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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.430.8094" ID-PMC="PMC4141162" ID-Pensoft-Pub="1313-2970-430-1" ID-Pensoft-UUID="FFE6FF8CFFD43B66335AD2203774DB49" ID-PubMed="25152663" ID-Zenodo-Dep="578441" ID-ZooBank="86F760563B8B49FB9C86FAD0DB0CBE8C" ModsDocID="1313-2970-430-1" checkinTime="1451245551152" checkinUser="pensoft" docAuthor="Erwin, Terry L. &amp; Zamorano, Laura S." docDate="2014" docId="07E9EE11E14FA9CA30C0EC4B123B6D5C" docLanguage="en" docName="ZooKeys 430: 1-108" docOrigin="ZooKeys 430" docPubDate="2014-08-01" docSource="http://dx.doi.org/10.3897/zookeys.430.8094" docTitle="Asklepia hilaris Erwin &amp; Zamorano 2014, comb. n." docType="treatment" docVersion="5" id="FFE6FF8CFFD43B66335AD2203774DB49" lastPageNumber="40" masterDocId="FFE6FF8CFFD43B66335AD2203774DB49" masterDocTitle="A synopsis of the tribe Lachnophorini, with a new genus of Neotropical distribution and a revision of the Neotropical genus Asklepia Liebke, 1938 (Insecta, Coleoptera, Carabidae)" masterLastPageNumber="108" masterPageNumber="1" pageNumber="39" updateTime="1668159067277" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A synopsis of the tribe Lachnophorini, with a new genus of Neotropical distribution and a revision of the Neotropical genus Asklepia Liebke, 1938 (Insecta, Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Erwin, Terry L.</mods:namePart>
<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamorano, Laura S.</mods:namePart>
<mods:affiliation>Research Student, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA &amp; Laboratorio de Zoologia Acuatica LAZOEA, Departamento de Ciencias Biologicas, Universidad de los Andes, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-08-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>430</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>108</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.430.8094</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.430.8094</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-430-1</mods:identifier>
<mods:identifier type="ZooBank">86F760563B8B49FB9C86FAD0DB0CBE8C</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFE6FF8CFFD43B66335AD2203774DB49</mods:identifier>
<mods:identifier type="Zenodo-Dep">578441</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152054784" LSID="urn:lsid:plazi:treatment:07E9EE11E14FA9CA30C0EC4B123B6D5C" httpUri="http://treatment.plazi.org/id/07E9EE11E14FA9CA30C0EC4B123B6D5C" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<subSubSection pageId="38" pageNumber="39" type="nomenclature">
<paragraph pageId="38" pageNumber="39">
<taxonomicName LSID="07E9EE11-E14F-A9CA-30C0-EC4B123B6D5C" authority="(Bates, 1871)" authorityName="Erwin &amp; Zamorano" authorityYear="2014" baseAuthorityName="Bates" baseAuthorityYear="1871" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Asklepia hilaris" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hilaris" status="comb. n.">
<pageBreakToken pageId="38" pageNumber="39" start="start">Asklepia</pageBreakToken>
hilaris (Bates, 1871)
</taxonomicName>
<taxonomicNameLabel pageId="38" pageNumber="39">comb. n.</taxonomicNameLabel>
Cheerful pattern-wing beetle
<figureCitation captionStart="Figure 3336" captionStartId="F10" captionText="Figure 33 - 36. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 33 Asklepia grammechrysea Zamorano &amp; Erwin, sp. n. ADP 133151, Pithecia, Peru 34 Asklepia hilaris (Bates, 1871), comb. n., Brazil ADP 132543, Sao Paulo de Olivenca, Brazil 35 Asklepia laetitia Zamorano &amp; Erwin, sp. n. ADP 109190, Leticia, Colombia 36 Asklepia lebioides (Bates, 1871), comb. n., ADP 109194, Santarem, Rio Tapajos, Brazil. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30640" pageId="38" pageNumber="39">Figs 34</figureCitation>
<figureCitation captionStart="Figure 7677" captionStartId="F21" captionText="Figure 76 - 77. 76 Distribution map for known localities of Asklepia geminata (Bates) 77 Distribution map for known localities of Asklepia species of the hilaris group." httpUri="https://binary.pensoft.net/fig/30651" pageId="38" pageNumber="39">, 77</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="reference_group">
<paragraph pageId="38" pageNumber="39">
<taxonomicName authorityName="Erwin &amp; Zamorano" authorityYear="2014" class="Insecta" family="Carabidae" genus="Eucaerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hilaris" status="comb. n.">Eucaerus hilaris</taxonomicName>
Bates, 1871:79.
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="holotype">
<paragraph pageId="38" pageNumber="39">Holotype.</paragraph>
<paragraph pageId="38" pageNumber="39">
<emphasis bold="true" pageId="38" pageNumber="39">Brazil</emphasis>
, Amazonas,
<normalizedToken originalValue="São">Sao</normalizedToken>
Paulo de
<normalizedToken originalValue="Olivença">Olivenca</normalizedToken>
,
<geoCoordinate degrees="3.4622" direction="south" orientation="latitude" precision="5" value="-3.4622">3.4622°S</geoCoordinate>
,
<geoCoordinate degrees="68.9499" direction="west" orientation="longitude" precision="5" value="-68.9499">68.9499°W</geoCoordinate>
, 77m, (H.W. Bates)(BMNH: ADP132543, male). This specimen labeled
<normalizedToken originalValue="“Holotype”">&quot;Holotype&quot;</normalizedToken>
by George E. Ball in 1972.
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="derivation of specific epithet">
<paragraph pageId="38" pageNumber="39">Derivation of specific epithet.</paragraph>
<paragraph pageId="38" pageNumber="39">
The specific epithet,
<emphasis italics="true" pageId="38" pageNumber="39">hilaris</emphasis>
, is a Latin adjective that adequately describes this species with gaily colorful elytra.
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="proposed english vernacular name">
<paragraph pageId="38" pageNumber="39">Proposed english vernacular name.</paragraph>
<paragraph pageId="38" pageNumber="39">Cheerful pattern-wing beetle.</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="diagnosis">
<paragraph pageId="38" pageNumber="39">Diagnosis.</paragraph>
<paragraph pageId="38" pageNumber="39">
With the attributes of the genus
<taxonomicName authorityName="Liebke" authorityYear="1938" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="38" pageNumber="39">Asklepia</emphasis>
</taxonomicName>
as described by
<bibRefCitation author="Liebke, M" journalOrPublisher="Embrik Strand" pageId="81" pageNumber="82" pagination="37 - 141" refId="B54" refString="Liebke, M, 1938. Denkschrift ueber die Carabiden-Tribus Colliurini. Festschrift zum 60. Geburtstage von Profesor Dr. Embrik Strand 4: 37 - 141" title="Denkschrift ueber die Carabiden-Tribus Colliurini. Festschrift zum 60. Geburtstage von Profesor Dr." volume="4" year="1938">Liebke (1938)</bibRefCitation>
and as noted above under the generic diagnosis, and small-sized for the genus (SBL = 2.336 mm). Adults with head and prothorax brunneus, elytral maculae fulvous or aurantiacus in some individuals; elytron brunneus with a rectangular flavous macula crossing basal lateral and proximal quadrants, macula not extended to lateral margin or sutural area, and a rounded flavous macula in the apical proximal quadrant, sutural area fulvous; metasternum, abdominal sterna III-VI, and epipleuron testaceous, abdominal sternum VII paler; legs testaceous; antennal scape and pedicel testaceous, antennomeres 3-6 deeply infuscated, 7-11 white. Dorsal surface devoid of microsculpture, surface luster very shiny. Pronotum cordiform, narrowly explanate; anterior angles feebly produced; lateral margin beaded; hind angle angulate, moderately prominent; median line moderately defined. Elytral interneurs evident as rows of discontinuous, coarse punctures widely spaced.
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="description">
<paragraph pageId="38" pageNumber="39">Description.</paragraph>
<paragraph pageId="38" pageNumber="39">
<emphasis bold="true" italics="true" pageId="38" pageNumber="39">Habitus</emphasis>
(
<figureCitation captionStart="Figure 3336" captionStartId="F10" captionText="Figure 33 - 36. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 33 Asklepia grammechrysea Zamorano &amp; Erwin, sp. n. ADP 133151, Pithecia, Peru 34 Asklepia hilaris (Bates, 1871), comb. n., Brazil ADP 132543, Sao Paulo de Olivenca, Brazil 35 Asklepia laetitia Zamorano &amp; Erwin, sp. n. ADP 109190, Leticia, Colombia 36 Asklepia lebioides (Bates, 1871), comb. n., ADP 109194, Santarem, Rio Tapajos, Brazil. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30640" pageId="38" pageNumber="39">Fig. 34</figureCitation>
).
<emphasis bold="true" pageId="38" pageNumber="39">
<emphasis italics="true" pageId="38" pageNumber="39">Size</emphasis>
:
</emphasis>
[See also
<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Asklepia hilaris (Bates, 1871), comb. n." httpUri="http://table.plazi.org/id/2E6ACB460777023242893FC9D21A9972" pageId="38" pageNumber="39" tableUuid="2E6ACB460777023242893FC9D21A9972">Table 6</tableCitation>
] Small in size for the genus; ABL = 2.464 mm, SBL = 2.336 mm, TW (total width) 1.233 mm, LP = 0.483 mm, WP = 0.617 mm, LE = 1.495 mm.
<emphasis bold="true" pageId="38" pageNumber="39">
<emphasis italics="true" pageId="38" pageNumber="39">Color</emphasis>
:
</emphasis>
See diagnosis above.
<emphasis bold="true" pageId="38" pageNumber="39">
<emphasis italics="true" pageId="38" pageNumber="39">Luster</emphasis>
:
</emphasis>
See diagnosis above.
<emphasis bold="true" italics="true" pageId="38" pageNumber="39">Head</emphasis>
(
<figureCitation captionStart="Figure 3336" captionStartId="F10" captionText="Figure 33 - 36. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 33 Asklepia grammechrysea Zamorano &amp; Erwin, sp. n. ADP 133151, Pithecia, Peru 34 Asklepia hilaris (Bates, 1871), comb. n., Brazil ADP 132543, Sao Paulo de Olivenca, Brazil 35 Asklepia laetitia Zamorano &amp; Erwin, sp. n. ADP 109190, Leticia, Colombia 36 Asklepia lebioides (Bates, 1871), comb. n., ADP 109194, Santarem, Rio Tapajos, Brazil. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30640" pageId="38" pageNumber="39">Fig. 34</figureCitation>
): as in description for genus above.
<emphasis bold="true" pageId="38" pageNumber="39">
<emphasis italics="true" pageId="38" pageNumber="39">Prothorax</emphasis>
.
</emphasis>
Pronotum (
<figureCitation captionStart="Figure 3336" captionStartId="F10" captionText="Figure 33 - 36. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 33 Asklepia grammechrysea Zamorano &amp; Erwin, sp. n. ADP 133151, Pithecia, Peru 34 Asklepia hilaris (Bates, 1871), comb. n., Brazil ADP 132543, Sao Paulo de Olivenca, Brazil 35 Asklepia laetitia Zamorano &amp; Erwin, sp. n. ADP 109190, Leticia, Colombia 36 Asklepia lebioides (Bates, 1871), comb. n., ADP 109194, Santarem, Rio Tapajos, Brazil. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30640" pageId="38" pageNumber="39">Fig. 34</figureCitation>
) moderately broad, as wide as head across eyes (WH/WP: 1.085), longer than head (LP/LH: 1.346), wider than long (W/L: 1.294); markedly cordiform and explanate, lateral margin beaded with seta at anterior third; apical margin straight; base markedly constricted; hind angle moderately produced and setose; median line moderately well defined, apical transverse impression punctate, punctures infuscated; surface smooth throughout.
<emphasis bold="true" pageId="38" pageNumber="39">
<emphasis italics="true" pageId="38" pageNumber="39">Pterothorax</emphasis>
.
</emphasis>
Normal for genus, see description for genus above. Elytra slightly convex; at apical third about same width as head across eyes (WH/TW: 1.099) and pronotum (WP/TW: 1.013). Elytral interneurs evident as rows of discontinuous, widely spaced, coarse punctures. Hind wings fully developed.
<emphasis bold="true" pageId="38" pageNumber="39">
<emphasis italics="true" pageId="38" pageNumber="39">Legs</emphasis>
.
</emphasis>
Overall, normal for genus, see description for genus above.
<emphasis bold="true" pageId="38" pageNumber="39">
<emphasis italics="true" pageId="38" pageNumber="39">Abdominal sterna</emphasis>
.
</emphasis>
Overall, normal for genus, see description for genus above.
<emphasis bold="true" pageId="38" pageNumber="39">
<emphasis italics="true" pageId="38" pageNumber="39">Male genitalia</emphasis>
.
</emphasis>
Dissected by G.E. Ball in 1972 at the BMNH, but the genitalia are missing from the microvial pinned under the holotype.
<emphasis bold="true" pageId="38" pageNumber="39">
<emphasis italics="true" pageId="38" pageNumber="39">Female genitalia</emphasis>
.
</emphasis>
Not investigated, presumably similar to that of
<taxonomicName authorityName="Erwin &amp; Zamorano" authorityYear="2014" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="demiti">
<emphasis italics="true" pageId="38" pageNumber="39">Asklepia demiti</emphasis>
</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="dispersal potential">
<paragraph pageId="39" pageNumber="40">
<pageBreakToken pageId="39" pageNumber="40" start="start">Dispersal</pageBreakToken>
potential.
</paragraph>
<paragraph pageId="39" pageNumber="40">These beetles are macropterous and probably capable of flight. They are moderately swift and agile runners.</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="distribution">
<paragraph pageId="39" pageNumber="40">Distribution.</paragraph>
<paragraph pageId="39" pageNumber="40">
(
<figureCitation captionStart="Figure 7677" captionStartId="F21" captionText="Figure 76 - 77. 76 Distribution map for known localities of Asklepia geminata (Bates) 77 Distribution map for known localities of Asklepia species of the hilaris group." httpUri="https://binary.pensoft.net/fig/30651" pageId="39" pageNumber="40">Fig. 77</figureCitation>
). This species has been found at only one location on the white-water system of the
<normalizedToken originalValue="Río">Rio</normalizedToken>
Amazonas drainage system. But that does not at all indicate its real distribution: as has been pointed out above, very small beetles are inadequately sampled, especially in the Neotropics.
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="way of life">
<paragraph pageId="39" pageNumber="40">Way of life.</paragraph>
<paragraph pageId="39" pageNumber="40">
See
<bibRefCitation author="Erwin, TL" journalOrPublisher="Revista Peruana de Entomologia" pageId="79" pageNumber="80" pagination="1 - 85" refId="B22" refString="Erwin, TL, 1991. Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Peru. Revista Peruana de Entomologia 33: 1 - 85" title="Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Peru." volume="33" year="1991">Erwin (1991)</bibRefCitation>
for a general description of the genus. No way of life information is available for this species other than that they occur in lowland Amazonia along the &quot;Rio Amazon.&quot;
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="other specimens examined">
<paragraph pageId="39" pageNumber="40">Other specimens examined.</paragraph>
<paragraph pageId="39" pageNumber="40">None.</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="note">
<paragraph pageId="39" pageNumber="40">Note.</paragraph>
<paragraph pageId="39" pageNumber="40">The holotype and only specimen seen by us is glued to a card and is somewhat damaged.</paragraph>
</subSubSection>
</treatment>
</document>