250 lines
25 KiB
XML
250 lines
25 KiB
XML
<document id="ED456CE8B67F6160CD396DA53E069066" ID-DOI="10.11646/zootaxa.3956.4.3" ID-GBIF-Dataset="ef601d63-c211-4a45-a530-88385b13d87e" ID-ISSN="1175-5326" ID-Zenodo-Dep="236658" ID-ZooBank="428F1742-286A-4550-8EF7-CD22686C664F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461057603347" checkinUser="plazi" docAuthor="Jiménez-Ferbans, Larry, Reyes-Castillo, Pedro & Schuster, Jack C." docDate="2015" docId="4268D77B8049FFEDFF36147BFAD6FD0C" docLanguage="en" docName="zt03956p512.pdf" docOrigin="Zootaxa 3956 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Passalus (Pertinax) doesburgi Boucher 2015, new combination" docType="treatment" docVersion="7" lastPageNumber="504" masterDocId="BE51AF038045FFE0FFA1163CFFA7FFC0" masterDocTitle="Passalidae (Coleoptera: Scarabaeoidea) of the Greater and Lesser Antilles" masterLastPageNumber="512" masterPageNumber="491" pageNumber="503" updateTime="1698581430319" updateUser="plazi">
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<mods:mods id="1DE954FDA6818FBE846240804FB49FA8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2FCD7A2305555B32653F6B2E5540DE5C">
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<mods:title id="520A3E8642806412908F867CBDDBEA0A">Passalidae (Coleoptera: Scarabaeoidea) of the Greater and Lesser Antilles</mods:title>
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</mods:titleInfo>
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<mods:name id="EFF34AE3C036EB143F91438AC612065D" type="personal">
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<mods:role id="66813BFB1A92A80FADCE9D34CD29B547">
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<mods:roleTerm id="4B7D73D31CB857A5556B72976E814B12">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E7E304E6BC5647D88C2180E785021361">Jiménez-Ferbans, Larry</mods:namePart>
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<mods:roleTerm id="DC68B25EB441EA6B33741FC21C6B532D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A3FC3780C2DF3EF0C9DB774BD76BA0C1">Reyes-Castillo, Pedro</mods:namePart>
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</mods:name>
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<mods:name id="A845857F14D07DB1D70D8D025911BDF2" type="personal">
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<mods:roleTerm id="C8F50E22A0FC2A2C46453F7DDD69E95F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="546300DA824F0C8DC044AB99B3032A28">Schuster, Jack C.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="66915DE81F6C4F6B99093D45A1DD6BC0">text</mods:typeOfResource>
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<mods:titleInfo id="4AB1AFD7C10B1858E90409585402BD93">
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<mods:title id="4EA9A8C78558D2480228A5F8EF25BE35">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D6FDB21EEFB931315DEE3042C51B27D1">
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<mods:date id="38B869CEC7F7A7B560C1521212AFEEE1">2015</mods:date>
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<mods:detail id="02B5FDB07F20380F3C1D7CE247F810F6" type="volume">
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<mods:number id="E79443C36DE1D35546E5384686F16678">3956</mods:number>
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</mods:detail>
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<mods:detail id="0424FCD0EC9190EAEA4026517B22E1CF" type="issue">
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<mods:number id="463AE830C3F56A9EFB92ECC3BEECFD7F">4</mods:number>
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<mods:classification id="A4A6B0484CF9D29A81B6B703AD309AFA">journal article</mods:classification>
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<mods:identifier id="054607C5B6D451E610AC75D31ABB9FCA" type="DOI">10.11646/zootaxa.3956.4.3</mods:identifier>
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<mods:identifier id="5D5DAF78027BD66299ECD0E01A169187" type="GBIF-Dataset">ef601d63-c211-4a45-a530-88385b13d87e</mods:identifier>
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<mods:identifier id="56B6703B8EF8854AF0C6889E85C68501" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8793E89A64EF9E6904885E6A622AE1B7" type="Zenodo-Dep">236658</mods:identifier>
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<mods:identifier id="258B37FD29EF77571CB227B07312D694" type="ZooBank">428F1742-286A-4550-8EF7-CD22686C664F</mods:identifier>
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<treatment id="4268D77B8049FFEDFF36147BFAD6FD0C" ID-DOI="http://doi.org/10.5281/zenodo.6103218" ID-GBIF-Taxon="119613546" ID-Zenodo-Dep="6103218" LSID="urn:lsid:plazi:treatment:4268D77B8049FFEDFF36147BFAD6FD0C" httpUri="http://treatment.plazi.org/id/4268D77B8049FFEDFF36147BFAD6FD0C" lastPageId="13" lastPageNumber="504" pageId="12" pageNumber="503">
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<subSubSection id="82DB35E68049FFECFF36147BFE89FD43" pageId="12" pageNumber="503" type="nomenclature">
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<paragraph id="CA7E666D8049FFECFF36147BFC0DFDA2" blockId="12.[151,938,583,643]" box="[151,938,583,610]" pageId="12" pageNumber="503">
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<heading id="9136D1018049FFECFF36147BFC0DFDA2" bold="true" box="[151,938,583,610]" fontSize="11" level="1" pageId="12" pageNumber="503" reason="1">
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<emphasis id="F8B5BA7F8049FFECFF36147BFC0DFDA2" bold="true" box="[151,938,583,610]" pageId="12" pageNumber="503">
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<taxonomicName id="0DC11DEE8049FFECFF36147BFD6AFDA2" authority="Boucher, 2015" authorityName="Boucher" authorityYear="2015" box="[151,717,583,610]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="doesburgi" status="comb. nov." subGenus="Pertinax">
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<emphasis id="F8B5BA7F8049FFECFF36147BFEA6FDA1" bold="true" box="[151,257,583,609]" italics="true" pageId="12" pageNumber="503">Passalus</emphasis>
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||
(
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<emphasis id="F8B5BA7F8049FFECFEB3147BFEDDFDA1" bold="true" box="[274,378,583,609]" italics="true" pageId="12" pageNumber="503">Pertinax</emphasis>
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)
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<emphasis id="F8B5BA7F8049FFECFE2A147BFDA5FDA1" bold="true" box="[395,514,583,609]" italics="true" pageId="12" pageNumber="503">doesburgi</emphasis>
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(Boucher, 2015)
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</taxonomicName>
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<taxonomicNameLabel id="E38607048049FFECFD751474FC0DFDA2" box="[724,938,584,610]" pageId="12" pageNumber="503">new combination</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="CA7E666D8049FFECFF361456FE89FD43" blockId="12.[151,938,583,643]" box="[151,302,618,643]" pageId="12" pageNumber="503">
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(
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<figureCitation id="52FA7AE88049FFECFF3E1456FF5BFD43" box="[159,252,618,643]" captionStart="FIGURES 14 – 17" captionStartId="11.[151,264,1738,1761]" captionTargetBox="[151,1435,373,1717]" captionTargetId="figure@11.[151,1435,373,1717]" captionTargetPageId="11" captionText="FIGURES 14 – 17. Head and pronotum, dorsal view. 14. Passalus pertyi. 15. Passalus doesburgi. 16. Paxillus jamaicensis. 17. Paxillus pentaphyllus. Scale bars: 1 mm." httpUri="https://zenodo.org/record/236663/files/figure.png" pageId="12" pageNumber="503">Figs. 15</figureCitation>
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,
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<figureCitation id="52FA7AE88049FFECFEA91457FE82FD43" box="[264,293,619,643]" captionStart="FIGURE 18" captionStartId="12.[151,250,1964,1986]" captionTargetBox="[310,1283,1023,1942]" captionTargetId="figure@12.[299,1287,1020,1943]" captionTargetPageId="12" captionText="FIGURE 18. Passalus doesburgi head and anterior part of pronotum. Scale bar: 1 mm." httpUri="https://zenodo.org/record/236664/files/figure.png" pageId="12" pageNumber="503">18</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="82DB35E68049FFECFF36148EFF5EFC07" pageId="12" pageNumber="503" type="materials_examined">
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<paragraph id="CA7E666D8049FFECFF36148EFD8BFD0B" blockId="12.[151,1436,690,967]" box="[151,556,690,715]" pageId="12" pageNumber="503">
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<emphasis id="F8B5BA7F8049FFECFF36148EFE8AFD0B" bold="true" box="[151,301,690,715]" pageId="12" pageNumber="503">
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<typeStatus id="157AD8CF8049FFECFF36148EFF75FD0B" box="[151,210,690,715]" pageId="12" pageNumber="503">Type</typeStatus>
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locality
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</emphasis>
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.
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<collectingCountry id="B2D626FD8049FFECFE9D148EFD8FFD0B" box="[316,552,690,715]" name="Dominican Republic" pageId="12" pageNumber="503">Dominican Republic</collectingCountry>
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.
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</paragraph>
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<paragraph id="CA7E666D8049FFECFF6614EAFF5EFC07" blockId="12.[151,1436,690,967]" pageId="12" pageNumber="503">
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<emphasis id="F8B5BA7F8049FFECFF6614EAFE15FD2F" bold="true" box="[199,434,726,751]" pageId="12" pageNumber="503">Material examined.</emphasis>
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<specimenCount id="DCC7ADE48049FFECFE1A14EBFDFEFD2F" box="[443,601,726,751]" pageId="12" pageNumber="503" type="generic">10 specimens</specimenCount>
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.
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<collectingCountry id="B2D626FD8049FFECFDC414EBFC39FD2F" box="[613,926,727,751]" name="Dominican Republic" pageId="12" pageNumber="503">DOMINICAN REPUBLIC</collectingCountry>
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: Pedernales, Las Abejas,
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<quantity id="0D39CB888049FFECFB6214EBFAABFD2F" box="[1219,1292,726,751]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="503" unit="km" value="38.0">38 km</quantity>
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NNW Cabo Rojo,
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<geoCoordinate id="AFF500AA8049FFECFF4214C7FEE6FCD3" box="[227,321,763,787]" direction="north" orientation="latitude" pageId="12" pageNumber="503" precision="925" value="18.15">18°09'N</geoCoordinate>
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,
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<geoCoordinate id="AFF500AA8049FFECFEF414C7FE1FFCD3" box="[341,440,763,787]" direction="west" orientation="longitude" pageId="12" pageNumber="503" precision="925" value="-71.63333">71°38'W</geoCoordinate>
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, Robert L. Davidson
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<quantity id="0D39CB888049FFECFD6414C6FCB5FCD3" box="[709,786,762,787]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="12" pageNumber="503" unit="m" value="1160.0">1160m</quantity>
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<date id="BE7F40AD8049FFECFC8114C6FC11FCD3" box="[800,950,762,787]" pageId="12" pageNumber="503" value="1987-07-13">13-July-1987</date>
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//
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<taxonomicName id="0DC11DEE8049FFECFC4314C7FAC2FCD3" box="[994,1381,763,787]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="dominicanus" subGenus="Pertinax">
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||
<emphasis id="F8B5BA7F8049FFECFC4314C7FBE1FCD3" box="[994,1094,763,787]" italics="true" pageId="12" pageNumber="503">Passalus</emphasis>
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||
(
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||
<emphasis id="F8B5BA7F8049FFECFBFF14C7FB19FCD2" box="[1118,1214,763,786]" italics="true" pageId="12" pageNumber="503">Pertinax</emphasis>
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||
)
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||
<emphasis id="F8B5BA7F8049FFECFB7714C7FAC2FCD3" box="[1238,1381,763,787]" italics="true" pageId="12" pageNumber="503">dominicanus</emphasis>
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</taxonomicName>
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van Doesburg det.: J. Schuster 1987 Lacks prominent internal tubercles (1♀
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<collectionCode id="ACD0FEA88049FFECFC1F1523FBB5FCF7" box="[958,1042,799,823]" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="12" pageNumber="503">UVGC</collectionCode>
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); Prov. Pedernales, Pedernales, Las Abejas, Sierra Bahoruco
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<date id="BE7F40AD8049FFECFE15157FFD91FC9B" box="[436,566,835,859]" pageId="12" pageNumber="503" value="1989-06-27">27.VI.1989</date>
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, M.N.H.N., col.: Matusik-Guerrero, 34956,
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<taxonomicName id="0DC11DEE8049FFECFB8A157FFA82FC9B" box="[1067,1317,835,859]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="dominicanus">
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<emphasis id="F8B5BA7F8049FFECFB8A157FFA82FC9B" box="[1067,1317,835,859]" italics="true" pageId="12" pageNumber="503">Passalus dominicanus</emphasis>
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</taxonomicName>
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det. Maes '95 (1♀
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<collectionCode id="ACD0FEA88049FFECFF5C155BFEE2FCBF" box="[253,325,871,895]" pageId="12" pageNumber="503">IEXA</collectionCode>
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); Pedernales, Las Abejas, Sierra Bahoruco
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<date id="BE7F40AD8049FFECFCE6155BFC48FCBF" box="[839,1007,871,895]" pageId="12" pageNumber="503" value="1989-06-18" valueMax="1989-06-28" valueMin="1989-06-18">18-28.VI.1989</date>
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, leg. Matusik //
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<taxonomicName id="0DC11DEE8049FFECFB18155AFE81FC63" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="dominicanus" subGenus="Pertinax">
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<emphasis id="F8B5BA7F8049FFECFB18155AFABAFCBE" box="[1209,1309,870,894]" italics="true" pageId="12" pageNumber="503">Passalus</emphasis>
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(
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<emphasis id="F8B5BA7F8049FFECFA93155BFA35FCBE" box="[1330,1426,871,894]" italics="true" pageId="12" pageNumber="503">Pertinax</emphasis>
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)
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<emphasis id="F8B5BA7F8049FFECFF3615B7FE81FC63" box="[151,294,907,931]" italics="true" pageId="12" pageNumber="503">dominicanus</emphasis>
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</taxonomicName>
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det.
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<bibRefCitation id="AE501B9C8049FFECFEC015B7FDE5FC63" author="Gillogly" box="[353,578,906,931]" pageId="12" pageNumber="503" refString="Ivie, M. & Gillogly, A. (1998) Passalidae of the West Indies. Available from: http: // wibeetles. org / polyphaga / scarabaeiformia / scarabaeoidea / passalidae. html (accessed 15 March 2015)" type="journal article" year="1998">A.R. Gillogly, 1998</bibRefCitation>
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||
//
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<taxonomicName id="0DC11DEE8049FFECFDC215B7FB93FC63" authority="Boucher" authorityName="Boucher" box="[611,1076,906,931]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="doesburgi" subGenus="Pertinax">
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<emphasis id="F8B5BA7F8049FFECFDC215B7FD60FC63" box="[611,711,907,931]" italics="true" pageId="12" pageNumber="503">Passalus</emphasis>
|
||
(
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<emphasis id="F8B5BA7F8049FFECFD7915B7FC9FFC62" box="[728,824,907,930]" italics="true" pageId="12" pageNumber="503">Pertinax</emphasis>
|
||
)
|
||
<emphasis id="F8B5BA7F8049FFECFCEA15B7FC1DFC63" box="[843,954,907,931]" italics="true" pageId="12" pageNumber="503">doesburgi</emphasis>
|
||
(Boucher)
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||
</taxonomicName>
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||
det.: Jiménez-Ferbans, 2015 (8
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<collectionCode id="ACD0FEA88049FFECFF361593FF4AFC07" LSID="urn:lsid:biocol.org:col:34795" box="[151,237,943,967]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="503">FMNH</collectionCode>
|
||
).
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</paragraph>
|
||
</subSubSection>
|
||
<caption id="9EBE36E58049FFECFF361190FBACF802" box="[151,1035,1964,1986]" httpUri="https://zenodo.org/record/236664/files/figure.png" pageId="12" pageNumber="503" targetBox="[310,1283,1023,1942]" targetPageId="12">
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||
<paragraph id="CA7E666D8049FFECFF361190FBACF802" blockId="12.[151,1035,1964,1986]" box="[151,1035,1964,1986]" pageId="12" pageNumber="503">
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||
<emphasis id="F8B5BA7F8049FFECFF361190FEB8F801" bold="true" box="[151,287,1964,1986]" pageId="12" pageNumber="503">FIGURE 18.</emphasis>
|
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<taxonomicName id="0DC11DEE8049FFECFE841190FE4CF802" box="[293,491,1964,1986]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="503" phylum="Arthropoda" rank="species" species="doesburgi">
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<emphasis id="F8B5BA7F8049FFECFE841190FE4CF802" box="[293,491,1964,1986]" italics="true" pageId="12" pageNumber="503">Passalus doesburgi</emphasis>
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</taxonomicName>
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head and anterior part of pronotum. Scale bar: 1 mm.
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</paragraph>
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</caption>
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<subSubSection id="82DB35E68048FFEDFF6616ABFE5DFE6D" pageId="13" pageNumber="504" type="diagnosis">
|
||
<paragraph id="CA7E666D8048FFEDFF6616ABFE5DFE6D" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="504">
|
||
<emphasis id="F8B5BA7F8048FFEDFF6616ABFE9DFF70" bold="true" box="[199,314,151,176]" pageId="13" pageNumber="504">Diagnosis</emphasis>
|
||
.
|
||
<quantity id="0D39CB888048FFEDFEE916ABFE4CFF6F" box="[328,491,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="2.84" metricValueMin="2.66" pageId="13" pageNumber="504" unit="mm" value="27.5" valueMax="28.4" valueMin="26.6">26.6–28.4 mm</quantity>
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mm total length. Clypeus hidden below frons, slightly visible dorsally, with anterior angles well developed, larger than mediofrontal tubercles. Frons border with notch, without secondary mediofrontal tubercles. Frontal fossae glabrous. Mediofrontal tubercles developed, not joined to interior tubercles. Internal tubercles smaller than mediofrontal tubercles. Central tubercle of MFS with apex not free; basolateral tubercles weak. Antennal club trilamellate, with short lamellae. Apex of lacinia bidentate. Median basal region of mentum protruding and pubescent posteriorly. Prosternal process rhomboidal, acute. Mesosternum glabrous without scars. Anterolateral metasternum and lateral groove glabrous, disc delimited by punctures in posterior area. Humeri and epipleura glabrous.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="82DB35E68048FFEDFF66178BFEE0FE34" pageId="13" pageNumber="504" type="distribution">
|
||
<paragraph id="CA7E666D8048FFEDFF66178BFEE0FE34" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="504">
|
||
<emphasis id="F8B5BA7F8048FFEDFF66178BFEC7FE10" bold="true" box="[199,352,439,464]" pageId="13" pageNumber="504">Distribution.</emphasis>
|
||
<collectingCountry id="B2D626FD8048FFEDFEC9178BFDF3FE10" box="[360,596,439,464]" name="Dominican Republic" pageId="13" pageNumber="504">Dominican Republic</collectingCountry>
|
||
, from mid-altitudes of Sierra Bahoruco, an area dominated by cloud forest (
|
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<bibRefCitation id="AE501B9C8048FFEDFF3F17E0FE9CFE34" author="Camara" box="[158,315,476,500]" pageId="13" pageNumber="504" refString="Camara, R. (2004) Escalonamiento bioclimatico, regimenes ecodinamicos y formaciones vegetales de la isla de La Espanola en Republica Dominicana. In: Pinto, J. (Ed.), Estudios en Biogeografia. Aster, Alarcon, Spain, pp. 39 - 55." type="book chapter" year="2004">Cámara 2004</bibRefCitation>
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||
).
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82DB35E68048FFEDFF6617C3FAD6FD0C" pageId="13" pageNumber="504" type="discussion">
|
||
<paragraph id="CA7E666D8048FFEDFF6617C3FAD6FD0C" blockId="13.[151,1437,151,717]" pageId="13" pageNumber="504">
|
||
<emphasis id="F8B5BA7F8048FFEDFF6617C3FEEFFDD8" bold="true" box="[199,328,511,536]" pageId="13" pageNumber="504">Comments</emphasis>
|
||
.
|
||
<bibRefCitation id="AE501B9C8048FFEDFEF617C3FDB0FDD8" author="Boucher" box="[343,535,511,536]" pageId="13" pageNumber="504" refString="Boucher, S. (2015 b) Homology versus homoplasy, a case of comprehensive study of male genitalia to reconstruct phylogeny of Passalidae of the Greater Antilles (Coleoptera, Scarabaeoidea). Coleopteres, 21, 1 - 17." type="journal article" year="2015" yearSuffix="b">Boucher (2015b)</bibRefCitation>
|
||
described the genus
|
||
<taxonomicName id="0DC11DEE8048FFEDFCA9143CFC43FDD8" authority="Boucher" authorityName="Boucher" box="[776,996,511,536]" class="Insecta" family="Passalidae" genus="Antillanax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F8B5BA7F8048FFEDFCA9143CFCDAFDD8" box="[776,893,512,536]" italics="true" pageId="13" pageNumber="504">Antillanax</emphasis>
|
||
Boucher
|
||
</taxonomicName>
|
||
for the species
|
||
<taxonomicName id="0DC11DEE8048FFEDFB3B143CFA32FDD8" box="[1178,1429,512,536]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="species" species="dominicanus">
|
||
<emphasis id="F8B5BA7F8048FFEDFB3B143CFA32FDD8" box="[1178,1429,512,536]" italics="true" pageId="13" pageNumber="504">Passalus dominicanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="F8B5BA7F8048FFEDFF361419FF51FDFC" box="[151,246,549,572]" italics="true" pageId="13" pageNumber="504">P. p e r t y i</emphasis>
|
||
, and
|
||
<taxonomicName id="0DC11DEE8048FFEDFEE51419FE72FDFC" box="[324,469,548,572]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="species" species="doesburgi">
|
||
<emphasis id="F8B5BA7F8048FFEDFEE51419FEF5FDFC" box="[324,338,549,572]" italics="true" pageId="13" pageNumber="504">P</emphasis>
|
||
.
|
||
<emphasis id="F8B5BA7F8048FFEDFEC71418FE72FDFC" box="[358,469,548,572]" italics="true" pageId="13" pageNumber="504">doesburgi</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<bibRefCitation id="AE501B9C8048FFEDFDC31418FC80FDFD" author="Boucher" box="[610,807,548,573]" pageId="13" pageNumber="504" refString="Boucher, S. (2015 b) Homology versus homoplasy, a case of comprehensive study of male genitalia to reconstruct phylogeny of Passalidae of the Greater Antilles (Coleoptera, Scarabaeoidea). Coleopteres, 21, 1 - 17." type="journal article" year="2015" yearSuffix="b">Boucher (2015b)</bibRefCitation>
|
||
stated “Following the phylogenetic advances on the Passalini,
|
||
<taxonomicName id="0DC11DEE8048FFEDFEAC1474FE25FDA0" box="[269,386,584,608]" class="Insecta" family="Passalidae" genus="Antillanax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F8B5BA7F8048FFEDFEAC1474FE25FDA0" box="[269,386,584,608]" italics="true" pageId="13" pageNumber="504">Antillanax</emphasis>
|
||
</taxonomicName>
|
||
is a possible relict of the “
|
||
<taxonomicName id="0DC11DEE8048FFEDFD711474FC97FD9F" box="[720,816,584,607]" class="Insecta" family="Passalidae" genus="Pertinax" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F8B5BA7F8048FFEDFD711474FC97FD9F" box="[720,816,584,607]" italics="true" pageId="13" pageNumber="504">Pertinax</emphasis>
|
||
</taxonomicName>
|
||
lineage” in the Greater Antilles”. This is somewhat congruent with the cladogram of
|
||
<bibRefCitation id="AE501B9C8048FFEDFDB71450FC51FD44" author="Jimenez-Ferbans" box="[534,1014,620,645]" pageId="13" pageNumber="504" refString="Jimenez-Ferbans, L. & Reyes-Castillo, P. (2014) Phylogeny, biogeography and description of Ameripassalus, a new Mesoamerican genus of Passalidae (Coleoptera). Invertebrate Systematics, 28, 124 - 144. http: // dx. doi. org / 10.1071 / IS 13009" type="journal article" year="2014">Jiménez-Ferbans & Reyes-Castillo (2014)</bibRefCitation>
|
||
, where
|
||
<taxonomicName id="0DC11DEE8048FFEDFBF01450FAEAFD44" box="[1105,1357,620,644]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="species" species="dominicanus">
|
||
<emphasis id="F8B5BA7F8048FFEDFBF01450FAEAFD44" box="[1105,1357,620,644]" italics="true" pageId="13" pageNumber="504">Passalus dominicanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0DC11DEE8048FFEDFA261451FF7FFD67" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="species" species="pertyi">
|
||
<emphasis id="F8B5BA7F8048FFEDFA261451FF7FFD67" italics="true" pageId="13" pageNumber="504">P. pertyi</emphasis>
|
||
</taxonomicName>
|
||
are nested within
|
||
<taxonomicName id="0DC11DEE8048FFEDFE1014ACFD34FD68" authority="(Pertinax)" baseAuthorityName="Pertinax" box="[433,659,656,680]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F8B5BA7F8048FFEDFE1014ACFDB2FD68" box="[433,533,656,680]" italics="true" pageId="13" pageNumber="504">Passalus</emphasis>
|
||
(
|
||
<emphasis id="F8B5BA7F8048FFEDFD8614ACFD20FD67" box="[551,647,656,679]" italics="true" pageId="13" pageNumber="504">Pertinax</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
, and related to
|
||
<taxonomicName id="0DC11DEE8048FFEDFCEB14ADFC68FD68" box="[842,975,656,680]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="F8B5BA7F8048FFEDFCEB14ADFCFFFD68" box="[842,856,657,680]" italics="true" pageId="13" pageNumber="504">P</emphasis>
|
||
.
|
||
<emphasis id="F8B5BA7F8048FFEDFCC914ADFC68FD68" box="[872,975,657,680]" italics="true" pageId="13" pageNumber="504">convexus</emphasis>
|
||
</taxonomicName>
|
||
. Since the genus
|
||
<taxonomicName id="0DC11DEE8048FFEDFB3E14ACFAB3FD68" box="[1183,1300,656,680]" class="Insecta" family="Passalidae" genus="Antillanax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F8B5BA7F8048FFEDFB3E14ACFAB3FD68" box="[1183,1300,656,680]" italics="true" pageId="13" pageNumber="504">Antillanax</emphasis>
|
||
</taxonomicName>
|
||
renders the subgenus
|
||
<taxonomicName id="0DC11DEE8048FFEDFEA71488FE44FD0C" authority="(Pertinax)" baseAuthorityName="Pertinax" box="[262,483,692,716]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F8B5BA7F8048FFEDFEA71488FECDFD0C" box="[262,362,692,716]" italics="true" pageId="13" pageNumber="504">Passalus</emphasis>
|
||
(
|
||
<emphasis id="F8B5BA7F8048FFEDFEDB1489FE7DFD0C" box="[378,474,693,716]" italics="true" pageId="13" pageNumber="504">Pertinax</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
paraphyletic, we hereby place
|
||
<taxonomicName id="0DC11DEE8048FFEDFCE11488FC12FD0C" box="[832,949,692,716]" class="Insecta" family="Passalidae" genus="Antillanax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F8B5BA7F8048FFEDFCE11488FC12FD0C" box="[832,949,692,716]" italics="true" pageId="13" pageNumber="504">Antillanax</emphasis>
|
||
</taxonomicName>
|
||
in synonymy with
|
||
<taxonomicName id="0DC11DEE8048FFEDFB2C1488FACBFD0C" authority="(Pertinax)" baseAuthorityName="Pertinax" box="[1165,1388,692,716]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F8B5BA7F8048FFEDFB2C1488FB56FD0C" box="[1165,1265,692,716]" italics="true" pageId="13" pageNumber="504">Passalus</emphasis>
|
||
(
|
||
<emphasis id="F8B5BA7F8048FFEDFAA01489FAC6FD0C" box="[1281,1377,693,716]" italics="true" pageId="13" pageNumber="504">Pertinax</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |