumber="119">D</emphasis>
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face;
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<emphasis bold="true" pageId="0" pageNumber="119">E</emphasis>
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left tegmen;
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<emphasis bold="true" pageId="0" pageNumber="119">F</emphasis>
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stridulatory file;
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<emphasis bold="true" pageId="0" pageNumber="119">G</emphasis>
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subgenital plate and cerci in ventral view;
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<emphasis bold="true" pageId="0" pageNumber="119">H</emphasis>
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left cercus in dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="119">I</emphasis>
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tip of abdomen in apical view;
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<emphasis bold="true" pageId="0" pageNumber="119">J</emphasis>
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titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Male.</emphasis>
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Tegmen with stridulatory area strongly projecting (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7E</figureCitation>
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). Stridulatory file about 1.80 mm long along diameter, with about 210 very dense teeth, plus at end 4 spaced, wider teeth that extend the file length to 1.86 mm along diameter; in middle of file with about 111 teeth at 1 mm or 55.5 teeth per 0.5 mm (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7F</figureCitation>
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). Tenth abdominal tergite little projecting between insertion of cerci; apical margin broadly concave. Epiproct Y-shaped; lateral margins projecting as rounded lobes, except in apical area (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7G, I</figureCitation>
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). Paraprocts with a small, compressed, obtuse projection (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7I</figureCitation>
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). Cerci short, conical; at apex with a stout, subacute, internal projection; basal of projection on internal side with another, small, triangular projection (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7H</figureCitation>
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). Subgenital plate broad, with indication of a faint medial carina; apex broad obtuse-angularly excised. Styli reduced to small tubercles (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7G</figureCitation>
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). Titillators subfused in middle; basal lobes, central area and base of apical lobes hyaline; apical lobes afterwards light brown, of roughly triangular outline and with wavy surface; margin crenulate, slightly bilobate with margin of external lobe provided with three acute teeth (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Depressacca kinabalu sp. nov. male holotype: A habitus dorsal view; B fastigium verticis lateral; C head, pronotum and base of tegmen lateral; D face; E left tegmen; F stridulatory file; G subgenital plate and cerci in ventral view; H left cercus in dorsal view; I tip of abdomen in apical view; J titillators of phallus. Abbreviations: 10 tenth abdominal tergite, c cercus, e epiproct, p paraproct, st stylus." httpUri="https://binary.pensoft.net/fig/487064" pageId="0" pageNumber="119">7J</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="coloration">
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<paragraph pageId="0" pageNumber="119">Coloration.</paragraph>
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<paragraph pageId="0" pageNumber="119">Discolored light yellowish brown; might have been green when alive.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="description">
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<paragraph pageId="0" pageNumber="119">Measurements of male.</paragraph>
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<paragraph pageId="0" pageNumber="119">Body: 17; pronotum: 5.7; tegmen: 10.2; hind femur: 7.8 mm.</paragraph>
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</subSubSection>
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</treatment>
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</subSection>
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</document> |