268 lines
31 KiB
XML
268 lines
31 KiB
XML
<document id="3791DB657209A4CE27E5BA1F133BE053" ID-DOI="10.5281/zenodo.213610" ID-GBIF-Dataset="636d954a-43e4-4da0-b8f5-715a28bcaab9" ID-ISSN="1175-5326" ID-Zenodo-Dep="213610" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460488474176" checkinUser="plazi" docAuthor="Bilton, David T & Perkins, Philip D" docDate="2012" docId="076887EAFF8BFFFFC4E2F97C254D5796" docLanguage="en" docName="zt03342p059.pdf" docOrigin="Zootaxa 3342" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pneuminion fontinalis Bilton & Perkins, 2012, new species" docType="treatment" docVersion="8" lastPageNumber="57" masterDocId="FB51FF92FF88FFF9C475FFC8276E5432" masterDocTitle="Two new species of Parhydraenini from South Africa (Coleoptera: Hydraenidae)" masterLastPageNumber="59" masterPageNumber="51" pageNumber="54" updateTime="1698309621046" updateUser="plazi">
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<mods:mods id="51D26C1C8D945E0E9FF8547F75F4272C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4EB358422E1D9EFB1A2231F69EA217A7">
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<mods:title id="8CB8A0724AF710EA1A14997CE162F253">Two new species of Parhydraenini from South Africa (Coleoptera: Hydraenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="44915E438004FF12906373DB77197888" type="personal">
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<mods:role id="46EF45A96323B10524AA8F208E52E015">
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<mods:roleTerm id="AD9F8DCDD31B2EEE94BFA872E169E4F6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FAD66C3B276433B13778023C86069D4C">Bilton, David T</mods:namePart>
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</mods:name>
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<mods:name id="E2C11571EB80B34346BE58D7612B3A2A" type="personal">
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<mods:roleTerm id="6A92B0CCED0A7B5B9E6370FEF047C6F5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9E4ACB47E2ACD2760AD460CD3DD76CAD">Perkins, Philip D</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="868060AB5C40D5B33C40CB8A5BAA513A">text</mods:typeOfResource>
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<mods:titleInfo id="80C0B5767E3B0A80B3ABB0357A32BB99">
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<mods:title id="25CD37ED2FA30CE057C1352B82F4EFB6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="980FC4D9BC81DD15FBFAEDB232CE9C30">
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<mods:date id="B1666AE1336A1CB92F066943A74EDB2E">2012</mods:date>
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<mods:detail id="A64B23B7F1D27530F5D9781586CB558E" type="volume">
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<mods:number id="AC83FA13E7CF6D9B416097D6D1B0E95D">3342</mods:number>
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<mods:start id="E261B975DACC814756E9489A72337AF2">51</mods:start>
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<mods:end id="4217A100762D21F46447DCEACD62C77F">59</mods:end>
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<mods:classification id="D1125942891227B879153B0216D6C1CB">journal article</mods:classification>
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<mods:identifier id="A20A1E6CC565324336041F64C23C5599" type="DOI">10.5281/zenodo.213610</mods:identifier>
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<mods:identifier id="4E5387BEB18DC7EE54A51B5771DFF336" type="GBIF-Dataset">636d954a-43e4-4da0-b8f5-715a28bcaab9</mods:identifier>
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<mods:identifier id="56ADCB16D44E39305AE76B455906B858" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="61C33FB6D058AD15523FECCE0AFBE0E3" type="Zenodo-Dep">213610</mods:identifier>
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<treatment id="076887EAFF8BFFFFC4E2F97C254D5796" ID-DOI="http://doi.org/10.5281/zenodo.6181037" ID-GBIF-Taxon="119519008" ID-Zenodo-Dep="6181037" LSID="urn:lsid:plazi:treatment:076887EAFF8BFFFFC4E2F97C254D5796" httpUri="http://treatment.plazi.org/id/076887EAFF8BFFFFC4E2F97C254D5796" lastPageId="6" lastPageNumber="57" pageId="3" pageNumber="54">
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<subSubSection id="C7DB6577FF8BFFFAC4E2F97C264952DD" pageId="3" pageNumber="54" type="nomenclature">
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<paragraph id="8F7E36FCFF8BFFFAC4E2F97C252852FC" blockId="3.[151,582,1716,1775]" box="[151,582,1716,1742]" pageId="3" pageNumber="54">
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<heading id="D4368190FF8BFFFAC4E2F97C252852FC" bold="true" box="[151,582,1716,1742]" fontSize="11" level="1" pageId="3" pageNumber="54" reason="1">
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<emphasis id="BDB5EAEEFF8BFFFAC4E2F97C252852FC" bold="true" box="[151,582,1716,1742]" pageId="3" pageNumber="54">
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<taxonomicName id="48C14D7FFF8BFFFAC4E2F97C26C952FC" ID-CoL="4KKV8" box="[151,423,1716,1742]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="fontinalis" status="sp. nov.">
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<emphasis id="BDB5EAEEFF8BFFFAC4E2F97C26C952FC" bold="true" box="[151,423,1716,1742]" italics="true" pageId="3" pageNumber="54">Pneuminion fontinalis</emphasis>
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="A6865795FF8BFFFAC5C3F97C252852FC" box="[438,582,1716,1742]" pageId="3" pageNumber="54" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8F7E36FCFF8BFFFAC4E2F91F264952DD" blockId="3.[151,582,1716,1775]" box="[151,295,1751,1775]" pageId="3" pageNumber="54">
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||
(
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<figureCitation id="17FA2A79FF8BFFFAC4EAF91F278352DD" box="[159,237,1751,1775]" captionStart="FIGURE 3" captionStartId="4.[151,250,1417,1440]" captionText="FIGURE 3. Pneuminion fontinalis n. sp., dorsal and lateral habitus of holotype." pageId="3" pageNumber="54">Figs. 3</figureCitation>
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&
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<figureCitation id="17FA2A79FF8BFFFAC565F91F267152DD" box="[272,287,1751,1775]" captionStart="FIGURE 4" captionStartId="5.[151,250,1476,1499]" captionText="FIGURE 4. Pneuminion fontinalis n. sp., aedeagus of holotype." pageId="3" pageNumber="54">4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C7DB6577FF8BFFFDC4E2F8D7237054C5" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54" type="materials_examined">
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<paragraph id="8F7E36FCFF8BFFFAC4E2F8D724B05391" blockId="3.[151,1436,1822,2027]" pageId="3" pageNumber="54">
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<emphasis id="BDB5EAEEFF8BFFFAC4E2F8D726AE5305" bold="true" box="[151,448,1822,1847]" pageId="3" pageNumber="54">
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<typeStatus id="507A885EFF8BFFFAC4E2F8D727BC5305" box="[151,210,1823,1847]" pageId="3" pageNumber="54">Type</typeStatus>
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Material.
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<typeStatus id="507A885EFF8BFFFAC526F8D726AE5305" box="[339,448,1823,1847]" pageId="3" pageNumber="54" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):
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<date id="FB7F103CFF8BFFFAC657F8D725F15305" box="[546,671,1823,1847]" pageId="3" pageNumber="54" value="2008-09-26">26/ix/2008</date>
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,
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<collectingCountry id="F7D6766CFF8BFFFAC6D9F8D7242F5305" box="[684,833,1823,1847]" name="South Africa" pageId="3" pageNumber="54">South Africa</collectingCountry>
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, Western Cape Province, madicolous seepage beside R46 road in Mitchell’s Pass, ca.
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<quantity id="48399B19FF8BFFFAC67EF88B25285369" box="[523,582,1859,1883]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="54" unit="km" value="5.0">5 km</quantity>
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SW of Ceres,
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<geoCoordinate id="EAF5503BFF8BFFFAC685F88B24DC5369" box="[752,946,1859,1883]" direction="south" orientation="latitude" pageId="3" pageNumber="54" precision="1" value="-33.40257">33° 24’ 09.24” S</geoCoordinate>
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,
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<geoCoordinate id="EAF5503BFF8BFFFAC7B5F88B23E85369" box="[960,1158,1859,1883]" direction="east" orientation="longitude" pageId="3" pageNumber="54" precision="1" value="19.285822">19° 17’ 08.96” E</geoCoordinate>
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,
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<quantity id="48399B19FF8BFFFAC0E6F88B23B35369" box="[1171,1245,1859,1883]" metricMagnitude="2" metricUnit="m" metricValue="3.86" pageId="3" pageNumber="54" unit="m" value="386.0">386 m</quantity>
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, D.T. Bilton leg. (
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<collectionCode id="E9D0AE39FF8BFFFAC4EAF8AF278C534D" box="[159,226,1895,1919]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="3" pageNumber="54">SAM</collectionCode>
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).
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<emphasis id="BDB5EAEEFF8BFFFAC481F8AF26F3534D" bold="true" box="[244,413,1894,1919]" pageId="3" pageNumber="54">
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<typeStatus id="507A885EFF8BFFFAC481F8AF2602534D" box="[244,364,1895,1919]" pageId="3" pageNumber="54" type="paratype">Paratypes</typeStatus>
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(7):
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</emphasis>
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||
<date id="FB7F103CFF8BFFFAC5D0F8AF254F534D" box="[421,545,1895,1919]" pageId="3" pageNumber="54" value="2009-09-27">27/ix/2009</date>
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,
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<collectingCountry id="F7D6766CFF8BFFFAC659F8AF25AE534D" box="[556,704,1895,1919]" name="South Africa" pageId="3" pageNumber="54">South Africa</collectingCountry>
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, Western Cape Province, Mitchell’s Pass on R46 ca.
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<quantity id="48399B19FF8BFFFAC166F8AF2222534D" box="[1299,1356,1895,1919]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="54" unit="km" value="5.0">5 km</quantity>
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SW of Ceres, wet rock faces, D.T. Bilton leg. (1
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<collectionCode id="E9D0AE39FF8BFFFAC61DF84325C95391" box="[616,679,1931,1955]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="3" pageNumber="54">SAM</collectionCode>
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, 1
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<collectionCode id="E9D0AE39FF8BFFFAC6BDF84324775391" box="[712,793,1931,1955]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="3" pageNumber="54">TMSA</collectionCode>
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, 1
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<collectionCode id="E9D0AE39FF8BFFFAC74DF84324165391" LSID="urn:lsid:biocol.org:col:33791" box="[824,888,1931,1955]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="54">MCZ</collectionCode>
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, 3 CBP).
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</paragraph>
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<paragraph id="8F7E36FCFF8BFFFDC4B2F866237054C5" blockId="3.[151,1436,1822,2027]" lastBlockId="4.[151,1436,151,247]" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54">
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<emphasis id="BDB5EAEEFF8BFFFAC4B2F86626BD53F5" bold="true" box="[199,467,1966,1991]" pageId="3" pageNumber="54">Differential Diagnosis.</emphasis>
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Members of this species will key to
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<taxonomicName id="48C14D7FFF8BFFFAC706F878235253F5" box="[883,1084,1967,1991]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="balfourbrownei">
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<emphasis id="BDB5EAEEFF8BFFFAC706F878235253F5" box="[883,1084,1967,1991]" italics="true" pageId="3" pageNumber="54">P. balfourbrownei</emphasis>
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</taxonomicName>
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in Perkins (2004).
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<taxonomicName id="48C14D7FFF8BFFFAC163F87822F553F5" box="[1302,1435,1967,1991]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="fontinalis" status="sp. nov.">
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<emphasis id="BDB5EAEEFF8BFFFAC163F87822F553F5" box="[1302,1435,1967,1991]" italics="true" pageId="3" pageNumber="54">P. fontinalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A6865795FF8BFFFAC4E2F81C27B953D9" box="[151,215,2004,2027]" pageId="3" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
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is slightly smaller and slightly narrower, and has the elytra slightly shorter relative to the length of the pronotum. The two species differ distinctly in the pronotal sculpture: the foveae are very shallow in
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<taxonomicName id="48C14D7FFF8CFFFDC08FFF5122115482" box="[1274,1407,152,176]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="fontinalis" status="sp. nov.">
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<emphasis id="BDB5EAEEFF8CFFFDC08FFF5122115482" box="[1274,1407,152,176]" italics="true" pageId="4" pageNumber="55">P. fontinalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A6865795FF8CFFFDC1F2FF5027D954E6" pageId="4" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="17FA2A79FF8CFFFDC4B0FF74266654E6" box="[197,264,188,212]" captionStart="FIGURE 3" captionStartId="4.[151,250,1417,1440]" captionText="FIGURE 3. Pneuminion fontinalis n. sp., dorsal and lateral habitus of holotype." pageId="4" pageNumber="55">Fig. 3</figureCitation>
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), whereas the foveae are deep in
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<taxonomicName id="48C14D7FFF8CFFFDC60FFF75242F54E6" box="[634,833,188,212]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="balfourbrownei">
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<emphasis id="BDB5EAEEFF8CFFFDC60FFF75242F54E6" box="[634,833,188,212]" italics="true" pageId="4" pageNumber="55">P. balfourbrownei</emphasis>
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</taxonomicName>
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. The aedeagi of the two species differ distinctly in the shapes of both the flagellum and the mainpiece (
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<figureCitation id="17FA2A79FF8CFFFDC6C3FF17259554C5" box="[694,763,223,247]" captionStart="FIGURE 4" captionStartId="5.[151,250,1476,1499]" captionText="FIGURE 4. Pneuminion fontinalis n. sp., aedeagus of holotype." pageId="4" pageNumber="55">Fig. 4</figureCitation>
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; Fig.
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<quantity id="48399B19FF8CFFFDC74FFF17241D54C5" box="[826,883,223,247]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="4" pageNumber="55" unit="in" value="14.0">14 in</quantity>
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Perkins 2004).
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</paragraph>
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</subSubSection>
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||
<caption id="DBBE6674FF8CFFFDC4E2FA4124AF51AD" box="[151,961,1417,1440]" pageId="4" pageNumber="55">
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<paragraph id="8F7E36FCFF8CFFFDC4E2FA4124AF51AD" blockId="4.[151,961,1417,1440]" box="[151,961,1417,1440]" pageId="4" pageNumber="55">
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||
<emphasis id="BDB5EAEEFF8CFFFDC4E2FA41267C51AC" bold="true" box="[151,274,1417,1440]" pageId="4" pageNumber="55">FIGURE 3.</emphasis>
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<taxonomicName id="48C14D7FFF8CFFFDC56CFA42269451AD" box="[281,506,1417,1439]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="fontinalis" status="sp. nov.">
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<emphasis id="BDB5EAEEFF8CFFFDC56CFA42269451AD" box="[281,506,1417,1439]" italics="true" pageId="4" pageNumber="55">Pneuminion fontinalis</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A6865795FF8CFFFDC58AFA42255951AD" box="[511,567,1418,1439]" pageId="4" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
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, dorsal and lateral habitus of holotype.
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</paragraph>
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</caption>
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<subSubSection id="C7DB6577FF8CFFFFC4B2FA2D26F954CA" lastPageId="6" lastPageNumber="57" pageId="4" pageNumber="55" type="description">
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||
<paragraph id="8F7E36FCFF8CFFFDC4B2FA2D266B53C7" blockId="4.[151,1436,1509,2037]" pageId="4" pageNumber="55">
|
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<emphasis id="BDB5EAEEFF8CFFFDC4B2FA2D26F251CC" bold="true" box="[199,412,1509,1534]" pageId="4" pageNumber="55">Description. Size:</emphasis>
|
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(length/width, mm)
|
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<typeStatus id="507A885EFF8CFFFDC6F8FA2D259A51CF" box="[653,756,1509,1533]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
|
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: body (length to elytral apices) 1.53/ 0.65, head 0.29/0.45, pronotum 0.37/0.56, elytra 0.90/0.65.
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<emphasis id="BDB5EAEEFF8CFFFDC624F9C025C05213" bold="true" box="[593,686,1544,1569]" pageId="4" pageNumber="55">Colour:</emphasis>
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Dorsum black, hind angles of pronotum dark piceous; legs and maxillary palpi dark brown; venter black.
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<emphasis id="BDB5EAEEFF8CFFFDC6C8F9E524685277" bold="true" box="[701,774,1581,1605]" pageId="4" pageNumber="55">Head:</emphasis>
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Labrum and clypeus weakly shining, with shallow microreticulation which is more strongly developed towards lateral margins. Surface finely punctate; punctures sparse on centre of clypeus, where they are spaced approximately two puncture-widths apart; denser towards lateral margins. Frons sparsely punctate, shining and weakly microreticulate. Punctures denser, and microreticulation more strongly developed towards margins of eyes. Punctures on frons deeper than on clypeus, and separated by ca. 1–1.5 puncture diameters. Fovea in front of ocelli large, approximately
|
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<date id="FB7F103CFF8CFFFDC7B9F929248152CB" box="[972,1007,1761,1785]" pageId="4" pageNumber="55">4X</date>
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size of ocellus; densely punctate and strongly microreticulate, with abrupt median margin; more open towards eyes, and extending as a shallow, heavily punctate and microreticulate area between ocelli and eyes, and posteriorly to the hind margin of the head. Dorsal hyaline setae conspicuous; slender and transparent, at least 4–5 puncture diameters in length.
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<emphasis id="BDB5EAEEFF8CFFFDC0C9F88522515357" bold="true" box="[1212,1343,1869,1893]" pageId="4" pageNumber="55">Pronotum:</emphasis>
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Surface shining, with traces of obsolete microreticulation; punctures in reliefs small and with a granule at the front margins, bearing a hyaline seta; granule often projecting posteriorly, giving punctures the appearance of being paired, or horseshoe-shaped. Midlongitudinal fovea weak; anterior and posterior fovea both well-marked and shallowly extending
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</paragraph>
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<caption id="DBBE6674FF8DFFFCC4E2FA0C244E51E9" box="[151,800,1476,1499]" pageId="5" pageNumber="56">
|
||
<paragraph id="8F7E36FCFF8DFFFCC4E2FA0C244E51E9" blockId="5.[151,800,1476,1499]" box="[151,800,1476,1499]" pageId="5" pageNumber="56">
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||
<emphasis id="BDB5EAEEFF8DFFFCC4E2FA0C267D51E8" bold="true" box="[151,275,1476,1499]" pageId="5" pageNumber="56">FIGURE 4.</emphasis>
|
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<taxonomicName id="48C14D7FFF8DFFFCC56CFA0E269451E9" box="[281,506,1477,1499]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="fontinalis" status="sp. nov.">
|
||
<emphasis id="BDB5EAEEFF8DFFFCC56CFA0E269451E9" box="[281,506,1477,1499]" italics="true" pageId="5" pageNumber="56">Pneuminion fontinalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A6865795FF8DFFFCC58AFA0E255951E9" box="[511,567,1478,1499]" pageId="5" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
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, aedeagus of holotype.
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||
</paragraph>
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</caption>
|
||
<paragraph id="8F7E36FCFF8DFFFFC4E2F9CA26F954CA" blockId="5.[151,1437,1538,2030]" lastBlockId="6.[151,1436,151,932]" lastPageId="6" lastPageNumber="57" pageId="5" pageNumber="56">
|
||
towards midline, deeper towards lateral margins where punctures are much larger and open posteriolaterally. Punctures towards lateral margins often confluent, appearing as lowered, heavily microreticulate areas. Pronotal postocular impression shallow. Setae on pronotal reliefs long and transparent, adpressed against the surface, and orientated antero-posteriorly, reaching back at least to the anterior margin of next puncture, and at least 4–5 puncture diameters in length. Wings full-length.
|
||
<emphasis id="BDB5EAEEFF8DFFFCC6CDF95B24635299" bold="true" box="[696,781,1683,1707]" pageId="5" pageNumber="56">Elytra:</emphasis>
|
||
Shining and lacking microreticulation. Setae of series and intervals equally developed, resulting in about 20 discrete linear rows on each elytron. Setae granule-based, transparent, whitish and closely adpressed to elytral surface; relatively long, reaching back to insertion point of next seta in each row. Serial punctures moderately large, shallow and slightly narrower than intervals. Elytra with narrow explanate margin. Very weak saddle in middle of elytra ca. 1/3 from base.
|
||
<emphasis id="BDB5EAEEFF8DFFFCC05CF8EA23CB5309" bold="true" box="[1065,1189,1826,1851]" pageId="5" pageNumber="56">Aedeagus:</emphasis>
|
||
total length
|
||
<quantity id="48399B19FF8DFFFCC144F8EA22F65308" box="[1329,1432,1826,1850]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="5" pageNumber="56" unit="mm" value="0.61">0.61 mm</quantity>
|
||
; mainpiece length
|
||
<quantity id="48399B19FF8DFFFCC52BF88E26A8536C" box="[350,454,1862,1886]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="56" unit="mm" value="0.44">0.44 mm</quantity>
|
||
. Duct weakly arcuate, approximately
|
||
<date id="FB7F103CFF8DFFFCC71BF88E24FF536C" box="[878,913,1862,1886]" pageId="5" pageNumber="56">2X</date>
|
||
length of mainpiece, thick at base, but slender over most of length, particularly apical third, with abruptly expanded weakly sclerotized apex (only visible with transmitted light—duct appearing finely pointed with incident light at lower magnification). Mainpiece evenly curved, with a bluntly pointed, deflexed tip; apical third grooved and laterally arched to guide duct. Parameres slender, approximately same length as main piece, with long setae on apical sections.
|
||
<emphasis id="BDB5EAEEFF8DFFFCC02FF81E23D453DC" bold="true" box="[1114,1210,2006,2030]" pageId="5" pageNumber="56">Female:</emphasis>
|
||
Pronotal postocular impression deeper and less symmetrical than in males. Explanate margin of elytra wider than in males, and elytral sides more strongly rounded.
|
||
<emphasis id="BDB5EAEEFF8EFFFFC59DFF73253154E6" bold="true" box="[488,607,187,212]" pageId="6" pageNumber="57">Variation:</emphasis>
|
||
The strength of the pronotal impressions and elytral saddle vary slightly between the
|
||
<typeStatus id="507A885EFF8EFFFFC551FF2826FD54CA" box="[292,403,224,248]" pageId="6" pageNumber="57" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C7DB6577FF8EFFFFC4B2FECC27BE550D" pageId="6" pageNumber="57" type="etymology">
|
||
<paragraph id="8F7E36FCFF8EFFFFC4B2FECC27BE550D" blockId="6.[151,1436,151,932]" pageId="6" pageNumber="57">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC4B2FECC263F552E" bold="true" box="[199,337,260,284]" pageId="6" pageNumber="57">Etymology:</emphasis>
|
||
Named in reference to the habitat—damp areas beside permanent films of water flowing over rock.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C7DB6577FF8EFFFFC4B2FE83254D5796" pageId="6" pageNumber="57" type="distribution">
|
||
<paragraph id="8F7E36FCFF8EFFFFC4B2FE83254D5796" blockId="6.[151,1436,151,932]" pageId="6" pageNumber="57">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC4B2FE83256C5556" bold="true" box="[199,514,331,356]" pageId="6" pageNumber="57">Distribution and Ecology:</emphasis>
|
||
So far only reported from the
|
||
<typeStatus id="507A885EFF8EFFFFC71FFE8524F45557" box="[874,922,333,357]" pageId="6" pageNumber="57">type</typeStatus>
|
||
locality, a hygropetric seepage in Mitchell’s Pass near Ceres in the Western Cape Province (
|
||
<figureCitation id="17FA2A79FF8EFFFFC6CDFEA7259055B5" box="[696,766,367,391]" captionStart="FIGURE 6" captionStartId="7.[151,250,1600,1623]" captionTargetBox="[275,1312,193,1576]" captionTargetId="figure@7.[275,1312,193,1579]" captionTargetPageId="7" captionText="FIGURE 6. Type Locality of Pneuminion fontinalis n. sp. Hygropetric habitat beside R 46 at Mitchell’s Pass, ca. 5 km SW of Ceres, Western Cape Province, South Africa. Arrow indicates area in which the Holotype was collected. Photo D. T. Bilton." httpUri="https://zenodo.org/record/213615/files/figure.png" pageId="6" pageNumber="57">Fig. 6</figureCitation>
|
||
). At Mitchell’s Pass
|
||
<taxonomicName id="48C14D7FFF8EFFFFC79AFEB9238755BA" box="[1007,1257,368,392]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="fontinalis" status="sp. nov.">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC79AFEB9238755BA" box="[1007,1257,368,392]" italics="true" pageId="6" pageNumber="57">Pneuminion fontinalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A6865795FF8EFFFFC087FEB9225E55BA" box="[1266,1328,369,392]" pageId="6" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
|
||
has been found on damp rock with algal biofilms and fine clay beside permanent and semi-permanent trickles of water, formed where springs flow over exposed rock in a road cutting. The species appears to be relatively rare; the eight specimens of the
|
||
<typeStatus id="507A885EFF8EFFFFC515FE1526FE55C7" box="[352,400,477,501]" pageId="6" pageNumber="57">type</typeStatus>
|
||
series being found in approximately 4 hours of sampling over two visits. The habitat was searched by eye, and beetles obtained with a finger or a small length of grass stem, used to winkle them out of crevices. Specimens were found on lower parts of the rock face, mostly ca.
|
||
<quantity id="48399B19FF8EFFFFC796FDEC234C560E" box="[995,1058,548,572]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="57" unit="m" value="0.5">0.5 m</quantity>
|
||
above ground level, in areas with fine biofilms, and surrounding grass and fern layers, providing partial shade. Most beetles were visible on the open rock face, but some were found inhabiting the overhangs of small crevices. The seepage habitats at Mitchell’s Pass host a rich hygropetric fauna, this being one of the original localities in which the recently described relictual water beetle family
|
||
<taxonomicName id="48C14D7FFF8EFFFFC546FD7C26D456FE" box="[307,442,692,716]" class="Insecta" family="Aspidytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="family">Aspidytidae</taxonomicName>
|
||
was found (
|
||
<bibRefCitation id="EB504B0DFF8EFFFFC633FD7C244A56FE" author="Ribera" box="[582,804,692,716]" pageId="6" pageNumber="57" refString="Ribera, I., Beutel, R. G., Balke, M. & Vogler, A. P. (2002) Discovery of Aspidytidae, a new family of aquatic Coleoptera. Proceedings of the Royal Society of London Series B, 269, 2351 - 2356." type="journal article" year="2002">
|
||
Ribera,
|
||
<emphasis id="BDB5EAEEFF8EFFFFC6D4FD7D25B456FE" box="[673,730,692,716]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
|
||
, 2002
|
||
</bibRefCitation>
|
||
; Alarie & Bilton, 2007). Larvae and adults of
|
||
<taxonomicName id="48C14D7FFF8EFFFFC144FD7C25EF56C2" authority="Ribera, Beutel, Balke & Vogler, 2002" authorityName="Ribera, Beutel, Balke & Vogler" authorityYear="2002" class="Insecta" family="Aspidytidae" genus="Aspidytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="niobe">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC144FD7C27BB56C2" italics="true" pageId="6" pageNumber="57">Aspidytes niobe</emphasis>
|
||
<bibRefCitation id="EB504B0DFF8EFFFFC4A9FD1025EF56C2" author="Ribera" box="[220,641,728,752]" pageId="6" pageNumber="57" refString="Ribera, I., Beutel, R. G., Balke, M. & Vogler, A. P. (2002) Discovery of Aspidytidae, a new family of aquatic Coleoptera. Proceedings of the Royal Society of London Series B, 269, 2351 - 2356." type="journal article" year="2002">Ribera, Beutel, Balke & Vogler, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
occur in wetter areas of seepage than
|
||
<taxonomicName id="48C14D7FFF8EFFFFC05BFD11224856C2" box="[1070,1318,728,752]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="fontinalis" status="sp. nov.">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC05BFD11224856C2" box="[1070,1318,728,752]" italics="true" pageId="6" pageNumber="57">Pneuminion fontinalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A6865795FF8EFFFFC15BFD11220256C2" box="[1326,1388,729,752]" pageId="6" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
|
||
, but the new species is microsympatric with
|
||
<taxonomicName id="48C14D7FFF8EFFFFC61CFD34230D5726" authority="Omer-Cooper, 1965" authorityName="Omer-Cooper" authorityYear="1965" box="[617,1123,764,788]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="nebulosus">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC61CFD34241D5726" box="[617,883,764,788]" italics="true" pageId="6" pageNumber="57">Canthyporus nebulosus</emphasis>
|
||
Omer-Cooper, 1965
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="48C14D7FFF8EFFFFC007FD3422275726" box="[1138,1353,764,788]" class="Insecta" family="Dytiscidae" genus="Africophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="jansei">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC007FD3422275726" box="[1138,1353,764,788]" italics="true" pageId="6" pageNumber="57">Africophilus jansei</emphasis>
|
||
</taxonomicName>
|
||
Omer- Cooper & Omer-Cooper, 1957,
|
||
<taxonomicName id="48C14D7FFF8EFFFFC588FCE824D0570A" authority="Komarek, 2004" authorityName="Komarek" authorityYear="2004" box="[509,958,800,824]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="glabriventris">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC588FCE8246B570A" box="[509,773,800,824]" italics="true" pageId="6" pageNumber="57">Anacaena glabriventris</emphasis>
|
||
Komarek, 2004
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="48C14D7FFF8EFFFFC7BCFCE8239A570A" authority="Komarek, 2004" authorityName="Komarek" authorityYear="2004" box="[969,1268,800,824]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="reducta">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC7BCFCE82350570A" box="[969,1086,800,824]" italics="true" pageId="6" pageNumber="57">A. reducta</emphasis>
|
||
Komarek, 2004
|
||
</taxonomicName>
|
||
and a range of SW Cape endemic
|
||
<taxonomicName id="48C14D7FFF8EFFFFC51BFC8C256E576E" box="[366,512,836,860]" class="Insecta" family="Hydraenidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="family">Hydraenidae</taxonomicName>
|
||
, including the relatively widespread congener
|
||
<taxonomicName id="48C14D7FFF8EFFFFC07CFC8D22F6576E" authority="Perkins, 1997" authorityName="Perkins" authorityYear="1997" box="[1033,1432,836,860]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="velamen">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC07CFC8D239A576E" box="[1033,1268,836,860]" italics="true" pageId="6" pageNumber="57">Pneuminion velamen</emphasis>
|
||
<bibRefCitation id="EB504B0DFF8EFFFFC08EFC8C22F6576E" author="Perkins" box="[1275,1432,836,860]" pageId="6" pageNumber="57" refString="Perkins, P. D. (1997) Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of Carnegie Museum, 66, 89 - 207." type="journal article" year="1997">Perkins, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="48C14D7FFF8EFFFFC4E2FCA0251857B2" authority="Perkins, 2005" authorityName="Perkins" authorityYear="2005" box="[151,630,872,896]" class="Insecta" family="Hydraenidae" genus="Coelometopon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="blinkwater">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC4E2FCA026AF57B2" box="[151,449,872,896]" italics="true" pageId="6" pageNumber="57">Coelometopon blinkwater</emphasis>
|
||
Perkins, 2005
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="48C14D7FFF8EFFFFC6FCFCA0230C57B2" authority="Perkins, 2005" authorityName="Perkins" authorityYear="2005" box="[649,1122,872,896]" class="Insecta" family="Hydraenidae" genus="Oomtelecopon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="sebastiani">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC6FCFCA024C057B2" box="[649,942,872,896]" italics="true" pageId="6" pageNumber="57">Oomtelecopon sebastiani</emphasis>
|
||
Perkins, 2005
|
||
</taxonomicName>
|
||
, and a further species of
|
||
<taxonomicName id="48C14D7FFF8EFFFFC4E2FC4426545796" box="[151,314,908,932]" class="Insecta" family="Hydraenidae" genus="Oomtelecopon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BDB5EAEEFF8EFFFFC4E2FC4426545796" box="[151,314,908,932]" italics="true" pageId="6" pageNumber="57">Oomtelecopon</emphasis>
|
||
</taxonomicName>
|
||
still to be described.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |