treatments-xml/data/C2/95/57/C29557916297C6AA559A00348F413623.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZBK="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. &amp; Bergsten, Johannes" docDate="2015" docId="C29557916297C6AA559A00348F413623" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus enigmaticus Bistroem, Nilsson &amp; Bergsten, 2015, sp. n." docType="treatment" docUuid="8A25143F-4849-495B-AF13-8820977C2934" docUuidSource="ZooBank" docVersion="5" lastPageNumber="129" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="128" updateTime="1668162284477" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bistroem, Olof</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nilsson, Anders N.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bergsten, Johannes</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>542</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>379</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.542.5975</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-542-1</mods:identifier>
<mods:identifier type="ZBK">026407877355425BAB10BF1674510F12</mods:identifier>
<mods:identifier type="ZooBank">026407877355425BAB10BF1674510F12</mods:identifier>
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<treatment ID-GBIF-Taxon="127862720" LSID="urn:lsid:zoobank.org:act:8A25143F-4849-495B-AF13-8820977C2934" httpUri="http://treatment.plazi.org/id/C29557916297C6AA559A00348F413623" lastPageId="128" lastPageNumber="129" pageId="127" pageNumber="128">
<subSubSection pageId="127" pageNumber="128" type="multiple">
<paragraph pageId="127" pageNumber="128">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
</subSubSection>
<subSubSection pageId="127" pageNumber="128" type="nomenclature">
<paragraph pageId="127" pageNumber="128">
<taxonomicName LSID="http://zoobank.org/8A25143F-4849-495B-AF13-8820977C2934" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus enigmaticus" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="enigmaticus">Laccophilus enigmaticus</taxonomicName>
<taxonomicNameLabel pageId="127" pageNumber="128">sp. n.</taxonomicNameLabel>
Figs 94-95, 285, 439, 555
</paragraph>
</subSubSection>
<subSubSection pageId="127" pageNumber="128" type="type locality">
<paragraph pageId="127" pageNumber="128">Type locality.</paragraph>
<paragraph pageId="127" pageNumber="128">Nigeria: Zaria province, Zaria.</paragraph>
</subSubSection>
<subSubSection pageId="127" pageNumber="128" type="type material">
<paragraph pageId="127" pageNumber="128">Type material</paragraph>
<paragraph pageId="127" pageNumber="128">(9 exs.): Holotype, male: &quot;Nigeria Zaria Prov., Zaria 5-6.III. 1949 B. Malkin / B. Malkin Coll. BMNH(E) 1956-234&quot; (BMNH). - Paratypes: Same data as holotype (3 exs. BMNH, 1 ex. MZH; habitus in Fig. 439); &quot;Sudan Bahr el Ghazal Wau 19.2. 1963 Linnavuori&quot; (1 ex. MZH); &quot;Sudan (Equatoria) Nzara 22.4. 1986 leg. Wewalka (Z11-13)&quot; (2 exs. CGW, 1 ex. MZH).</paragraph>
</subSubSection>
<subSubSection pageId="128" pageNumber="129" type="diagnosis">
<paragraph pageId="128" pageNumber="129">
<pageBreakToken pageId="128" pageNumber="129" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="128" pageNumber="129">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus enigmaticus" order="Coleoptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="enigmaticus">Laccophilus enigmaticus</taxonomicName>
is especially characterized by comparatively big sized body, peculiar, uniform elytral colour pattern in combination with the penis, which is different in comparison to all other recognized species in Africa; inner outline curvature medially somewhat enlarged, forming a distinct but smooth expansion of penis. Apex of penis broadly but rather indistinctly hooked.
</paragraph>
</subSubSection>
<subSubSection pageId="128" pageNumber="129" type="description">
<paragraph pageId="128" pageNumber="129">Description.</paragraph>
<paragraph pageId="128" pageNumber="129">Body length 4.3-4.6 mm, width 2.3-2.5 mm. Dorsal, colour pattern of body almost uniform, exhibits only minor variation (Fig. 439).</paragraph>
<paragraph pageId="128" pageNumber="129">Head: Pale ferrugineous. Slightly mat, microsculptured. Reticulation double; large meshes distinctly more strongly developed than small meshes. Large meshes may contain 2-5 small meshes. Impunctate, except at eyes; with fine, irregular punctures, the area of which extends a short distance towards middle of head-disc.</paragraph>
<paragraph pageId="128" pageNumber="129">Pronotum: Pale ferrugineous, frontally and basally in middle with narrow, often indistinct, ferrugineous to dark ferrugineous marking. Slightly mat, microsculptured. Reticulation double. Large meshes clearly, more strongly developed than small meshes. Large meshes may contain 2-5 small meshes. Almost impunctate, except frontally and discally; with fine, sparse and irregular punctures.</paragraph>
<paragraph pageId="128" pageNumber="129">Elytra: Pale ferrugineous, with dense, almost uniform, dark brown to dark ferrugineous irrorations (Fig. 439). Rather shiny, although microsculptured. Reticulation double. Large meshes in part quite indistinct. When discernible large meshes may contain 2-5 small meshes. Discally with fine, irregular row of punctures. Scattered, sparse punctures may also be discerned outside discal row-area. Pre-apical, lateral row of punctures fine, provided with some hairs.</paragraph>
<paragraph pageId="128" pageNumber="129">Ventral aspect: Pale ferrugineous. Almost impunctate; apical ventrite with some scattered, rather fine punctures. Apical ventrite asymmetric having a small, sharp knob on one side (Fig. 92). Rather shiny, very finely and in part indistinctly microsculptured. Ventrites with very fine, slightly curved striae. Metacoxal plates with shallow, especially in posterior half, reduced and indistinct, furrows, which are almost transversely located. Prosternal process quite slender, posteriorly slightly extended, apically pointed.</paragraph>
<paragraph pageId="128" pageNumber="129">Legs: Pale ferrugineous. Pro- and mesotarsus slightly enlarged, with distinct suckers.</paragraph>
<paragraph pageId="128" pageNumber="129">Male genitalia: Penis in lateral aspect peculiar, being quite slender and curved, provided with a broad extension on inner outline of penis. Extreme apex broadly but moderately hooked (Fig. 285).</paragraph>
<paragraph pageId="128" pageNumber="129">Female: Pro- and mesotarsus slender. Apical ventrite as in Fig. 93.</paragraph>
</subSubSection>
<subSubSection pageId="128" pageNumber="129" type="etymology">
<paragraph pageId="128" pageNumber="129">Etymology.</paragraph>
<paragraph pageId="128" pageNumber="129">
The species name enigmaticus is a Latin adjective meaning
<normalizedToken originalValue="“enigmatic”">&quot;enigmatic&quot;</normalizedToken>
and refers to the situation that, externally,
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus enigmaticus" order="Coleoptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="enigmaticus">Laccophilus enigmaticus</taxonomicName>
resembles of some other African species but regarding male genitalia, it deviates strongly from the corresponding features in those species.
</paragraph>
</subSubSection>
<subSubSection pageId="128" pageNumber="129" type="distribution">
<paragraph pageId="128" pageNumber="129">Distribution.</paragraph>
<paragraph pageId="128" pageNumber="129">Nigeria, Sudan (Fig. 555).</paragraph>
</subSubSection>
<subSubSection pageId="128" pageNumber="129" type="collecting circumstances">
<paragraph pageId="128" pageNumber="129">Collecting circumstances.</paragraph>
<paragraph pageId="128" pageNumber="129">Unknown, no information available.</paragraph>
</subSubSection>
</treatment>
</document>