375 lines
43 KiB
XML
375 lines
43 KiB
XML
<document id="3FF3E704C8BCF8B6C65C73D9D297EBBC" ID-DOI="10.5281/zenodo.198925" ID-GBIF-Dataset="b168cdce-1f22-42a6-ace6-f5ae1856cde3" ID-ISSN="1175-5326" ID-Zenodo-Dep="198925" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460317550632" checkinUser="plazi" docAuthor="Thandar, Ahmed S., Zettler, Michael L. & Arumugam, Preyan" docDate="2010" docId="5E0B315DFFD03E0EFF287AAE28DCF95C" docLanguage="en" docName="zt02655p024.pdf" docOrigin="Zootaxa 2655" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ocnus placominutus Thandar, Zettler & Arumugam, 2010, n. sp." docType="treatment" docVersion="10" lastPageNumber="4" masterDocId="A2324925FFD13E0DFFBF7E0B282DFFF0" masterDocTitle="Additions to the sea cucumber fauna of Namibia and Angola, with descriptions of new taxa (Echinodermata: Holothuroidea)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="2" updateTime="1698255494658" updateUser="plazi">
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<mods:mods id="C32B3EB80B349A8ACD36828B05047059" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1579A80205B1E643E3F424B453D06F5A">
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<mods:title id="59A04DE1CC1242030833D54E2B8BD02F">Additions to the sea cucumber fauna of Namibia and Angola, with descriptions of new taxa (Echinodermata: Holothuroidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="11A13B50598092A0F4A79C11E4C3233C" type="personal">
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<mods:role id="431BF666B199F2045E64BDD645697D8A">
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<mods:roleTerm id="38BD16942FA65D748B2826943323795B">Author</mods:roleTerm>
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<mods:namePart id="11EB1A0D2B89FB1E294AC0254EEE4EFF">Thandar, Ahmed S.</mods:namePart>
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</mods:name>
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<mods:name id="9AE16EABA74B6099EF605D51318127CC" type="personal">
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<mods:role id="19011AE26B7116288EC49CBDA7538326">
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<mods:roleTerm id="D016B396D3866CF7F1812BCB9A35C3DC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="06B053E38F87A0A291501D824DE0C268">Zettler, Michael L.</mods:namePart>
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</mods:name>
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<mods:name id="38EE1C29CA0F26B5F2B49E0233F48EC6" type="personal">
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<mods:role id="FA4D55F0767A0100F45E272CE33662E8">
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<mods:roleTerm id="C065BA790564E1DEB0597F79AD1B2511">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="71978201612CF3DD21BC382CE3DCBF8F">Arumugam, Preyan</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1C19D219DF67205C9ED7144A22B30196">text</mods:typeOfResource>
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<mods:titleInfo id="C7D3690F296ED261E10A7536A8240521">
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<mods:title id="599A1E9204A6A787D27DEAD43C348951">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="39674C9526C1BA2E293B57BF649C73BF">
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<mods:date id="C8549B03C16CD75ECD20F1A2EBF4F3BD">2010</mods:date>
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<mods:detail id="09E7029C6E99944210AD4B401B2A18AA" type="volume">
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<mods:number id="C8622AA03F1D0062A2F3D6898C99D90E">2655</mods:number>
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<mods:start id="08F65E32EA6030D0628937A3A41709FF">1</mods:start>
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<mods:end id="B0A1C3ECE64E8C2D98CF0DB091D38447">24</mods:end>
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<mods:classification id="B49FD269D99E6ED98FEFBA29BCAB3068">journal article</mods:classification>
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<mods:identifier id="CA931AF5A644813D43223CD2E65679A7" type="DOI">10.5281/zenodo.198925</mods:identifier>
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<mods:identifier id="718212EB1C70800DFDA30402FBD6884B" type="GBIF-Dataset">b168cdce-1f22-42a6-ace6-f5ae1856cde3</mods:identifier>
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<mods:identifier id="22934860B90E561CDD2DFAA928E3E90B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2165A2AC0101A1A80EA36ACE9A7C3404" type="Zenodo-Dep">198925</mods:identifier>
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<treatment id="5E0B315DFFD03E0EFF287AAE28DCF95C" ID-DOI="http://doi.org/10.5281/zenodo.6204523" ID-GBIF-Taxon="119401935" ID-Zenodo-Dep="6204523" LSID="urn:lsid:zoobank.org:act:9F60A61B-9ED0-4743-AC3C-73A39E2B9EEC" httpUri="http://treatment.plazi.org/id/5E0B315DFFD03E0EFF287AAE28DCF95C" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection id="9EB8D3C0FFD03E0CFF287AAE28D7FB12" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="D61D804BFFD03E0CFF287AAE29F7FB4F" blockId="1.[151,474,1189,1250]" box="[151,474,1189,1215]" pageId="1" pageNumber="2">
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<heading id="8D553727FFD03E0CFF287AAE29F7FB4F" bold="true" box="[151,474,1189,1215]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="E4D65C59FFD03E0CFF287AAE29F7FB4F" bold="true" box="[151,474,1189,1215]" pageId="1" pageNumber="2">
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<taxonomicName id="11A2FBC8FFD03E0CFF287AAE29BDFB4F" ID-CoL="48H5P" box="[151,400,1189,1215]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="2" phylum="Echinodermata" rank="species" species="placominutus" status="sp. nov.">
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<emphasis id="E4D65C59FFD03E0CFF287AAE29BDFB4F" bold="true" box="[151,400,1189,1215]" italics="true" pageId="1" pageNumber="2">Ocnus placominutus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FFE5E122FFD03E0CFE277AAE29F7FB4F" box="[408,474,1189,1215]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D61D804BFFD03E0CFF287AC328D7FB12" blockId="1.[151,474,1189,1250]" box="[151,250,1224,1250]" pageId="1" pageNumber="2">
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<figureCitation id="4E999CCEFFD03E0CFF287AC328D7FB12" box="[151,250,1224,1250]" captionStart="FIGURE 1" captionStartId="2.[151,255,1922,1946]" captionTargetBox="[170,1424,185,1895]" captionTargetId="figure@2.[167,1426,178,1910]" captionTargetPageId="2" captionText="FIGURE 1. Ocnus placominutus n. sp. Holotype, SAM A 28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire." httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="1" pageNumber="2">Figure 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9EB8D3C0FFD03E0CFF287B192B29F9C2" pageId="1" pageNumber="2" type="diagnosis">
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<paragraph id="D61D804BFFD03E0CFF287B192B29F9C2" blockId="1.[151,1437,1298,2034]" pageId="1" pageNumber="2">
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<emphasis id="E4D65C59FFD03E0CFF287B192930FADC" bold="true" box="[151,285,1298,1324]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
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A small cucumariid, up to
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<quantity id="115A2DAEFFD03E0CFDCB7B192AE3FADC" box="[628,718,1298,1324]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="2" unit="mm" value="10.0">10 mm</quantity>
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, with slightly U-shaped, angulated body. Mouth and anus surrounded by valves. Tentacles 10, ventral two reduced. Tube feet restricted to radii in single rows, ± 10 per ambulacrum; interambulacra naked. Skin rigid. Radial and interradial plates of calcareous ring anteriorly cleft. Polian vesicle single. Ossicles of body wall of three
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<typeStatus id="09193EE9FFD03E0CFCE97B892BB7FA6C" box="[854,922,1410,1436]" pageId="1" pageNumber="2">types</typeStatus>
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: external layer of minute, flat baskets with spiky margin and a spinulated base; a layer of large, multilayered, knobbed, imbricating plates with minute holes; and a layer of minute, complex plates, similar in structure to large plates. Tube feet deposits table-like rods, frequently knobbed; end-plates stellate. Tentacle deposits include rods of the pedicel
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<typeStatus id="09193EE9FFD03E0CFB657BF92D20F9FC" box="[1242,1293,1522,1548]" pageId="1" pageNumber="2">type</typeStatus>
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and oblong perforated plates, both spinulated or feebly-knobbed.
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</paragraph>
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</subSubSection>
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<subSubSection id="9EB8D3C0FFD03E0CFF797836295CF98C" pageId="1" pageNumber="2" type="etymology">
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<paragraph id="D61D804BFFD03E0CFF797836295CF98C" blockId="1.[151,1437,1298,2034]" pageId="1" pageNumber="2">
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<emphasis id="E4D65C59FFD03E0CFF797836297EF9A7" bold="true" box="[198,339,1597,1623]" pageId="1" pageNumber="2">Etymology.</emphasis>
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The species is so named because of its characteristic minute plates beneath the large, imbricating plates.
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</paragraph>
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</subSubSection>
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<subSubSection id="9EB8D3C0FFD03E0CFF79788C2CD9F937" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph id="D61D804BFFD03E0CFF79788C2CD9F937" blockId="1.[151,1437,1298,2034]" pageId="1" pageNumber="2">
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<emphasis id="E4D65C59FFD03E0CFF79788C2992F951" bold="true" box="[198,447,1671,1697]" pageId="1" pageNumber="2">Material examined.</emphasis>
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<typeStatus id="09193EE9FFD03E0CFE7378832A15F951" box="[460,568,1672,1697]" pageId="1" pageNumber="2" type="holotype">
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<emphasis id="E4D65C59FFD03E0CFE7378832A15F951" box="[460,568,1672,1697]" italics="true" pageId="1" pageNumber="2">Holotype</emphasis>
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</typeStatus>
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(female),
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<collectionCode id="B0B3188EFFD03E0CFD0578832AD7F952" box="[698,762,1672,1698]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="1" pageNumber="2">SAM</collectionCode>
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A28092, AHAB-9, stn. 88, dredge,
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<collectingCountry id="AEB5C0DBFFD03E0CFB7E78832D37F952" box="[1217,1306,1672,1698]" name="Angola" pageId="1" pageNumber="2">Angola</collectingCountry>
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,
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<geoCoordinate id="B396E68CFFD03E0CFA9578832DBBF952" box="[1322,1430,1672,1698]" direction="south" orientation="latitude" pageId="1" pageNumber="2" precision="55" value="-7.235">7.235° S</geoCoordinate>
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,
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<geoCoordinate id="B396E68CFFD03E0CFF2878A62923F937" box="[151,270,1709,1735]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="55" value="12.684">12.684° E</geoCoordinate>
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,
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<date id="A21CA68BFFD03E0CFEA478A629BCF937" box="[283,401,1709,1735]" pageId="1" pageNumber="2" value="2004-05-28">28.v.2004</date>
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, 41 m, Forster & Zettler;
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<typeStatus id="09193EE9FFD03E0CFD0278A62B05F936" box="[701,808,1709,1734]" pageId="1" pageNumber="2" type="paratype">
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<emphasis id="E4D65C59FFD03E0CFD0278A62B05F936" box="[701,808,1709,1734]" italics="true" pageId="1" pageNumber="2">Paratype</emphasis>
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</typeStatus>
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,
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<collectionCode id="B0B3188EFFD03E0CFC8878A62B5AF937" box="[823,887,1709,1735]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="1" pageNumber="2">SAM</collectionCode>
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A28093, same data as
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<typeStatus id="09193EE9FFD03E0CFB3978A62CDDF937" box="[1158,1264,1709,1735]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="9EB8D3C0FFD03E0EFF7978D92BE0FC17" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
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<paragraph id="D61D804BFFD03E0CFF7978D92B1CF802" blockId="1.[151,1437,1298,2034]" pageId="1" pageNumber="2">
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<emphasis id="E4D65C59FFD03E0CFF7978D929DCF91C" bold="true" box="[198,497,1746,1772]" pageId="1" pageNumber="2">
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Description of
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<typeStatus id="09193EE9FFD03E0CFEC078D929C0F91C" box="[383,493,1746,1772]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Specimen small, slightly U-shaped, length along ventral surface
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<quantity id="115A2DAEFFD03E0CFB4E78D92D15F91C" box="[1265,1336,1746,1772]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="2" unit="mm" value="9.0">9 mm</quantity>
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, breadth in mid-body
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<quantity id="115A2DAEFFD03E0CFE9078F329A4F8E2" box="[303,393,1784,1810]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="2" unit="mm" value="1.5">1.5 mm</quantity>
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(
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<figureCitation id="4E999CCEFFD03E0CFE2778F329D7F8E2" box="[408,506,1784,1810]" captionStart="FIGURE 1" captionStartId="2.[151,255,1922,1946]" captionTargetBox="[170,1424,185,1895]" captionTargetId="figure@2.[167,1426,178,1910]" captionTargetPageId="2" captionText="FIGURE 1. Ocnus placominutus n. sp. Holotype, SAM A 28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire." httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="1" pageNumber="2">
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Figure
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<date id="A21CA68BFFD03E0CFE5178F329D7F8E2" box="[494,506,1784,1810]" pageId="1" pageNumber="2">1</date>
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</figureCitation>
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I). Form slightly angular in the radii; preserved colouration off-white, uniform. Mouth anterior, terminal, covered by valves; tentacles retracted; anus also terminal, surrounded by valves; anal teeth absent. Tentacles 10, well-branched, ventral two slightly reduced. Tube feet restricted to radii in single rows, scanty, ± 10 per ambulacrum, longer in the ventral radii, shorter dorsally, most extending rigidly from body wall, longest one ≈
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<quantity id="115A2DAEFFD03E0CFE4479862A4BF857" box="[507,614,1933,1959]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="1" pageNumber="2" unit="mm" value="0.83">0.83 mm</quantity>
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, suckers present, of same diameter as tube foot. Interambulacra naked, papillae/warts/tubercles absent. Skin rough, slightly rigid, packed with ossicles of which the large plates are clearly visible through stereoscope as imbricating scales.
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</paragraph>
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<caption id="82DDD0C3FFD33E0FFF2879892B81F829" httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="2" pageNumber="3" targetBox="[170,1424,185,1895]" targetPageId="2">
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<paragraph id="D61D804BFFD33E0FFF2879892B81F829" blockId="2.[151,1436,1922,2009]" pageId="2" pageNumber="3">
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<emphasis id="E4D65C59FFD33E0FFF2879892A66F86A" bold="true" box="[151,587,1922,1946]" pageId="2" pageNumber="3">
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FIGURE 1.
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<taxonomicName id="11A2FBC8FFD33E0FFE9D79892A2BF86A" box="[290,518,1922,1946]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="placominutus" status="sp. nov.">
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<emphasis id="E4D65C59FFD33E0FFE9D79892A2BF86A" bold="true" box="[290,518,1922,1946]" italics="true" pageId="2" pageNumber="3">Ocnus placominutus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FFE5E122FFD33E0FFDB179892A66F86A" box="[526,587,1922,1946]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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Holotype, SAM A28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire.
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</paragraph>
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</caption>
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<paragraph id="D61D804BFFD23E0EFF797E932A2FFE47" blockId="3.[151,1437,152,1708]" pageId="3" pageNumber="4">
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Calcareous ring simple (
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<figureCitation id="4E999CCEFFD23E0EFE567E932A66FF42" box="[489,587,152,178]" captionStart="FIGURE 1" captionStartId="2.[151,255,1922,1946]" captionTargetBox="[170,1424,185,1895]" captionTargetId="figure@2.[167,1426,178,1910]" captionTargetPageId="2" captionText="FIGURE 1. Ocnus placominutus n. sp. Holotype, SAM A 28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire." httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="3" pageNumber="4">Figure 1</figureCitation>
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H), well calcified, easily fragmented. Radial plates slightly longer than interradial plates, anteriorly bifid, without posterior prolongations; interradial plates also anteriorly bifid, all plates with a posterior notch or concavity, that of the radial plate deeper. Polian vesicle single, short, saccular, slightly to left of ventral mesentery; stone canal thin, straight, remarkably short; madreporite well calcified, exact shape not discernable. Longitudinal muscles as thin single strands, retractors also thin, originating anteriorly. Respiratory trees confined to posterior quarter of body, both poorly branched, especially the left one, right one also much reduced. Gonad as a simple tuft of short tubules attached anteriorly, full of eggs in various stages of development.
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</paragraph>
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<paragraph id="D61D804BFFD23E0EFF797FC928CCFC6C" blockId="3.[151,1437,152,1708]" pageId="3" pageNumber="4">
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Ossicles of body wall characteristically of three
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<typeStatus id="09193EE9FFD23E0EFCB87FC92B64FE2C" box="[775,841,450,476]" pageId="3" pageNumber="4">types</typeStatus>
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: external layer of minute (14–22 µm, mean 17.25 µm, ± 2.57, n = 20) flat baskets with a rather spiky margin and a spinulated base perforated by four or more holes (
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<figureCitation id="4E999CCEFFD23E0EFF5C7C062969FDD7" box="[227,324,525,551]" captionStart="FIGURE 1" captionStartId="2.[151,255,1922,1946]" captionTargetBox="[170,1424,185,1895]" captionTargetId="figure@2.[167,1426,178,1910]" captionTargetPageId="2" captionText="FIGURE 1. Ocnus placominutus n. sp. Holotype, SAM A 28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire." httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="3" pageNumber="4">Figure 1</figureCitation>
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C); followed by a layer of large (224–382 µm, mean 273.95 µm, ± 44.28, n = 10) multilayered, knobbed plates with minute holes (
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<figureCitation id="4E999CCEFFD23E0EFD8C7C392AB9FDBC" box="[563,660,562,588]" captionStart="FIGURE 1" captionStartId="2.[151,255,1922,1946]" captionTargetBox="[170,1424,185,1895]" captionTargetId="figure@2.[167,1426,178,1910]" captionTargetPageId="2" captionText="FIGURE 1. Ocnus placominutus n. sp. Holotype, SAM A 28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire." httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="3" pageNumber="4">Figure 1</figureCitation>
|
||
A); and a layer of minute (65–141 µm, mean 96.45 µm, ± 20.3, n = 10) complex plates, similar in structure and form to the large plates, but extremely diminutive (
|
||
<figureCitation id="4E999CCEFFD23E0EFAAD7C532D58FD82" box="[1298,1397,600,626]" captionStart="FIGURE 1" captionStartId="2.[151,255,1922,1946]" captionTargetBox="[170,1424,185,1895]" captionTargetId="figure@2.[167,1426,178,1910]" captionTargetPageId="2" captionText="FIGURE 1. Ocnus placominutus n. sp. Holotype, SAM A 28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire." httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="3" pageNumber="4">Figure 1</figureCitation>
|
||
B). Tube feet deposits mostly in the form of table-like rods (82–135 µm, mean 114.83 µm, ± 15.01, n = 20) with a curved base, perforated at ends and in the middle which appears in a spire-like extension (
|
||
<figureCitation id="4E999CCEFFD23E0EFB6E7CA92D19FD4C" box="[1233,1332,674,700]" captionStart="FIGURE 1" captionStartId="2.[151,255,1922,1946]" captionTargetBox="[170,1424,185,1895]" captionTargetId="figure@2.[167,1426,178,1910]" captionTargetPageId="2" captionText="FIGURE 1. Ocnus placominutus n. sp. Holotype, SAM A 28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire." httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="3" pageNumber="4">Figure 1</figureCitation>
|
||
F), most rods finely knobbed, rarely smooth but sometimes irregular, plate-like with several holes. End-plates (
|
||
<figureCitation id="4E999CCEFFD23E0EFAF17CC3288FFCF7" captionStart="FIGURE 1" captionStartId="2.[151,255,1922,1946]" captionTargetBox="[170,1424,185,1895]" captionTargetId="figure@2.[167,1426,178,1910]" captionTargetPageId="2" captionText="FIGURE 1. Ocnus placominutus n. sp. Holotype, SAM A 28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire." httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="3" pageNumber="4">Figure 1</figureCitation>
|
||
G) very characteristic, of stellate form, with large medial perforations and smaller ones in the arms, all arms with jagged margins. Tentacle deposits include rods (73–130 µm, mean 95.25 µm, ± 16.12, n = 10) and plates (24–41 µm, mean 36 µm, ± 5.23, n = 10), the former (
|
||
<figureCitation id="4E999CCEFFD23E0EFCB37D332B40FCA2" box="[780,877,824,850]" captionStart="FIGURE 1" captionStartId="2.[151,255,1922,1946]" captionTargetBox="[170,1424,185,1895]" captionTargetId="figure@2.[167,1426,178,1910]" captionTargetPageId="2" captionText="FIGURE 1. Ocnus placominutus n. sp. Holotype, SAM A 28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire." httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="3" pageNumber="4">Figure 1</figureCitation>
|
||
E) of the pedicel
|
||
<typeStatus id="09193EE9FFD23E0EFB867D332C46FCA2" box="[1081,1131,824,850]" pageId="3" pageNumber="4">type</typeStatus>
|
||
and the latter (
|
||
<figureCitation id="4E999CCEFFD23E0EFAA67D332D54FCA2" box="[1305,1401,824,850]" captionStart="FIGURE 1" captionStartId="2.[151,255,1922,1946]" captionTargetBox="[170,1424,185,1895]" captionTargetId="figure@2.[167,1426,178,1910]" captionTargetPageId="2" captionText="FIGURE 1. Ocnus placominutus n. sp. Holotype, SAM A 28092. A. Large knobbed body wall plates; B. minute body wall plates; C. flat baskets from the body wall; D. oblong rods from base of the tentacles; E. plates from base of tentacles; F. rods from tube feet; G. endplate; H. calcareous ring; I. specimen entire." httpUri="https://zenodo.org/record/198926/files/figure.png" pageId="3" pageNumber="4">Figure 1</figureCitation>
|
||
D) as oblong straight or slightly curved and/or branched perforated deposits, the medial perforations being the larger.
|
||
</paragraph>
|
||
<paragraph id="D61D804BFFD23E0EFF797DAC2BE0FC17" blockId="3.[151,1437,152,1708]" pageId="3" pageNumber="4">
|
||
<emphasis id="E4D65C59FFD23E0EFF797DAC29D4FC31" bold="true" box="[198,505,935,961]" pageId="3" pageNumber="4">
|
||
Description of
|
||
<typeStatus id="09193EE9FFD23E0EFE3E7DAC29D9FC31" box="[385,500,935,961]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Length
|
||
<quantity id="115A2DAEFFD23E0EFDE27DA32A8AFC32" box="[605,679,936,962]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="4" unit="mm" value="4.0">4 mm</quantity>
|
||
. Ossicles of body wall similar to those of
|
||
<typeStatus id="09193EE9FFD23E0EFB207DA32D2BFC32" box="[1183,1286,936,962]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
but those of the tube feet frequently without knobs as they are not fully developed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9EB8D3C0FFD23E0EFF797DF929DBFBC2" pageId="3" pageNumber="4" type="distribution">
|
||
<paragraph id="D61D804BFFD23E0EFF797DF92ABFFBFC" blockId="3.[151,1437,152,1708]" box="[198,658,1010,1036]" pageId="3" pageNumber="4">
|
||
<emphasis id="E4D65C59FFD23E0EFF797DF9294AFBFC" bold="true" box="[198,359,1010,1036]" pageId="3" pageNumber="4">Distribution.</emphasis>
|
||
<typeStatus id="09193EE9FFD23E0EFED27DF9298AFBFC" box="[365,423,1010,1036]" pageId="3" pageNumber="4">Type</typeStatus>
|
||
locality only,
|
||
<specimenCount id="C0A44BC2FFD23E0EFDF07DF92ABFFBFC" box="[591,658,1010,1036]" pageId="3" pageNumber="4" type="generic">
|
||
<quantity id="115A2DAEFFD23E0EFDF07DF92AA3FBFC" box="[591,654,1010,1036]" metricMagnitude="1" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="4" unit="m" value="41.0">41 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
<paragraph id="D61D804BFFD23E0EFF797A1C29DBFBC2" blockId="3.[151,1437,152,1708]" box="[198,502,1047,1074]" pageId="3" pageNumber="4">
|
||
<emphasis id="E4D65C59FFD23E0EFF797A1C2903FBC1" bold="true" box="[198,302,1047,1073]" pageId="3" pageNumber="4">Habitat.</emphasis>
|
||
Sandy substrate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9EB8D3C0FFD23E0EFF797A3628DCF95C" pageId="3" pageNumber="4" type="discussion">
|
||
<paragraph id="D61D804BFFD23E0EFF797A362C41FA3C" blockId="3.[151,1437,152,1708]" pageId="3" pageNumber="4">
|
||
<emphasis id="E4D65C59FFD23E0EFF797A36296DFBA7" bold="true" box="[198,320,1085,1111]" pageId="3" pageNumber="4">Remarks.</emphasis>
|
||
The specimens in hand satisfy the diagnosis of
|
||
<taxonomicName id="11A2FBC8FFD23E0EFCC07A362BE7FBA6" box="[895,970,1085,1110]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="E4D65C59FFD23E0EFCC07A362BE7FBA6" box="[895,970,1085,1110]" italics="true" pageId="3" pageNumber="4">Ocnus</emphasis>
|
||
</taxonomicName>
|
||
as amended by
|
||
<bibRefCitation id="B233FDBAFFD23E0EFB317A362D67FBA7" author="Thandar" box="[1166,1354,1085,1111]" pageId="3" pageNumber="4" refString="Thandar, A. S. (2008) Additions to the holothuroid fauna of the southern African temperate faunistic provinces, with descriptions of new species. Zootaxa, 1697, 1 - 57." type="journal article" year="2008">Thandar (2008)</bibRefCitation>
|
||
. In the structure of the calcareous ring and the presence of three
|
||
<typeStatus id="09193EE9FFD23E0EFCF97A692BA9FB8C" box="[838,900,1122,1148]" pageId="3" pageNumber="4">types</typeStatus>
|
||
of body wall ossicles it comes close to most species currently classified in
|
||
<taxonomicName id="11A2FBC8FFD23E0EFE407A832A67FB51" box="[511,586,1160,1185]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="E4D65C59FFD23E0EFE407A832A67FB51" box="[511,586,1160,1185]" italics="true" pageId="3" pageNumber="4">Ocnus</emphasis>
|
||
</taxonomicName>
|
||
, notably
|
||
<taxonomicName id="11A2FBC8FFD23E0EFD047A832BD9FB52" authority="Forbes, 1841" authorityName="Forbes" authorityYear="1841" box="[699,1012,1160,1186]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="brunneus">
|
||
<emphasis id="E4D65C59FFD23E0EFD047A832B63FB51" box="[699,846,1160,1185]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFD047A832AFEFB51" box="[699,723,1160,1185]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. brunneus
|
||
</emphasis>
|
||
<bibRefCitation id="B233FDBAFFD23E0EFCE97A832BD9FB52" author="Forbes" box="[854,1012,1160,1186]" pageId="3" pageNumber="4" refString="Forbes, E. (1841) A History of British Starfishes and other Animals of the Class Echinodermata. John van Voorst, London, 167 pp." type="book" year="1841">Forbes, 1841</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11A2FBC8FFD23E0EFBBE7A832D0DFB52" authority="Forbes, 1841" authorityName="Forbes" authorityYear="1841" box="[1025,1312,1160,1186]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="lacteus">
|
||
<emphasis id="E4D65C59FFD23E0EFBBE7A832C57FB51" box="[1025,1146,1160,1185]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFBBE7A832C34FB51" box="[1025,1049,1160,1185]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. lacteus
|
||
</emphasis>
|
||
<bibRefCitation id="B233FDBAFFD23E0EFB3D7A832D0DFB52" author="Forbes" box="[1154,1312,1160,1186]" pageId="3" pageNumber="4" refString="Forbes, E. (1841) A History of British Starfishes and other Animals of the Class Echinodermata. John van Voorst, London, 167 pp." type="book" year="1841">Forbes, 1841</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11A2FBC8FFD23E0EFA927A832968FB37" authority="Brandt, 1835" authorityName="Brandt" authorityYear="1835" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="planci">
|
||
<emphasis id="E4D65C59FFD23E0EFA927A832DB1FB51" box="[1325,1436,1160,1185]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFA927A832D6BFB51" box="[1325,1350,1160,1185]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. planci
|
||
</emphasis>
|
||
(Brandt, 1835)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11A2FBC8FFD23E0EFEEE7AA62AAFFB37" authority="Dendy, 1897" authorityName="Dendy" authorityYear="1897" box="[337,642,1197,1223]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="calcareus">
|
||
<emphasis id="E4D65C59FFD23E0EFEEE7AA629C8FB36" box="[337,485,1197,1222]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFEEE7AA62944FB36" box="[337,361,1197,1222]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. calcareus
|
||
</emphasis>
|
||
Dendy, 1897
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11A2FBC8FFD23E0EFD317AA62B92FB37" authority="Theel, 1886" authorityName="Theel" authorityYear="1886" box="[654,959,1197,1223]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="capensis">
|
||
<emphasis id="E4D65C59FFD23E0EFD317AA62B3BFB36" box="[654,790,1197,1222]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFD317AA62A8BFB36" box="[654,678,1197,1222]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. capensis
|
||
</emphasis>
|
||
(Théel, 1886)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="11A2FBC8FFD23E0EFC757AA62D6AFB37" authority="Cherbonnier, 1953" authorityName="Cherbonnier" authorityYear="1953" box="[970,1351,1197,1223]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="corbulus">
|
||
<emphasis id="E4D65C59FFD23E0EFC757AA62C79FB36" box="[970,1108,1197,1222]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFC757AA62BCFFB36" box="[970,994,1197,1222]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. corbulus
|
||
</emphasis>
|
||
(Cherbonnier, 1953)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11A2FBC8FFD23E0EFAC07AA629BCFB1C" authority="Thandar, 2008" authorityName="Thandar" authorityYear="2008" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="rowei">
|
||
<emphasis id="E4D65C59FFD23E0EFAC07AA628F4FB1C" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFAC07AA62DBAFB36" box="[1407,1431,1197,1222]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. rowei
|
||
</emphasis>
|
||
<bibRefCitation id="B233FDBAFFD23E0EFF5D7AD929BCFB1C" author="Thandar" box="[226,401,1234,1260]" pageId="3" pageNumber="4" refString="Thandar, A. S. (2008) Additions to the holothuroid fauna of the southern African temperate faunistic provinces, with descriptions of new species. Zootaxa, 1697, 1 - 57." type="journal article" year="2008">Thandar, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
, but not
|
||
<taxonomicName id="11A2FBC8FFD23E0EFE447AD82B36FB1C" authority="Bell, 1883" authorityName="Bell" authorityYear="1883" box="[507,795,1234,1260]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="vicarius">
|
||
<emphasis id="E4D65C59FFD23E0EFE447AD82AADFB1C" box="[507,640,1235,1260]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFE447AD82A39FB1C" box="[507,532,1235,1260]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. vicarius
|
||
</emphasis>
|
||
(Bell, 1883)
|
||
</taxonomicName>
|
||
, which is characterised by cup-like baskets.
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFA8E7AD92D7FFB1C" box="[1329,1362,1234,1260]" name="American Samoa" pageId="3" pageNumber="4">As</collectingCountry>
|
||
far as satisfying the characters of the
|
||
<typeStatus id="09193EE9FFD23E0EFDB77AF32A16FAE2" box="[520,571,1272,1298]" pageId="3" pageNumber="4">type</typeStatus>
|
||
species depends on which species is really the
|
||
<typeStatus id="09193EE9FFD23E0EFBD37AF32CB2FAE2" box="[1132,1183,1272,1298]" pageId="3" pageNumber="4">type</typeStatus>
|
||
species of the genus.
|
||
<bibRefCitation id="B233FDBAFFD23E0EFF287B162911FAC7" author="Forbes" box="[151,316,1309,1335]" pageId="3" pageNumber="4" refString="Forbes, E. (1841) A History of British Starfishes and other Animals of the Class Echinodermata. John van Voorst, London, 167 pp." type="book" year="1841">Forbes (1841)</bibRefCitation>
|
||
chose
|
||
<taxonomicName id="11A2FBC8FFD23E0EFE337B162A31FAC6" box="[396,540,1309,1334]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="brunneus">
|
||
<emphasis id="E4D65C59FFD23E0EFE337B162A31FAC6" box="[396,540,1309,1334]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFE337B162989FAC6" box="[396,420,1309,1334]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. brunneus
|
||
</emphasis>
|
||
</taxonomicName>
|
||
as the
|
||
<typeStatus id="09193EE9FFD23E0EFDD27B162AB2FAC7" box="[621,671,1309,1335]" pageId="3" pageNumber="4">type</typeStatus>
|
||
species of his genus but according to
|
||
<bibRefCitation id="B233FDBAFFD23E0EFBE67B162D23FAC7" author="Panning" box="[1113,1294,1309,1335]" pageId="3" pageNumber="4" refString="Panning, A. (1971) Bemerkungen uber die Holothurien _ Families Cucumaridae (Ordnung Dendrochirota) 6. Teil (Schluss). Mitteilungen aus dem Hamburgischen Zoologischen museum und Institut, 67, 29 - 51." type="journal article" year="1971">Panning (1971)</bibRefCitation>
|
||
<taxonomicName id="11A2FBC8FFD23E0EFAAB7B162DACFAC6" box="[1300,1409,1309,1334]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="brunneus">
|
||
<emphasis id="E4D65C59FFD23E0EFAAB7B162DACFAC6" box="[1300,1409,1309,1334]" italics="true" pageId="3" pageNumber="4">brunneus</emphasis>
|
||
</taxonomicName>
|
||
is a synonym of
|
||
<taxonomicName id="11A2FBC8FFD23E0EFEF37B482A4DFAAC" authority="Brandt, 1835" authorityName="Brandt" authorityYear="1835" box="[332,608,1346,1372]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="planci">
|
||
<emphasis id="E4D65C59FFD23E0EFEF37B4829B6FAAC" box="[332,411,1347,1372]" italics="true" pageId="3" pageNumber="4">planci</emphasis>
|
||
(Brandt, 1835)
|
||
</taxonomicName>
|
||
, while
|
||
<bibRefCitation id="B233FDBAFFD23E0EFD027B492B49FAAC" author="Rowe" box="[701,868,1346,1372]" pageId="3" pageNumber="4" refString="Rowe, F. W. E. (1970) A note on the British species of Cucumarians involving the erection of two new nominal genera. Journal of the Marine Biological Association of the United Kingdom, 50, 683 - 387." type="journal volume" year="1970">Rowe (1970)</bibRefCitation>
|
||
, following
|
||
<bibRefCitation id="B233FDBAFFD23E0EFC497B492CF7FAAC" author="Mortensen" box="[1014,1242,1346,1372]" pageId="3" pageNumber="4" refString="Mortensen, T. (1927) Handbook of the Echinoderms of the British Isles. Oxford University Press, Oxford, 471 pp." type="book" year="1927">Mortensen (1927)</bibRefCitation>
|
||
, suggested that
|
||
<taxonomicName id="11A2FBC8FFD23E0EFF287B63292BFA71" box="[151,262,1384,1409]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="brunneus">
|
||
<emphasis id="E4D65C59FFD23E0EFF287B63292BFA71" box="[151,262,1384,1409]" italics="true" pageId="3" pageNumber="4">brunneus</emphasis>
|
||
</taxonomicName>
|
||
is a synonym of
|
||
<taxonomicName id="11A2FBC8FFD23E0EFE697B632A06FA71" box="[470,555,1384,1409]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="lacteus">
|
||
<emphasis id="E4D65C59FFD23E0EFE697B632A06FA71" box="[470,555,1384,1409]" italics="true" pageId="3" pageNumber="4">lacteus</emphasis>
|
||
</taxonomicName>
|
||
and hence
|
||
<taxonomicName id="11A2FBC8FFD23E0EFD0B7B632B24FA71" box="[692,777,1384,1409]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="lacteus">
|
||
<emphasis id="E4D65C59FFD23E0EFD0B7B632B24FA71" box="[692,777,1384,1409]" italics="true" pageId="3" pageNumber="4">lacteus</emphasis>
|
||
</taxonomicName>
|
||
should replace
|
||
<taxonomicName id="11A2FBC8FFD23E0EFC777B632C1AFA71" box="[968,1079,1384,1409]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="brunneus">
|
||
<emphasis id="E4D65C59FFD23E0EFC777B632C1AFA71" box="[968,1079,1384,1409]" italics="true" pageId="3" pageNumber="4">brunneus</emphasis>
|
||
</taxonomicName>
|
||
as the
|
||
<typeStatus id="09193EE9FFD23E0EFB327B632CEDFA72" box="[1165,1216,1384,1410]" pageId="3" pageNumber="4">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="11A2FBC8FFD23E0EFAF77B632DB9FA71" box="[1352,1428,1384,1409]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="E4D65C59FFD23E0EFAF77B632DB9FA71" box="[1352,1428,1384,1409]" italics="true" pageId="3" pageNumber="4">Ocnus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="B233FDBAFFD23E0EFF287B862970FA57" author="McKenzie" box="[151,349,1421,1447]" pageId="3" pageNumber="4" refString="McKenzie, J. D. (1984) Description of a neotype for the holothurian Ocnus brunneus (Forbes MS in Thompson 1840) from Strangford Lough, Northern Ireland (Holothuroidea: Dendrochirotida). Bulletin of the British Museum of Natural History (Zoology), 47 (5), 265 - 272." type="journal article" year="1984">McKenzie (1984</bibRefCitation>
|
||
&
|
||
<bibRefCitation id="B233FDBAFFD23E0EFE3C7B862992FA57" author="McKenzie" box="[387,447,1421,1447]" pageId="3" pageNumber="4" refString="McKenzie, J. D. (1991) The taxonomy and natural history of north European dendrochirote holothurians (Echinodermata). Journal of Natural History, 25, 123 - 171." type="journal article" year="1991">1991</bibRefCitation>
|
||
) as the last reviser, attempted to separate the above three species and his suggestion should be accepted that
|
||
<taxonomicName id="11A2FBC8FFD23E0EFE0F7BB82A30FA3C" box="[432,541,1459,1484]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="brunneus">
|
||
<emphasis id="E4D65C59FFD23E0EFE0F7BB82A30FA3C" box="[432,541,1459,1484]" italics="true" pageId="3" pageNumber="4">brunneus</emphasis>
|
||
</taxonomicName>
|
||
must be retained as the
|
||
<typeStatus id="09193EE9FFD23E0EFC867BB92B46FA3C" box="[825,875,1458,1484]" pageId="3" pageNumber="4">type</typeStatus>
|
||
species of this genus.
|
||
</paragraph>
|
||
<paragraph id="D61D804BFFD23E0EFF797BD328DCF95C" blockId="3.[151,1437,152,1708]" pageId="3" pageNumber="4">
|
||
In the presence of a single row of tube feet, multilocular plates in the body wall together with other small plates/buttons, the new species agrees with both
|
||
<taxonomicName id="11A2FBC8FFD23E0EFD597BF62B51F9E6" box="[742,892,1533,1558]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="brunneus">
|
||
<emphasis id="E4D65C59FFD23E0EFD597BF62B51F9E6" box="[742,892,1533,1558]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFD597BF62AD2F9E6" box="[742,767,1533,1558]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. brunneus
|
||
</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11A2FBC8FFD23E0EFC077BF62C1FF9E6" box="[952,1074,1533,1558]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="lacteus">
|
||
<emphasis id="E4D65C59FFD23E0EFC077BF62C1FF9E6" box="[952,1074,1533,1558]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFC077BF62BFCF9E6" box="[952,977,1533,1558]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. lacteus
|
||
</emphasis>
|
||
</taxonomicName>
|
||
. However, its large plates are always knobbed and never smooth and its other plates are minute and, unlike those of
|
||
<taxonomicName id="11A2FBC8FFD23E0EFB2D78282D27F9CC" box="[1170,1290,1571,1596]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="lacteus">
|
||
<emphasis id="E4D65C59FFD23E0EFB2D78282D27F9CC" box="[1170,1290,1571,1596]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="AEB5C0DBFFD23E0EFB2D78282C87F9CC" box="[1170,1194,1571,1596]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
|
||
. lacteus
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, never fourholed but complex and multilocular, resembling in form the large plates. The tube feet ossicles are also different from those of the other species of
|
||
<taxonomicName id="11A2FBC8FFD23E0EFD1878662ADEF976" box="[679,755,1645,1670]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="E4D65C59FFD23E0EFD1878662ADEF976" box="[679,755,1645,1670]" italics="true" pageId="3" pageNumber="4">Ocnus</emphasis>
|
||
</taxonomicName>
|
||
but the end-plates resemble those of both
|
||
<taxonomicName id="11A2FBC8FFD23E0EFB4678662D44F976" box="[1273,1385,1645,1670]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="brunneus">
|
||
<emphasis id="E4D65C59FFD23E0EFB4678662D44F976" box="[1273,1385,1645,1670]" italics="true" pageId="3" pageNumber="4">brunneus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="11A2FBC8FFD23E0EFF28789828C7F95C" box="[151,234,1683,1708]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="lacteus">
|
||
<emphasis id="E4D65C59FFD23E0EFF28789828C7F95C" box="[151,234,1683,1708]" italics="true" pageId="3" pageNumber="4">lacteus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |