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<document id="6E49A1D9A605530B22741839E61E1C07" ID-DOI="10.11646/zootaxa.4021.4.6" ID-GBIF-Dataset="dc544c22-fb6e-4504-922c-7ce2ecb41956" ID-ISSN="1175-5326" ID-Zenodo-Dep="245304" ID-ZooBank="40A75A81-9658-4ABD-BDD6-1C9DE3E7D517" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461277520292" checkinUser="plazi" docAuthor="Gorochov, Andrej V., Dawwrueng, Pattarawich &amp; Artchawakom, Taksin" docDate="2015" docId="03CB2265972A3E58FF58B8A2FADAD979" docLanguage="en" docName="zt04021p577.pdf" docOrigin="Zootaxa 4021 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Capnogryllacris (Capnogryllacris) martha Griffini 1914, comb. nov." docType="treatment" docVersion="8" lastPageNumber="572" masterDocId="FFF25A1D972C3E5FFFCFBD50FFB4DF74" masterDocTitle="Study of Gryllacridinae (Orthoptera: Stenopelmatidae) from Thailand and adjacent countries: the genera Ultragryllacris gen. nov. and Capnogryllacris" masterLastPageNumber="577" masterPageNumber="565" pageNumber="571" updateTime="1698613265158" updateUser="plazi">
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<mods:title id="BB06EEBC2D5A1659876EFBA8FD5F7B5F">Study of Gryllacridinae (Orthoptera: Stenopelmatidae) from Thailand and adjacent countries: the genera Ultragryllacris gen. nov. and Capnogryllacris</mods:title>
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<mods:namePart id="8B0DE3CAAEE50ED6AF1060BE9E34C4E1">Gorochov, Andrej V.</mods:namePart>
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<mods:namePart id="7A7DC46944B8B523A1F9381609A806DC">Dawwrueng, Pattarawich</mods:namePart>
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<treatment id="03CB2265972A3E58FF58B8A2FADAD979" ID-DOI="http://doi.org/10.5281/zenodo.6108753" ID-GBIF-Taxon="119639949" ID-Zenodo-Dep="6108753" LSID="urn:lsid:plazi:treatment:03CB2265972A3E58FF58B8A2FADAD979" httpUri="http://treatment.plazi.org/id/03CB2265972A3E58FF58B8A2FADAD979" lastPageId="7" lastPageNumber="572" pageId="6" pageNumber="571">
<subSubSection id="C378C0F8972A3E59FF58B8A2FEE5D95A" pageId="6" pageNumber="571" type="nomenclature">
<paragraph id="8BDD9373972A3E59FF58B8A2FC59D979" blockId="6.[151,1005,1522,1582]" box="[151,1005,1522,1549]" pageId="6" pageNumber="571">
<heading id="D095241F972A3E59FF58B8A2FC59D979" bold="true" box="[151,1005,1522,1549]" fontSize="11" level="1" pageId="6" pageNumber="571" reason="1">
<emphasis id="B9164F61972A3E59FF58B8A2FC59D979" bold="true" box="[151,1005,1522,1549]" pageId="6" pageNumber="571">
<taxonomicName id="4C62E8F0972A3E59FF58B8A2FCEDD979" authority="Griffini, 1914" authorityName="Griffini" authorityYear="1914" box="[151,857,1522,1549]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="martha" status="comb. nov." subGenus="Capnogryllacris">
<emphasis id="B9164F61972A3E59FF58B8A2FEEDD978" bold="true" box="[151,345,1522,1548]" italics="true" pageId="6" pageNumber="571">Capnogryllacris</emphasis>
(
<emphasis id="B9164F61972A3E59FEA5B8A2FD98D978" bold="true" box="[362,556,1522,1548]" italics="true" pageId="6" pageNumber="571">Capnogryllacris</emphasis>
)
<emphasis id="B9164F61972A3E59FDF3B8A2FD27D978" bold="true" box="[572,659,1522,1548]" italics="true" pageId="6" pageNumber="571">martha</emphasis>
(
<bibRefCitation id="EFF3EE82972A3E59FD6CB8A3FCE5D979" author="Griffini" box="[675,849,1523,1549]" pageId="6" pageNumber="571" refString="Griffini A. (1914) Sopra due Gryllacris del Museo di Budapest. Annales Musei Nationalis Hungarici, 12, 249 - 260." type="journal article" year="1914">Griffini, 1914</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A225F21A972A3E59FCA8B8A3FC59D979" box="[871,1005,1523,1549]" pageId="6" pageNumber="571">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD9373972A3E59FF58BB45FEE5D95A" blockId="6.[151,1005,1522,1582]" box="[151,337,1557,1582]" pageId="6" pageNumber="571">(Figs 38, 45, 52)</paragraph>
</subSubSection>
<subSubSection id="C378C0F8972A3E59FF58BB0EFCB4D9E6" pageId="6" pageNumber="571" type="reference_group">
<paragraph id="8BDD9373972A3E59FF58BB0EFE42D907" blockId="6.[151,768,1629,1682]" box="[151,502,1629,1651]" pageId="6" pageNumber="571">
=
<taxonomicName id="4C62E8F0972A3E59FF64BB0DFE42D907" authority="Griffini, 1914" authorityName="Griffini" authorityYear="1914" box="[171,502,1629,1651]" class="Insecta" family="Haglidae" genus="Gryllacris" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="martha">
<emphasis id="B9164F61972A3E59FF64BB0DFED7D906" box="[171,355,1629,1650]" italics="true" pageId="6" pageNumber="571">Gryllacris martha</emphasis>
<bibRefCitation id="EFF3EE82972A3E59FEA6BB0DFE42D907" author="Griffini" box="[361,502,1629,1651]" pageId="6" pageNumber="571" refString="Griffini A. (1914) Sopra due Gryllacris del Museo di Budapest. Annales Musei Nationalis Hungarici, 12, 249 - 260." type="journal article" year="1914">Griffini, 1914</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BDD9373972A3E59FF58BB2DFCB4D9E6" blockId="6.[151,768,1629,1682]" box="[151,768,1660,1682]" pageId="6" pageNumber="571">
=
<taxonomicName id="4C62E8F0972A3E59FF64BB2CFD80D9E6" authority="Karny, 1937" authorityName="Karny" authorityYear="1937" box="[171,564,1660,1682]" class="Insecta" family="Haglidae" genus="Marthogryllacris" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="martha">
<emphasis id="B9164F61972A3E59FF64BB2CFE1DD9E5" box="[171,425,1660,1681]" italics="true" pageId="6" pageNumber="571">Marthogryllacris martha</emphasis>
:
<bibRefCitation id="EFF3EE82972A3E59FE78BB2DFD80D9E6" author="Karny" box="[439,564,1660,1682]" pageId="6" pageNumber="571" refString="Karny, H. H. (1937) Orthoptera Fam. Gryllacrididae subfamiliae omnes. Genera Insectorum, Fascicule 206. A. F. Devrien, Brussels, 317 pp." type="book" year="1937">Karny, 1937</bibRefCitation>
</taxonomicName>
(not
<bibRefCitation id="EFF3EE82972A3E59FDA6BB2CFD4DD9E6" author="Li" box="[617,761,1660,1682]" pageId="6" pageNumber="571" refString="Li, M., Fang, Ya., Liu, X. &amp; Li, K. (2014) Review of the species of the genus Marthogryllacris (Orthoptera, Gryllacrididae, Gryllacridinae). Zootaxa, 3889 (2), 277 - 288. http: // dx. doi. org / 10.11646 / zootaxa. 3889.2.6" type="journal article" year="2014">
Li
<emphasis id="B9164F61972A3E59FD49BB2DFD0DD9E5" box="[646,697,1660,1682]" italics="true" pageId="6" pageNumber="571">et al.</emphasis>
, 2014
</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C378C0F8972A3E58FF58BBEEFDCADB43" lastPageId="7" lastPageNumber="572" pageId="6" pageNumber="571" type="materials_examined">
<paragraph id="8BDD9373972A3E59FF58BBEEFE01D98F" blockId="6.[151,1437,1726,2003]" pageId="6" pageNumber="571">
<emphasis id="B9164F61972A3E59FF58BBEEFED5D9A3" bold="true" box="[151,353,1726,1751]" pageId="6" pageNumber="571">Material studied</emphasis>
.
<typeStatus id="54D92DD1972A3E59FEBCBB90FE66D9AC" box="[371,466,1728,1752]" pageId="6" pageNumber="571" type="neotype">Neotype</typeStatus>
(here designated)—male;
<collectingCountry id="F375D3E3972A3E59FCC7BBEEFCF5D9A3" box="[776,833,1726,1751]" name="Laos" pageId="6" pageNumber="571">
<emphasis id="B9164F61972A3E59FCC7BBEEFCF5D9A3" bold="true" box="[776,833,1726,1751]" pageId="6" pageNumber="571">Laos</emphasis>
</collectingCountry>
, Khammouan Prov., Boulapha Distr., Bandoy Vill.,
<date id="FFDCB5B3972A3E59FF58BBB3FF42D98F" box="[151,246,1763,1787]" pageId="6" pageNumber="571" value="2009-06">VI.2009</date>
, N. Orlov (
<collectionCode id="ED730BB6972A3E59FEBBBBB3FE1CD98F" box="[372,424,1763,1787]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="571">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8BDD9373972A3E58FF08BA58FEFDDF8C" blockId="6.[151,1437,1726,2003]" lastBlockId="7.[151,1437,151,1550]" lastPageId="7" lastPageNumber="572" pageId="6" pageNumber="571">
<emphasis id="B9164F61972A3E59FF08BA58FF48D86B" box="[199,252,1800,1823]" italics="true" pageId="6" pageNumber="571">Note</emphasis>
. This species was described by
<bibRefCitation id="EFF3EE82972A3E59FDBFBA56FCADD86B" author="Griffini" box="[624,793,1798,1823]" pageId="6" pageNumber="571" refString="Griffini A. (1914) Sopra due Gryllacris del Museo di Budapest. Annales Musei Nationalis Hungarici, 12, 249 - 260." type="journal article" year="1914">Griffini (1914)</bibRefCitation>
from a single male with the following geographic data: “
<collectingCountry id="F375D3E3972A3E59FF6EBA7BFF4CD837" box="[161,248,1835,1859]" name="Vietnam" pageId="6" pageNumber="571">Annam</collectingCountry>
,
<collectingCountry id="F375D3E3972A3E59FEC3BA7BFEF3D837" box="[268,327,1835,1859]" name="Laos" pageId="6" pageNumber="571">Laos</collectingCountry>
”.
<bibRefCitation id="EFF3EE82972A3E59FEAABA7BFDB0D837" author="Karny" box="[357,516,1835,1859]" pageId="6" pageNumber="571" refString="Karny, H. H. (1937) Orthoptera Fam. Gryllacrididae subfamiliae omnes. Genera Insectorum, Fascicule 206. A. F. Devrien, Brussels, 317 pp." type="book" year="1937">Karny (1937)</bibRefCitation>
mentioned this species as originating from “
<collectingCountry id="F375D3E3972A3E59FBE0BA7BFBDCD837" box="[1071,1128,1835,1859]" name="Laos" pageId="6" pageNumber="571">Laos</collectingCountry>
(
<collectingCountry id="F375D3E3972A3E59FBB0BA7BFB6DD837" box="[1151,1241,1835,1859]" name="Vietnam" pageId="6" pageNumber="571">Annam</collectingCountry>
)”. However, the
<taxonomicName id="4C62E8F0972A3E59FF58BA1EFEA4D813" box="[151,272,1870,1895]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
Species File (Eades
<emphasis id="B9164F61972A3E59FDCFBA00FD8ED813" box="[512,570,1871,1895]" italics="true" pageId="6" pageNumber="571">et al.</emphasis>
, 2015) gives another
<typeStatus id="54D92DD1972A3E59FCFABA00FCD1D81C" box="[821,869,1872,1896]" pageId="6" pageNumber="571">type</typeStatus>
locality for this species: “
<collectingCountry id="F375D3E3972A3E59FB5EBA1EFB41D813" box="[1169,1269,1870,1895]" name="Vietnam" pageId="6" pageNumber="571">Vietnam</collectingCountry>
,
<collectingCountry id="F375D3E3972A3E59FACDBA1FFAEED813" box="[1282,1370,1871,1895]" name="Vietnam" pageId="6" pageNumber="571">Annam</collectingCountry>
”. The latter indication is most probably erroneous, because in some old maps of
<collectingCountry id="F375D3E3972A3E59FC39BA22FB74D8FF" box="[1014,1216,1906,1931]" name="Vietnam" pageId="6" pageNumber="571">French Indochina</collectingCountry>
, the central part of recent
<collectingCountry id="F375D3E3972A3E59FF2CBAC6FEF6D8DB" box="[227,322,1942,1967]" name="Vietnam" pageId="6" pageNumber="571">Vietnam</collectingCountry>
(Trung Bo) and southern half of recent
<collectingCountry id="F375D3E3972A3E59FCCABAC8FC8FD8DB" box="[773,827,1944,1967]" name="Laos" pageId="6" pageNumber="571">Laos</collectingCountry>
were united under the name
<collectingCountry id="F375D3E3972A3E59FB4BBAC7FB6ED8DB" box="[1156,1242,1943,1967]" name="Vietnam" pageId="6" pageNumber="571">Annam</collectingCountry>
; thus, it seems to me, that Griffinis species originates from this half of
<collectingCountry id="F375D3E3972A3E59FD3DBAEBFC9CD8A6" box="[754,808,1979,2002]" name="Laos" pageId="6" pageNumber="571">Laos</collectingCountry>
which was included in “
<collectingCountry id="F375D3E3972A3E59FBF3BAEBFB27D8A7" box="[1084,1171,1979,2003]" name="Vietnam" pageId="6" pageNumber="571">Annam</collectingCountry>
” in accordance to these maps. Also, it is necessary to note that many forest genera of Ensifera in this part of
<collectingCountry id="F375D3E3972B3E58FB89BDC8FBC8DFDB" box="[1094,1148,152,175]" name="Laos" pageId="7" pageNumber="572">Laos</collectingCountry>
and in Vietnamese Trung Bo are represented by different species, as most parts of these territories are separated from each other by a long mountain ridge.
</paragraph>
<paragraph id="8BDD9373972B3E58FF08BC55FCAFDD6C" blockId="7.[151,1437,151,1550]" pageId="7" pageNumber="572">
Because the
<typeStatus id="54D92DD1972B3E58FE91BC55FE3ADE69" box="[350,398,261,285]" pageId="7" pageNumber="572">type</typeStatus>
material on this species is destroyed (see the notes to
<taxonomicName id="4C62E8F0972B3E58FBD0BC54FB63DE68" box="[1055,1239,260,284]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B9164F61972B3E58FBD0BC54FB63DE68" box="[1055,1239,260,284]" italics="true" pageId="7" pageNumber="572">Capnogryllacris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225F21A972B3E58FB2CBC55FABBDE69" box="[1251,1295,260,285]" pageId="7" pageNumber="572" sensu="lato">s. l.</taxonomicNameLabel>
above), the synonymization of
<taxonomicName id="4C62E8F0972B3E58FEBEBC78FE57DE34" box="[369,483,296,320]" class="Insecta" family="Haglidae" genus="Marthogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="martha">
<emphasis id="B9164F61972B3E58FEBEBC78FE57DE34" box="[369,483,296,320]" italics="true" pageId="7" pageNumber="572">C. martha</emphasis>
</taxonomicName>
from
<collectingCountry id="F375D3E3972B3E58FDE6BC78FDD4DE4B" box="[553,608,296,319]" name="Laos" pageId="7" pageNumber="572">Laos</collectingCountry>
with
<taxonomicName id="4C62E8F0972B3E58FD6EBC78FB99DE34" authority="Gorochov, 2003" authorityName="Gorochov" authorityYear="2003" box="[673,1069,295,320]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="erythrocephala">
<emphasis id="B9164F61972B3E58FD6EBC78FCD9DE34" box="[673,877,296,320]" italics="true" pageId="7" pageNumber="572">C. erythrocephala</emphasis>
<bibRefCitation id="EFF3EE82972B3E58FCBBBC77FB99DE34" author="Gorochov" box="[884,1069,295,320]" pageId="7" pageNumber="572" refString="Gorochov, A. V. (2003) Contribution to the knowledge of the fauna and systematics of the Stenopelmatoidea (Orthoptera) of Indochina and some other territories. IV. Entomologicheskoe obozrenie [Revue de Entomologie], 82 (3), 629 - 649. [in Russian]" type="journal article" year="2003">Gorochov, 2003</bibRefCitation>
</taxonomicName>
,
<emphasis id="B9164F61972B3E58FBF6BC77FB05DE34" bold="true" box="[1081,1201,295,320]" pageId="7" pageNumber="572">sp. ressur.</emphasis>
from
<collectingCountry id="F375D3E3972B3E58FB38BC77FAE2DE34" box="[1271,1366,295,320]" name="Vietnam" pageId="7" pageNumber="572">Vietnam</collectingCountry>
(
<bibRefCitation id="EFF3EE82972B3E58FAAABC77FF4ADE10" author="Li" pageId="7" pageNumber="572" refString="Li, M., Fang, Ya., Liu, X. &amp; Li, K. (2014) Review of the species of the genus Marthogryllacris (Orthoptera, Gryllacrididae, Gryllacridinae). Zootaxa, 3889 (2), 277 - 288. http: // dx. doi. org / 10.11646 / zootaxa. 3889.2.6" type="journal article" year="2014">
Li
<emphasis id="B9164F61972B3E58FA47BC79FF00DE10" italics="true" pageId="7" pageNumber="572">et al.</emphasis>
, 2014
</bibRefCitation>
) seems to us very problematic. Moreover, there are some differences in the colouration and body size between these species which follow from the decriptions by
<bibRefCitation id="EFF3EE82972B3E58FC8ABC3FFC58DEFC" author="Griffini" box="[837,1004,367,392]" pageId="7" pageNumber="572" refString="Griffini A. (1914) Sopra due Gryllacris del Museo di Budapest. Annales Musei Nationalis Hungarici, 12, 249 - 260." type="journal article" year="1914">Griffini (1914)</bibRefCitation>
and
<bibRefCitation id="EFF3EE82972B3E58FBEABC3FFB53DEFC" author="Gorochov" box="[1061,1255,367,392]" pageId="7" pageNumber="572" refString="Gorochov, A. V. (2003) Contribution to the knowledge of the fauna and systematics of the Stenopelmatoidea (Orthoptera) of Indochina and some other territories. IV. Entomologicheskoe obozrenie [Revue de Entomologie], 82 (3), 629 - 649. [in Russian]" type="journal article" year="2003">Gorochov (2003)</bibRefCitation>
as well as from the brief comparison below. In this connection, the fixation of Griffinis species name with help of the
<typeStatus id="54D92DD1972B3E58FA8CBCC5FA28DED9" box="[1347,1436,405,429]" pageId="7" pageNumber="572" type="neotype">neotype</typeStatus>
designation will be very useful for stability of the
<taxonomicName id="4C62E8F0972B3E58FD01BCE7FCC4DEA4" box="[718,880,439,464]" class="Insecta" family="Gryllacrididae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="subFamily" subFamily="Gryllacridinae">Gryllacridinae</taxonomicName>
classification (this species is also
<typeStatus id="54D92DD1972B3E58FB3BBCE8FA90DEA4" box="[1268,1316,440,464]" pageId="7" pageNumber="572">type</typeStatus>
species of the generic name
<taxonomicName id="4C62E8F0972B3E58FE93BC8CFDA9DE80" box="[348,541,476,500]" class="Insecta" family="Haglidae" genus="Marthogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="B9164F61972B3E58FE93BC8CFDA9DE80" box="[348,541,476,500]" italics="true" pageId="7" pageNumber="572">Marthogryllacris</emphasis>
</taxonomicName>
). The specimen, designated here as
<typeStatus id="54D92DD1972B3E58FC7EBC8DFBB9DE81" box="[945,1037,477,501]" pageId="7" pageNumber="572" type="neotype">neotype</typeStatus>
, originates from the
<typeStatus id="54D92DD1972B3E58FB3CBC8DFA97DE81" box="[1267,1315,477,501]" pageId="7" pageNumber="572">type</typeStatus>
locality of
<taxonomicName id="4C62E8F0972B3E58FF58BF50FEBCDD6C" box="[151,264,512,536]" class="Insecta" family="Haglidae" genus="Marthogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="martha">
<emphasis id="B9164F61972B3E58FF58BF50FEBCDD6C" box="[151,264,512,536]" italics="true" pageId="7" pageNumber="572">C. martha</emphasis>
</taxonomicName>
and is in accordance to its original description.
</paragraph>
<paragraph id="8BDD9373972B3E58FF08BF74FDCADB43" blockId="7.[151,1437,151,1550]" pageId="7" pageNumber="572">
<emphasis id="B9164F61972B3E58FF08BF74FE2ADD48" box="[199,414,548,572]" italics="true" pageId="7" pageNumber="572">Brief redescription</emphasis>
. Male (
<typeStatus id="54D92DD1972B3E58FE33BF74FDEFDD48" box="[508,603,548,572]" pageId="7" pageNumber="572" type="neotype">neotype</typeStatus>
). Colouration and structure of body more or less similar to those of
<taxonomicName id="4C62E8F0972B3E58FA4DBF74FEF5DD14" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="erythrocephala">
<emphasis id="B9164F61972B3E58FA4DBF74FEF5DD14" italics="true" pageId="7" pageNumber="572">C. erythrocephala</emphasis>
</taxonomicName>
and
<taxonomicName id="4C62E8F0972B3E58FEB6BF18FD40DD14" authority="Gorochov, 2003" authorityName="Gorochov" authorityYear="2003" box="[377,756,583,608]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="phaeocephala">
<emphasis id="B9164F61972B3E58FEB6BF18FD83DD14" box="[377,567,584,608]" italics="true" pageId="7" pageNumber="572">C. phaeocephala</emphasis>
<bibRefCitation id="EFF3EE82972B3E58FDF0BF17FD40DD14" author="Gorochov" box="[575,756,583,608]" pageId="7" pageNumber="572" refString="Gorochov, A. V. (2003) Contribution to the knowledge of the fauna and systematics of the Stenopelmatoidea (Orthoptera) of Indochina and some other territories. IV. Entomologicheskoe obozrenie [Revue de Entomologie], 82 (3), 629 - 649. [in Russian]" type="journal article" year="2003">Gorochov, 2003</bibRefCitation>
</taxonomicName>
but with following differences: head more similar to that of
<taxonomicName id="4C62E8F0972B3E58FF58BF3CFD38DDF0" authority="Gorochov, 2003" authorityName="Gorochov" authorityYear="2003" box="[151,652,620,645]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="subSpecies" species="erythrocephala" subSpecies="borealis">
<emphasis id="B9164F61972B3E58FF58BF3CFE7DDDF0" box="[151,457,620,644]" italics="true" pageId="7" pageNumber="572">C. erythrocephala borealis</emphasis>
<bibRefCitation id="EFF3EE82972B3E58FE1DBF3CFD38DDF0" author="Gorochov" box="[466,652,620,645]" pageId="7" pageNumber="572" refString="Gorochov, A. V. (2003) Contribution to the knowledge of the fauna and systematics of the Stenopelmatoidea (Orthoptera) of Indochina and some other territories. IV. Entomologicheskoe obozrenie [Revue de Entomologie], 82 (3), 629 - 649. [in Russian]" type="journal article" year="2003">Gorochov, 2003</bibRefCitation>
</taxonomicName>
,
<emphasis id="B9164F61972B3E58FD56BF3CFC90DDF1" bold="true" box="[665,804,620,645]" pageId="7" pageNumber="572">stat. ressur.</emphasis>
(Northern
<collectingCountry id="F375D3E3972B3E58FC6ABF3CFBB2DDF1" box="[933,1030,620,645]" name="Vietnam" pageId="7" pageNumber="572">Vietnam</collectingCountry>
) in colouration (Fig. 38), i. e. with dorsal and anterior parts of epicranium as well as clypeus light brown, but with lateral parts including genae, mandibles and labrum dark brown (
<bibRefCitation id="EFF3EE82972B3E58FD88BFE4FD0EDD84" author="Griffini" pageId="7" pageNumber="572" refString="Griffini A. (1914) Sopra due Gryllacris del Museo di Budapest. Annales Musei Nationalis Hungarici, 12, 249 - 260." type="journal article" year="1914" yearSuffix="p">Griffini, 1914: p. 257—“caput occipute, vertice, fronte, clypeoque totis testaceis; labro, mandibulis, genisque totis atris”</bibRefCitation>
); but pronotum more similar to that of
<taxonomicName id="4C62E8F0972B3E58FBA1BF88FA9FDD84" box="[1134,1323,728,752]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="phaeocephala">
<emphasis id="B9164F61972B3E58FBA1BF88FA9FDD84" box="[1134,1323,728,752]" italics="true" pageId="7" pageNumber="572">C. phaeocephala</emphasis>
</taxonomicName>
(
<figureCitation id="13598FF6972B3E58FAF4BF87FA3BDD84" box="[1339,1423,727,752]" captionStart="FIGURES 52 55" captionStartId="12.[151,264,1510,1533]" captionTargetBox="[191,1381,187,1480]" captionTargetId="figure@12.[182,1387,181,1492]" captionTargetPageId="12" captionText="FIGURES 52 55. Capnogryllacris Karny, male body from above (left wings spread): 52, C. martha (Griff.); 53, C. erythocephala erythrocephala Gor.; 54, C. rubrocellata Gor.; 55, C. phaeocephala cambodiensis subsp. nov." httpUri="https://zenodo.org/record/245308/files/figure.png" pageId="7" pageNumber="572">Fig. 52</figureCitation>
), as its colouration dark brown with small reddish brown median spot along anterior edge and with light brown transverse band along posterior edge running from posterior parts of lateral lobes to posterior part of disc (
<bibRefCitation id="EFF3EE82972B3E58FAF1BE4FFEBDDCF4" author="Griffini" pageId="7" pageNumber="572" refString="Griffini A. (1914) Sopra due Gryllacris del Museo di Budapest. Annales Musei Nationalis Hungarici, 12, 249 - 260." type="journal article" year="1914" yearSuffix="p">Griffini, 1914: p. 257—“pronotum atrum, metazona et parte postica loborum lateralium testaceis, margine antico latiuscula castaneo”</bibRefCitation>
); hind wings with thin darkened (greyish) stripes along crossveins similar to that of
<taxonomicName id="4C62E8F0972B3E58FB3ABE38FA20DCF4" box="[1269,1428,872,896]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="subSpecies" species="erythrocephala" subSpecies="borealis">
<emphasis id="B9164F61972B3E58FB3ABE38FA20DCF4" box="[1269,1428,872,896]" italics="true" pageId="7" pageNumber="572">C. e. borealis</emphasis>
</taxonomicName>
; tegmina somewhat widened to distal part, distinctly wider than in both above-mentioned species (for comparison see
<figureCitation id="13598FF6972B3E58FF09BEE2FEC8DCBF" box="[198,380,946,971]" captionStart="FIGURES 52 55" captionStartId="12.[151,264,1510,1533]" captionTargetBox="[191,1381,187,1480]" captionTargetId="figure@12.[182,1387,181,1492]" captionTargetPageId="12" captionText="FIGURES 52 55. Capnogryllacris Karny, male body from above (left wings spread): 52, C. martha (Griff.); 53, C. erythocephala erythrocephala Gor.; 54, C. rubrocellata Gor.; 55, C. phaeocephala cambodiensis subsp. nov." httpUri="https://zenodo.org/record/245308/files/figure.png" pageId="7" pageNumber="572">Figs 52, 53, 55</figureCitation>
); hooks of 9th abdominal tergite clearly different from those of latter representatives, with distinctly thickened proximal part and longer and narrower rest part (for comparison see
<figureCitation id="13598FF6972B3E58FBB3BE86FB4ADC9B" box="[1148,1278,982,1007]" captionStart="FIGURES 40 51" captionStartId="11.[151,264,1857,1880]" captionTargetBox="[156,1433,206,1829]" captionTargetId="figure@11.[151,1436,193,1837]" captionTargetPageId="11" captionText="FIGURES 40 51. Capnogryllacris Karny, male (40 50) and female (51): 40 44, C. sakaerat sp. nov. (40 42, holotype; 43, 44, paratype); 45, C. martha (Griff.); 45, C. erythrocephala erythrocephala Gor.; 47; C. e. borealis Gor.; 48, C. phaeocephala cambodiensis subsp. nov.; 49, C. borneoensis pileata Gor.; 50, C. fumigata miniata Karny; 51, C. khmerica Gor. Body from side (40); right tegmen in rest position from side and slightly above (41, 43); specimen in living condition (42, 44); abdominal apex from below (45 50); genital plate from below (51)." httpUri="https://zenodo.org/record/245307/files/figure.png" pageId="7" pageNumber="572">Figs 4548</figureCitation>
); genital plate with styles intermediate in length between those of these species and with posteromedian notch clearly less angular than in both these species (see
<figureCitation id="13598FF6972B3E58FE3FB94EFDC5DB43" box="[496,625,1054,1079]" captionStart="FIGURES 40 51" captionStartId="11.[151,264,1857,1880]" captionTargetBox="[156,1433,206,1829]" captionTargetId="figure@11.[151,1436,193,1837]" captionTargetPageId="11" captionText="FIGURES 40 51. Capnogryllacris Karny, male (40 50) and female (51): 40 44, C. sakaerat sp. nov. (40 42, holotype; 43, 44, paratype); 45, C. martha (Griff.); 45, C. erythrocephala erythrocephala Gor.; 47; C. e. borealis Gor.; 48, C. phaeocephala cambodiensis subsp. nov.; 49, C. borneoensis pileata Gor.; 50, C. fumigata miniata Karny; 51, C. khmerica Gor. Body from side (40); right tegmen in rest position from side and slightly above (41, 43); specimen in living condition (42, 44); abdominal apex from below (45 50); genital plate from below (51)." httpUri="https://zenodo.org/record/245307/files/figure.png" pageId="7" pageNumber="572">Figs 4548</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C378C0F8972B3E58FF08B912FCD7DA7B" pageId="7" pageNumber="572" type="description">
<paragraph id="8BDD9373972B3E58FF08B912FA35DB2F" blockId="7.[151,1437,151,1550]" box="[199,1409,1090,1115]" pageId="7" pageNumber="572">
<emphasis id="B9164F61972B3E58FF08B912FEC2DB2F" bold="true" box="[199,374,1090,1115]" pageId="7" pageNumber="572">Measurements</emphasis>
(in mm). Length of body 32; body with wings 56; pronotum 8.8; tegmen 42; hind femur 24.
</paragraph>
<paragraph id="8BDD9373972B3E58FF08B936FCD7DA7B" blockId="7.[151,1437,151,1550]" pageId="7" pageNumber="572">
<emphasis id="B9164F61972B3E58FF08B936FEEEDB0B" bold="true" box="[199,346,1126,1151]" pageId="7" pageNumber="572">Comparison</emphasis>
. This species differs from
<taxonomicName id="4C62E8F0972B3E58FD48B937FCE6DB0B" box="[647,850,1127,1151]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="erythrocephala">
<emphasis id="B9164F61972B3E58FD48B937FCE6DB0B" box="[647,850,1127,1151]" italics="true" pageId="7" pageNumber="572">C. erythrocephala</emphasis>
</taxonomicName>
in the absence of large yellowish or reddish area on the pronotal disc along its anterior edge; from
<taxonomicName id="4C62E8F0972B3E58FD60B9DBFC2DDBD7" box="[687,921,1163,1187]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="subSpecies" species="erythrocephala" subSpecies="erythrocephala">
<emphasis id="B9164F61972B3E58FD60B9DBFC2DDBD7" box="[687,921,1163,1187]" italics="true" pageId="7" pageNumber="572">C. e. erythrocephala</emphasis>
</taxonomicName>
and
<taxonomicName id="4C62E8F0972B3E58FC1BB9DBFB27DBD7" box="[980,1171,1163,1187]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="phaeocephala">
<emphasis id="B9164F61972B3E58FC1BB9DBFB27DBD7" box="[980,1171,1163,1187]" italics="true" pageId="7" pageNumber="572">C. phaeocephala</emphasis>
</taxonomicName>
, in a larger body, light brown (not obviously reddish, black or dark brown) coloration of the head most part, and narrower darkened stripes situated in the hind wings along their crossveins; and from both these species, in a clearly wider distal tegminal half and in the male abdominal characters listed in the redescription.
</paragraph>
</subSubSection>
<subSubSection id="C378C0F8972B3E58FF08B84AFADAD979" pageId="7" pageNumber="572" type="discussion">
<paragraph id="8BDD9373972B3E58FF08B84AFADAD979" blockId="7.[151,1437,151,1550]" pageId="7" pageNumber="572">
<emphasis id="B9164F61972B3E58FF08B84AFE8FDA47" bold="true" box="[199,315,1306,1331]" pageId="7" pageNumber="572">Remarks.</emphasis>
It is necessary to note that
<taxonomicName id="4C62E8F0972B3E58FDBBB84BFD53DA47" box="[628,743,1307,1331]" class="Insecta" family="Haglidae" genus="Marthogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="martha">
<emphasis id="B9164F61972B3E58FDBBB84BFD53DA47" box="[628,743,1307,1331]" italics="true" pageId="7" pageNumber="572">C. martha</emphasis>
</taxonomicName>
, recorded by
<bibRefCitation id="EFF3EE82972B3E58FC4AB84AFB86DA47" box="[901,1074,1306,1331]" pageId="7" pageNumber="572" refString="Li, M., Fang, Ya., Liu, X. &amp; Li, K. (2014) Review of the species of the genus Marthogryllacris (Orthoptera, Gryllacrididae, Gryllacridinae). Zootaxa, 3889 (2), 277 - 288. http: // dx. doi. org / 10.11646 / zootaxa. 3889.2.6" type="journal article">
Li
<emphasis id="B9164F61972B3E58FC6AB84BFC6BDA47" box="[933,991,1307,1331]" italics="true" pageId="7" pageNumber="572">et al.</emphasis>
(2014)
</bibRefCitation>
as “
<taxonomicName id="4C62E8F0972B3E58FBAAB84BFAC6DA47" box="[1125,1394,1307,1331]" class="Insecta" family="Stenopelmatidae" genus="Martogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="martha">
<emphasis id="B9164F61972B3E58FBAAB84BFAC6DA47" box="[1125,1394,1307,1331]" italics="true" pageId="7" pageNumber="572">Martogryllacris martha</emphasis>
</taxonomicName>
” in Northern
<collectingCountry id="F375D3E3972B3E58FECBB86EFED7DA23" box="[260,355,1342,1367]" name="Vietnam" pageId="7" pageNumber="572">Vietnam</collectingCountry>
(Ba Vi), is most probably
<taxonomicName id="4C62E8F0972B3E58FD5EB86FFC9CDA23" box="[657,808,1343,1367]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="subSpecies" species="erythrocephala" subSpecies="borealis">
<emphasis id="B9164F61972B3E58FD5EB86FFC9CDA23" box="[657,808,1343,1367]" italics="true" pageId="7" pageNumber="572">C. e. borealis</emphasis>
</taxonomicName>
for the following reasons: we have one similar male of this subspecies from the same locality; the female genital plate, pictured by these authors, differs from that of
<taxonomicName id="4C62E8F0972B3E58FAA7B833FEF5DAEB" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="subSpecies" species="erythrocephala" subSpecies="erythrocephala">
<emphasis id="B9164F61972B3E58FAA7B833FEF5DAEB" italics="true" pageId="7" pageNumber="572">C. e. erythrocephala</emphasis>
</taxonomicName>
in a more widely truncate apex; and other females of
<taxonomicName id="4C62E8F0972B3E58FC75B8D7FBE2DAEB" box="[954,1110,1415,1439]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="subSpecies" species="erythrocephala" subSpecies="borealis">
<emphasis id="B9164F61972B3E58FC75B8D7FBE2DAEB" box="[954,1110,1415,1439]" italics="true" pageId="7" pageNumber="572">C. e. borealis</emphasis>
</taxonomicName>
are unknown. However, the species indicated in the same paper as “
<taxonomicName id="4C62E8F0972B3E58FD99B8FBFCD8DAB7" box="[598,876,1451,1475]" class="Insecta" family="Stenopelmatidae" genus="Martogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B9164F61972B3E58FD99B8FBFCD8DAB7" box="[598,876,1451,1475]" italics="true" pageId="7" pageNumber="572">Martogryllacris borealis</emphasis>
</taxonomicName>
” for
<collectingCountry id="F375D3E3972B3E58FC67B8FAFC5FDAB7" box="[936,1003,1450,1475]" name="China" pageId="7" pageNumber="572">China</collectingCountry>
is possibly another species, because it differs from
<taxonomicName id="4C62E8F0972B3E58FEE5B89FFE72DA93" box="[298,454,1487,1511]" class="Insecta" family="Haglidae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="572" phylum="Arthropoda" rank="subSpecies" species="erythrocephala" subSpecies="borealis">
<emphasis id="B9164F61972B3E58FEE5B89FFE72DA93" box="[298,454,1487,1511]" italics="true" pageId="7" pageNumber="572">C. e. borealis</emphasis>
</taxonomicName>
in a distinctly shallower posteromedian notch of the male genital plate as well as in clearly different hooks of the male 9th abdominal tergite. Thus, we restore here subspecies status for this taxon.
</paragraph>
</subSubSection>
</treatment>
</document>