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<document id="FB8007B1533F6982FB2470C5936B6886" ID-CLB-Dataset="107004" ID-DOI="10.11646/zootaxa.5167.1.1" ID-GBIF-Dataset="86fc8470-92ab-4040-9a60-9c984cceff65" ID-ISSN="1175-5326" ID-Zenodo-Dep="6876483" ID-ZooBank="4C140613-04F6-4227-B084-45851F42E039" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1658418473528" checkinUser="plazi" docAuthor="Shinohara, Akihiko, Kramp, Katja &amp; Taeger, Andreas" docDate="2022" docId="FB3C87F1F23FAC57FF67FF04FD1CAF54" docLanguage="en" docName="zootaxa.5167.1.1.pdf" docOrigin="Zootaxa 5167 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5167.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pamphilius graciloides Shinohara &amp; Kramp &amp; Taeger 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="81" masterDocId="0705FF89F270AC07FFF0FF93FFAFAB03" masterDocTitle="The Pamphiliinae of the Russian Far East and Korea (Hymenoptera, Pamphiliidae)" masterLastPageNumber="251" masterPageNumber="1" pageNumber="80" updateTime="1699361468654" updateUser="ExternalLinkService">
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<mods:title id="7C6B07A03BBA90EC0882286846DBA112">The Pamphiliinae of the Russian Far East and Korea (Hymenoptera, Pamphiliidae)</mods:title>
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<taxonomicName id="B4954D64F23FAC48FF67FF04FE02ABB1" authority="Shinohara &amp; Kramp &amp; Taeger, 2022" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[151,429,151,178]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="graciloides" status="sp. nov.">
<emphasis id="41E1EAF5F23FAC48FF67FF04FE02ABB1" bold="true" box="[151,429,151,178]" italics="true" pageId="79" pageNumber="80">Pamphilius graciloides</emphasis>
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<subSubSection id="3B8F656CF23FAC48FF67FF28FCEBABD5" box="[151,836,187,214]" pageId="79" pageNumber="80" type="description">
<paragraph id="732A36E7F23FAC48FF67FF28FCEBABD5" blockId="79.[151,836,151,214]" box="[151,836,187,214]" pageId="79" pageNumber="80">
(
<figureCitation id="EBAE2A62F23FAC48FF6FFF28FF59ABD5" box="[159,246,187,214]" captionStart="FIGURE 75" captionStartId="164.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-26@164.[151,1435,181,1828]" captionTargetPageId="164" captionText="FIGURE 75. Pamphilius graciloides, female, Russia (paratype) (a) head &amp; thorax dorsally, (b) head &amp; thorax ventrolaterally, (c) sawsheath laterally, (d) face, (e) antenna, (f) fore wing." figureDoi="http://doi.org/10.5281/zenodo.6876687" httpUri="https://zenodo.org/record/6876687/files/figure.png" pageId="79" pageNumber="80">Figs 75</figureCitation>
,
<figureCitation id="EBAE2A62F23FAC48FEF2FF28FEB0ABD6" box="[258,287,187,213]" captionStart="FIGURE 76" captionStartId="165.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-19@165.[151,1435,181,1828]" captionTargetPageId="165" captionText="FIGURE 76. Pamphilius graciloides, male, Korea (holotype) (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) antenna, (g) genitalia dorsally, (h) genitalia ventrally, (i) claw." figureDoi="http://doi.org/10.5281/zenodo.6876689" httpUri="https://zenodo.org/record/6876689/files/figure.png" pageId="79" pageNumber="80">76</figureCitation>
) (https://doi.org/10.6084/m9.figshare.11405232)
</paragraph>
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<subSubSection id="3B8F656CF23FAC48FF67FE93FD3CAA35" pageId="79" pageNumber="80" type="reference_group">
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<treatmentCitationGroup id="538511C9F23FAC48FF67FE93FD3CAA35" pageId="79" pageNumber="80">
<treatmentCitation id="F23410F6F23FAC48FF67FE93FDEBAA1B" author="Shinohara, A." box="[151,580,256,280]" page="110" pageId="79" pageNumber="80" year="2001">
<taxonomicName id="B4954D64F23FAC48FF67FE93FDEBAA1B" authority=": Shinohara, 2001: 110" authorityName=": Shinohara" authorityPageNumber="110" authorityYear="2001" box="[151,580,256,280]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F23FAC48FF67FE93FEF2AA1B" box="[151,349,256,280]" italics="true" pageId="79" pageNumber="80">Pamphilius gracilis</emphasis>
:
<bibRefCitation id="17044B16F23FAC48FE9BFE93FDEBAA1B" author="Shinohara, A." box="[363,580,256,280]" pageId="79" pageNumber="80" pagination="99 - 116" refId="ref76721" refString="Shinohara, A. (2001) The group of Pamphilius sylvaticus (Hymenoptera, Pamphiliidae): Two new species, new collection records, and a key to Palearctic species. Japanese Journal of Systematic Entomology, 7, 99 - 116." type="journal article" year="2001">Shinohara, 2001: 110</bibRefCitation>
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, 114, 115;
<treatmentCitation id="F23410F6F23FAC48FD41FE93FC36AA1B" author="Shinohara, A." box="[689,921,256,280]" page="430" pageId="79" pageNumber="80" year="2002">
<bibRefCitation id="17044B16F23FAC48FD41FE93FC36AA1B" author="Shinohara, A." box="[689,921,256,280]" pageId="79" pageNumber="80" pagination="359 - 438" refId="ref76809" refString="Shinohara, A. (2002 b) Systematics of the leaf-rolling or webspinning sawfly subfamily Pamphiliinae: a preliminary overview. In: Viitasaari, M. (Ed.), Sawflies (Hymenoptera, Symphyta) I. Tremex Press, Helsinki, pp. 359 - 438." type="book chapter" year="2002">Shinohara, 2002b: 430</bibRefCitation>
</treatmentCitation>
; Shinohara, 2004: 265;
<treatmentCitation id="F23410F6F23FAC48FB78FE93FAC3AA14" author="Taeger, A. &amp; Blank, S. M. &amp; Liston, A. D." box="[1160,1388,255,280]" page="86" pageId="79" pageNumber="80" year="2010">
<bibRefCitation id="17044B16F23FAC48FB78FE93FAC3AA14" author="Taeger, A. &amp; Blank, S. M. &amp; Liston, A. D." box="[1160,1388,255,280]" pageId="79" pageNumber="80" pagination="1 - 1064" refId="ref80219" refString="Taeger, A., Blank, S. M. &amp; Liston, A. D. (2010) World catalog of Symphyta (Hymenoptera). Zootaxa, 2580 (1), 1 - 1064. https: // doi. org / 10.11646 / zootaxa. 2580.1.1" type="journal article" year="2010">
Taeger
<emphasis id="41E1EAF5F23FAC48FB22FE93FAAAAA1B" box="[1234,1285,256,280]" italics="true" pageId="79" pageNumber="80">et al.</emphasis>
, 2010: 86
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F23410F6F23FAC48FA86FE93FEF9AA35" author="Lee, J. - W. &amp; Choi, J. - K. &amp; Park, B." page="9" pageId="79" pageNumber="80" year="2019">
<bibRefCitation id="17044B16F23FAC48FA86FE93FEF9AA35" author="Lee, J. - W. &amp; Choi, J. - K. &amp; Park, B." pageId="79" pageNumber="80" pagination="1 - 96" refId="ref74711" refString="Lee, J. - W., Choi, J. - K. &amp; Park, B. (2019) Synoptic list of Symphyta (Hymenoptera) in Korea. Journal of Species Research, 8, 1 - 96." type="journal article" year="2019">
Lee
<emphasis id="41E1EAF5F23FAC48FF37FE8CFF55AA35" box="[199,250,286,310]" italics="true" pageId="79" pageNumber="80">et al.</emphasis>
, 2019: 9
</bibRefCitation>
</treatmentCitation>
. Not
<bibRefCitation id="17044B16F23FAC48FE7CFE8DFDEFAA35" author="Shinohara, A." box="[396,576,286,310]" pageId="79" pageNumber="80" pagination="323 - 351" refId="ref76025" refString="Shinohara, A. (1985 a) Web-spinning sawflies of the sylvaticus group of Pamphilius (Hymenoptera, Pamphiliidae). Systematic Entomology, 10, 323 - 351. https: // doi. org / 10.1111 / j. 1365 - 3113.1985. tb 00141. x" type="journal article" year="1985">Shinohara, 1985a</bibRefCitation>
, in part.
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</paragraph>
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<subSubSection id="3B8F656CF23FAC48FF67FEF7FCC3AD21" pageId="79" pageNumber="80" type="description">
<paragraph id="732A36E7F23FAC48FF67FEF7FB71A9C1" blockId="79.[151,1437,356,2039]" pageId="79" pageNumber="80">
<emphasis id="41E1EAF5F23FAC48FF67FEF7FE28AA7C" bold="true" box="[151,391,356,383]" pageId="79" pageNumber="80">Description. Female</emphasis>
(
<figureCitation id="EBAE2A62F23FAC48FE66FEF7FE44AA7C" box="[406,491,356,383]" captionStart="FIGURE 75" captionStartId="164.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-26@164.[151,1435,181,1828]" captionTargetPageId="164" captionText="FIGURE 75. Pamphilius graciloides, female, Russia (paratype) (a) head &amp; thorax dorsally, (b) head &amp; thorax ventrolaterally, (c) sawsheath laterally, (d) face, (e) antenna, (f) fore wing." figureDoi="http://doi.org/10.5281/zenodo.6876687" httpUri="https://zenodo.org/record/6876687/files/figure.png" pageId="79" pageNumber="80">Fig. 75</figureCitation>
). Length about 11.5 mm. Head black, with following pale yellow: linear mark along median anterior margin of clypeus, paired spots on frons, spot on upper margin of inner orbit, spot on posterior margin of postocular area, elongate spot on gena along lower outer orbit; mandible black, with outer basal part and subapical part pale yellow and apex dark rufous; antenna black. Thorax black, with following pale yellow: large spot on ventral margin of lateral pronotum, posterolateral corner of dorsal pronotum, tegula, posterior half of mesoscutal middle lobe, mesoscutellum, minute spot on anterodorsal corner of mesepisternum, metascutellum; legs pale yellow, with fore coxa (except for narrow apical margin), fore trochanter and basal half of fore femur (except for anterior surface of trochantellus), mid coxa (except for apical margin) and hind coxa (except for broad apical part) black. Wings hyaline, slightly tinted with blackish brown; veins and stigma black. Abdomen black, with outer margin of each laterotergite and posterior margin of each sternum (more broadly on sterna 47) pale yellow.
</paragraph>
<paragraph id="732A36E7F23FAC48FF37FD5EFE01AF05" blockId="79.[151,1437,356,2039]" pageId="79" pageNumber="80">
Frons rather strongly swollen; ocellar basin subtriangular, represented as broad furrow around median ocellus; upper frons not distinctly notched medially; frontoclypeal crest rather flat, rounded, swollen between antennal sockets; facial crest strongly convex, rounded. Upper part of head behind transverse and lateral transverse sutures smooth, weakly coriaceous, with sparse and small punctures; frons, area from facial crest to lateral transverse suture, and paraantennal field shallowly coarsely rugose; clypeus with distinct medium-sized punctures, with interspaces coriaceous medially and coarsely rugose laterally; gena coriaceous, with rather small, often ill-defined punctures; head covered with rather short silvery hairs before crassa. Antennae with 23 antennomeres; flagellomere 1 1.1 × length of flagellomere 2. Forewing with cell C pilose all over. Ovipositor sheath as in
<figureCitation id="EBAE2A62F23FAC48FBBFFC5BFB02A8E0" box="[1103,1197,968,995]" captionStart="FIGURE 75" captionStartId="164.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-26@164.[151,1435,181,1828]" captionTargetPageId="164" captionText="FIGURE 75. Pamphilius graciloides, female, Russia (paratype) (a) head &amp; thorax dorsally, (b) head &amp; thorax ventrolaterally, (c) sawsheath laterally, (d) face, (e) antenna, (f) fore wing." figureDoi="http://doi.org/10.5281/zenodo.6876687" httpUri="https://zenodo.org/record/6876687/files/figure.png" pageId="79" pageNumber="80">Fig. 75c</figureCitation>
; appendage large and elongate, setose apically.
</paragraph>
<paragraph id="732A36E7F23FAC48FF37FB82FADEAE6D" blockId="79.[151,1437,356,2039]" pageId="79" pageNumber="80">
<emphasis id="41E1EAF5F23FAC48FF37FB82FEABAF28" bold="true" box="[199,260,1041,1067]" pageId="79" pageNumber="80">Male</emphasis>
(
<figureCitation id="EBAE2A62F23FAC48FEE2FB83FECBAF29" box="[274,356,1040,1066]" captionStart="FIGURE 76" captionStartId="165.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-19@165.[151,1435,181,1828]" captionTargetPageId="165" captionText="FIGURE 76. Pamphilius graciloides, male, Korea (holotype) (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) antenna, (g) genitalia dorsally, (h) genitalia ventrally, (i) claw." figureDoi="http://doi.org/10.5281/zenodo.6876689" httpUri="https://zenodo.org/record/6876689/files/figure.png" pageId="79" pageNumber="80">Fig. 76</figureCitation>
). Length about
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. Head black; pale yellow are frontal surface before line connecting facial crests, minute spot at upper margin of inner orbit (sometimes missing) and most of gena and malar space (median part of malar space always black); mandible pale yellow, inner half usually black and apex dark rufous; scape pale yellow, with large black spot above; pedicel and flagellum blackish brown. Thorax black, with following pale yellow: large spot on ventral margin of lateral pronotum (sometimes small), posterolateral corner of dorsal pronotum (often narrowly), tegula, posterior half of mesoscutal middle lobe, mesoscutellum, broad anterior margin and large posterolateral spot on mesepisternum (often reduced or missing), metascutellum, and large spot on metepisternum; legs pale yellow, with most of coxae black; fore femur often obscurely marked with black posteriorly. Wings hyaline, slightly stained with blackish brown; veins and stigma black. Abdomen black, with most of laterotergites, broad posterior margin of each sternum and entire subgenital plate pale yellow. Genitalia black; harpe largely pale brown.
</paragraph>
<paragraph id="732A36E7F23FAC48FF37FAEAFCC3AD21" blockId="79.[151,1437,356,2039]" pageId="79" pageNumber="80">
Structure generally similar to that of female. Frons very strongly swollen; ocellar basin often indistinct anteriorly; upper frons very shallowly notched medially; facial crest very strongly convex, bluntly carinate. Antennae with 2223 antennomeres; flagellomere 1 1.01.1 × length of flagellomere 2 and slightly thinner in lateral view than the latter, with apex weakly oblique. Subgenital plate with apical margin broadly truncated. Genitalia as in
<figureCitation id="EBAE2A62F23FAC48FAC5FA77FF07AD21" captionStart="FIGURE 76" captionStartId="165.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-19@165.[151,1435,181,1828]" captionTargetPageId="165" captionText="FIGURE 76. Pamphilius graciloides, male, Korea (holotype) (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) antenna, (g) genitalia dorsally, (h) genitalia ventrally, (i) claw." figureDoi="http://doi.org/10.5281/zenodo.6876689" httpUri="https://zenodo.org/record/6876689/files/figure.png" pageId="79" pageNumber="80">
Figs
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, h
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; penis valve short and elongate-subtriangular seen from above.
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF23FAC48FF37F9BFFBD0AC88" pageId="79" pageNumber="80" type="materials_examined">
<paragraph id="732A36E7F23FAC48FF37F9BFFBD0AC88" blockId="79.[151,1437,356,2039]" pageId="79" pageNumber="80">
<emphasis id="41E1EAF5F23FAC48FF37F9BFFE1BAD45" bold="true" box="[199,436,1580,1606]" pageId="79" pageNumber="80">Material examined.</emphasis>
Eighteen specimens.
<materialsCitation id="C3FD3CBAF23FAC48FD42F9BEFC0EAD68" ID-GBIF-Occurrence="3866537368" collectingDate="1996-06-06" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" elevation="1300" latitude="43.102" location="South Korea" longLatPrecision="68" longitude="132.786" pageId="79" pageNumber="80" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Primorskiy" typeStatus="holotype">
<emphasis id="41E1EAF5F23FAC48FD42F9BEFC87AD44" bold="true" box="[690,808,1581,1607]" pageId="79" pageNumber="80">
<typeStatus id="AC2E8845F23FAC48FD42F9BEFC8CAD44" box="[690,803,1581,1607]" pageId="79" pageNumber="80" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="6593FD6EF23FAC48FCC2F9BEFCE8AD44" box="[818,839,1581,1607]" pageId="79" pageNumber="80" type="male"></specimenCount>
(
<figureCitation id="EBAE2A62F23FAC48FCAAF9BFFC1EAD45" box="[858,945,1580,1606]" captionStart="FIGURE 76" captionStartId="165.[151,250,1851,1875]" captionTargetBox="[151,1435,181,1828]" captionTargetId="figure-19@165.[151,1435,181,1828]" captionTargetPageId="165" captionText="FIGURE 76. Pamphilius graciloides, male, Korea (holotype) (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) antenna, (g) genitalia dorsally, (h) genitalia ventrally, (i) claw." figureDoi="http://doi.org/10.5281/zenodo.6876689" httpUri="https://zenodo.org/record/6876689/files/figure.png" pageId="79" pageNumber="80">Fig. 76</figureCitation>
), Mirugam (Bukdaesa),
<quantity id="B46D9B02F23FAC48FB3BF9BEFA82AD44" box="[1227,1325,1581,1607]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="79" pageNumber="80" unit="m" value="1300.0">
<elevation id="F8B8D1D4F23FAC48FB3BF9BEFA82AD44" box="[1227,1325,1581,1607]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="79" pageNumber="80" unit="m" value="1300.0">1,300 m</elevation>
</quantity>
, Odaesan Mts., Gangwon-do,
<collectingCountry id="0B827677F23FAC48FE88F9C3FDA8AD68" box="[376,519,1616,1643]" name="South Korea" pageId="79" pageNumber="80">South Korea</collectingCountry>
,
<date id="072B1027F23FAC48FDE3F9C3FD3AAD69" box="[531,661,1616,1643]" pageId="79" pageNumber="80" value="1996-06-06">
<collectingDate id="176FE9CFF23FAC48FDE3F9C3FD3AAD69" box="[531,661,1616,1643]" pageId="79" pageNumber="80" value="1996-06-06">6. VI. 1996</collectingDate>
</date>
, A.
<collectorName id="DE605331F23FAC48FD31F9C3FC9CAD69" box="[705,819,1616,1642]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FCB1F9C2FC3AAD68" box="[833,917,1617,1643]" collectionName="Japan, Tokyo, National Science Museum (Natural History)" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
).
</materialsCitation>
<emphasis id="41E1EAF5F23FAC48FC58F9C2FAC4AD69" bold="true" box="[936,1387,1616,1643]" pageId="79" pageNumber="80">
<typeStatus id="AC2E8845F23FAC48FC58F9C2FB8BAD68" box="[936,1060,1617,1643]" pageId="79" pageNumber="80" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="0B827677F23FAC48FBC0F9C3FB34AD69" box="[1072,1179,1616,1642]" name="Russia" pageId="79" pageNumber="80">RUSSIA</collectingCountry>
:
<collectingRegion id="B151F805F23FAC48FB57F9C3FAC4AD69" box="[1191,1387,1616,1642]" country="Russia" name="Primorskiy" pageId="79" pageNumber="80">Primorskij Kraj</collectingRegion>
</emphasis>
:
<specimenCount id="6593FD6EF23FAC48FA89F9C2FA33AD68" box="[1401,1436,1617,1643]" pageId="79" pageNumber="80" type="female">1♀</specimenCount>
(DEI-GISHym 88016),
<collectorName id="DE605331F23FAC48FE6FF9E7FD87AD8D" box="[415,552,1652,1678]" pageId="79" pageNumber="80">Anisimovka</collectorName>
<quantity id="B46D9B02F23FAC48FDDFF9E6FDC8AD8D" box="[559,615,1652,1679]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="79" pageNumber="80" unit="km" value="7.0">7 km</quantity>
S,
<location id="764A603CF23FAC48FD7BF9E6FCBCAD8D" LSID="urn:lsid:plazi:treatment:FB3C87F1F23FAC57FF67FF04FD1CAF54:764A603CF23FAC48FD7BF9E6FCBCAD8D" box="[651,787,1652,1679]" country="Russia" latitude="43.102" longLatPrecision="68" longitude="132.786" name="Mt. Litovka" pageId="79" pageNumber="80" stateProvince="Primorskiy">Mt. Litovka</location>
,
<quantity id="B46D9B02F23FAC48FCEDF9E6FCC1AD8C" box="[797,878,1653,1679]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="79" pageNumber="80" unit="m" value="1200.0">
<elevation id="F8B8D1D4F23FAC48FCEDF9E6FCC1AD8C" box="[797,878,1653,1679]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="79" pageNumber="80" unit="m" value="1200.0">1200m</elevation>
</quantity>
,
<geoCoordinate id="16A15020F23FAC48FC89F9E6FC4AAD8C" box="[889,997,1653,1679]" degrees="43.102" direction="north" orientation="latitude" pageId="79" pageNumber="80" precision="55" value="43.102">43.102°N</geoCoordinate>
<geoCoordinate id="16A15020F23FAC48FC1CF9E7FBC9AD8D" box="[1004,1126,1652,1678]" degrees="132.786" direction="east" orientation="longitude" pageId="79" pageNumber="80" precision="55" value="132.786">132.786°E</geoCoordinate>
,
<date id="072B1027F23FAC48FB80F9E7FB50AD8D" box="[1136,1279,1652,1679]" pageId="79" pageNumber="80" value="2016-06-16">
16.
<collectionCode id="1584AE22F23FAC48FB69F9E6FB15AD8C" box="[1177,1210,1653,1679]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="79" pageNumber="80">VI</collectionCode>
. 2016
</date>
,
<collectionCode id="1584AE22F23FAC48FAF9F9E6FAB2AD8C" box="[1289,1309,1653,1679]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="79" pageNumber="80" type="Museum">V</collectionCode>
.
<materialsCitation id="C3FD3CBAF23FAC48FAD8F9E7FB6CADB0" ID-GBIF-Occurrence="3866537386" collectionCode="NSMT" collectorName="Loktionov &amp; Mirugam" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCount="2" specimenCount-male="2" stateProvince="Gangwon-do" typeStatus="paratype">
<collectorName id="DE605331F23FAC48FAD8F9E7FA32AD8D" box="[1320,1437,1652,1678]" pageId="79" pageNumber="80">Loktionov</collectorName>
(
<collectionCode id="1584AE22F23FAC48FF6FF90AFF5CADB0" box="[159,243,1689,1715]" collectionName="Japan, Tokyo, National Science Museum (Natural History)" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
).
<emphasis id="41E1EAF5F23FAC48FEF6F90BFD2EADB1" bold="true" box="[262,641,1688,1714]" pageId="79" pageNumber="80">
<collectingCountry id="0B827677F23FAC48FEF6F90BFE74ADB1" box="[262,475,1688,1714]" name="South Korea" pageId="79" pageNumber="80">SOUTH KOREA</collectingCountry>
:
<collectingRegion id="B151F805F23FAC48FE17F90BFD2EADB1" box="[487,641,1688,1714]" country="South Korea" name="Gangwon-do" pageId="79" pageNumber="80">Gangwon-do</collectingRegion>
</emphasis>
:
<specimenCount id="6593FD6EF23FAC48FD61F90AFD19ADB0" box="[657,694,1689,1715]" pageId="79" pageNumber="80" type="male">2♂</specimenCount>
,
<collectorName id="DE605331F23FAC48FD32F90BFC85ADB1" box="[706,810,1688,1714]" pageId="79" pageNumber="80">Mirugam</collectorName>
(
<collectingMunicipality id="934EAC9DF23FAC48FCC9F90BFC03ADB1" box="[825,940,1688,1714]" pageId="79" pageNumber="80">Bukdaesa</collectingMunicipality>
),
<quantity id="B46D9B02F23FAC48FC30F90AFBB1ADB0" box="[960,1054,1689,1715]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="79" pageNumber="80" unit="m" value="1300.0">
<elevation id="F8B8D1D4F23FAC48FC30F90AFBB1ADB0" box="[960,1054,1689,1715]" metricMagnitude="0" metricUnit="m" metricValue="1.3" pageId="79" pageNumber="80" unit="m" value="1.3">1,300 m</elevation>
</quantity>
,
<location id="764A603CF23FAC48FBDBF90BFB6CADB0" LSID="urn:lsid:plazi:treatment:FB3C87F1F23FAC57FF67FF04FD1CAF54:764A603CF23FAC48FBDBF90BFB6CADB0" box="[1067,1219,1688,1715]" country="South Korea" municipality="Bukdaesa" name="Odaesan Mts" pageId="79" pageNumber="80" stateProvince="Gangwon-do">Odaesan Mts</location>
</materialsCitation>
.,
<date id="072B1027F23FAC48FB24F90BFAF8ADB0" box="[1236,1367,1688,1715]" pageId="79" pageNumber="80" value="1997-06-06">
6.
<collectionCode id="1584AE22F23FAC48FB00F90AFABEADB0" box="[1264,1297,1689,1715]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="79" pageNumber="80">VI</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FAE0F90AFAF8ADB0" ID-GBIF-Occurrence="3866537478" box="[1296,1367,1689,1715]" collectionCode="NSMT" collectorName="Kim" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCount="1" specimenCount-male="1" stateProvince="Gangwon-do" typeStatus="paratype">. 1997</materialsCitation>
</date>
, J.-W.
<collectorName id="DE605331F23FAC48FF67F92FFF66ADD5" box="[151,201,1724,1750]" pageId="79" pageNumber="80">Kim</collectorName>
(
<collectionCode id="1584AE22F23FAC48FF28F92EFE83ADD4" box="[216,300,1725,1751]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
);
<specimenCount id="6593FD6EF23FAC48FECFF92EFECBADD4" box="[319,356,1725,1751]" pageId="79" pageNumber="80" type="male">1♂</specimenCount>
, same locality,
<date id="072B1027F23FAC48FDE2F92EFD3AADD4" box="[530,661,1725,1751]" pageId="79" pageNumber="80" value="2002-05-23">
23.
<collectionCode id="1584AE22F23FAC48FDCBF92EFDE0ADD4" box="[571,591,1725,1751]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="79" pageNumber="80" type="Museum">V</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FDA0F92EFD3AADD4" ID-GBIF-Occurrence="3866537427" box="[592,661,1725,1751]" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCount="1" specimenCount-male="1" stateProvince="Gangwon-do" typeStatus="paratype">. 2002</materialsCitation>
</date>
, A.
<collectorName id="DE605331F23FAC48FD31F92FFC9CADD5" box="[705,819,1724,1750]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FCB2F92EFC39ADD4" box="[834,918,1725,1751]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
);
<specimenCount id="6593FD6EF23FAC48FC59F92EFC62ADD4" box="[937,973,1725,1751]" pageId="79" pageNumber="80" type="male">1♂</specimenCount>
, same locality,
<date id="072B1027F23FAC48FB8BF92EFB50ADD4" box="[1147,1279,1725,1751]" pageId="79" pageNumber="80" value="2002-05-29">
29.
<collectionCode id="1584AE22F23FAC48FB55F92EFB16ADD4" box="[1189,1209,1725,1751]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="79" pageNumber="80" type="Museum">V</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FB49F92EFB50ADD4" ID-GBIF-Occurrence="3866537428" box="[1209,1279,1725,1751]" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCode="NSMT 30603" specimenCount="4" specimenCount-male="4" stateProvince="Gangwon-do" typeStatus="paratype">. 2002</materialsCitation>
</date>
, A.
<collectorName id="DE605331F23FAC48FADBF92FFA32ADD5" box="[1323,1437,1724,1750]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FF6FF972FF5CADF8" box="[159,243,1761,1787]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
);
<specimenCount id="6593FD6EF23FAC48FEF7F972FE85ADF8" box="[263,298,1761,1787]" pageId="79" pageNumber="80" type="male">4♂</specimenCount>
(incl.
<specimenCode id="23339E9CF23FAC48FE85F972FDBBADF9" box="[373,532,1760,1787]" collectionCode="NSMT" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT 30603</specimenCode>
), same locality,
<date id="072B1027F23FAC48FD3BF972FCE0ADF8" box="[715,847,1761,1787]" pageId="79" pageNumber="80" value="2002-06-04">
4.
<collectionCode id="1584AE22F23FAC48FD18F972FCA6ADF8" box="[744,777,1761,1787]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="79" pageNumber="80">VI</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FCF9F972FCE0ADF8" ID-GBIF-Occurrence="3866537360" box="[777,847,1761,1787]" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCode="NSMT 30747, 30748" specimenCount="2" specimenCount-male="2" stateProvince="Gangwon-do" typeStatus="paratype">. 2002</materialsCitation>
</date>
, A.
<collectorName id="DE605331F23FAC48FC8DF973FC40ADF9" box="[893,1007,1760,1786]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FC0FF972FBFCADF8" box="[1023,1107,1761,1787]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
);
<specimenCount id="6593FD6EF23FAC48FB97F972FB25ADF8" box="[1127,1162,1761,1787]" pageId="79" pageNumber="80" type="male">2♂</specimenCount>
(
<specimenCode id="23339E9CF23FAC48FB6AF972FA95ADF8" box="[1178,1338,1761,1787]" collectionCode="NSMT" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT 30747</specimenCode>
,
<specimenCode id="23339E9CF23FAC48FAB6F972FA3FADF8" box="[1350,1424,1761,1787]" collectionCode="NSMT" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">30748</specimenCode>
), same locality,
<date id="072B1027F23FAC48FECDF897FE6BAC1C" box="[317,452,1796,1823]" pageId="79" pageNumber="80" value="2008-05-26">
26.
<collectionCode id="1584AE22F23FAC48FE98F896FED3AC1C" box="[360,380,1797,1823]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="79" pageNumber="80" type="Museum">V</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FE8DF896FE6BAC1C" ID-GBIF-Occurrence="3866537399" box="[381,452,1797,1823]" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCode="NSMT 30850" specimenCount="1" specimenCount-male="1" stateProvince="Gangwon-do" typeStatus="paratype">. 2008</materialsCitation>
</date>
, A.
<collectorName id="DE605331F23FAC48FE05F897FDC8AC1D" box="[501,615,1796,1822]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FD88F896FD63AC1C" box="[632,716,1797,1823]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
);
<specimenCount id="6593FD6EF23FAC48FD11F896FCABAC1C" box="[737,772,1797,1823]" pageId="79" pageNumber="80" type="male">1♂</specimenCount>
(
<specimenCode id="23339E9CF23FAC48FCE5F896FC18AC1C" box="[789,951,1796,1823]" collectionCode="NSMT" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT 30850</specimenCode>
), same locality,
<date id="072B1027F23FAC48FB83F896FB55AC1C" box="[1139,1274,1797,1823]" pageId="79" pageNumber="80" value="2009-05-27">
27.
<collectionCode id="1584AE22F23FAC48FB6EF896FB1DAC1C" box="[1182,1202,1797,1823]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="79" pageNumber="80" type="Museum">V</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FB43F896FB55AC1C" ID-GBIF-Occurrence="3866537471" box="[1203,1274,1797,1823]" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCode="NSMT 30851" specimenCount="1" specimenCount-male="1" stateProvince="Gangwon-do" typeStatus="paratype">. 2009</materialsCitation>
</date>
, A.
<collectorName id="DE605331F23FAC48FADBF897FA32AC1D" box="[1323,1437,1796,1822]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FF6FF8BAFF5CAC40" box="[159,243,1833,1859]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
);
<specimenCount id="6593FD6EF23FAC48FEF6F8BAFE86AC40" box="[262,297,1833,1859]" pageId="79" pageNumber="80" type="male">1♂</specimenCount>
(
<specimenCode id="23339E9CF23FAC48FEC7F8BAFE78AC40" box="[311,471,1832,1859]" collectionCode="NSMT" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT 30851</specimenCode>
), same locality,
<date id="072B1027F23FAC48FD7AF8BAFCA2AC40" box="[650,781,1833,1859]" pageId="79" pageNumber="80" value="2009-05-31">
31.
<collectionCode id="1584AE22F23FAC48FD43F8BAFD68AC40" box="[691,711,1833,1859]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="79" pageNumber="80" type="Museum">V</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FD38F8BAFCA2AC40" ID-GBIF-Occurrence="3866537397" box="[712,781,1833,1859]" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCode="NSMT 30852" specimenCount="1" specimenCount-male="1" stateProvince="Gangwon-do" typeStatus="paratype">. 2009</materialsCitation>
</date>
, A.
<collectorName id="DE605331F23FAC48FCC8F8BBFC05AC41" box="[824,938,1832,1858]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FC48F8BAFBA3AC40" box="[952,1036,1833,1859]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
);
<specimenCount id="6593FD6EF23FAC48FBEFF8BAFBEDAC40" box="[1055,1090,1833,1859]" pageId="79" pageNumber="80" type="male">1♂</specimenCount>
(
<specimenCode id="23339E9CF23FAC48FBA0F8BAFB5FAC40" box="[1104,1264,1832,1859]" collectionCode="NSMT" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT 30852</specimenCode>
), same locality,
<date id="072B1027F23FAC48FF67F8DEFEB4AC64" box="[151,283,1869,1895]" pageId="79" pageNumber="80" value="2009-06-04">
4.
<collectionCode id="1584AE22F23FAC48FF44F8DEFF7AAC64" box="[180,213,1869,1895]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="79" pageNumber="80">VI</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FF25F8DEFEB4AC64" ID-GBIF-Occurrence="3866537307" box="[213,283,1869,1895]" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCount="1" specimenCount-male="1" stateProvince="Gangwon-do" typeStatus="paratype">. 2009</materialsCitation>
</date>
, A.
<collectorName id="DE605331F23FAC48FEB9F8DFFE14AC65" box="[329,443,1868,1894]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FE3BF8DEFDB0AC64" box="[459,543,1869,1895]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
);
<specimenCount id="6593FD6EF23FAC48FDC3F8DEFDF7AC64" box="[563,600,1869,1895]" pageId="79" pageNumber="80" type="male">1♂</specimenCount>
, same locality,
<date id="072B1027F23FAC48FCF8F8DFFC23AC64" box="[776,908,1868,1895]" pageId="79" pageNumber="80" value="2009-06-05">
5.
<collectionCode id="1584AE22F23FAC48FCD5F8DEFCE9AC64" box="[805,838,1869,1895]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="79" pageNumber="80">VI</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FCB6F8DEFC23AC64" ID-GBIF-Occurrence="3866537339" box="[838,908,1869,1895]" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCount="1" specimenCount-male="1" stateProvince="Gangwon-do" typeStatus="paratype">. 2009</materialsCitation>
</date>
, A.
<collectorName id="DE605331F23FAC48FC4AF8DFFB83AC65" box="[954,1068,1868,1894]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FBCCF8DEFB3FAC64" box="[1084,1168,1869,1895]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
);
<specimenCount id="6593FD6EF23FAC48FB54F8DEFB66AC64" box="[1188,1225,1869,1895]" pageId="79" pageNumber="80" type="male">1♂</specimenCount>
, same locality,
<date id="072B1027F23FAC48FA89F8DEFF51AC88" pageId="79" pageNumber="80" value="2010-06-13">
13.
<collectionCode id="1584AE22F23FAC48FF67F8E2FF17AC88" box="[151,184,1905,1931]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="79" pageNumber="80">VI</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FF48F8E2FF51AC88" ID-GBIF-Occurrence="3866537378" box="[184,254,1905,1931]" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" elevation="1" location="Odaesan Mts" municipality="Bukdaesa" pageId="79" pageNumber="80" specimenCount="1" specimenCount-male="1" stateProvince="Gangwon-do" typeStatus="paratype">. 2010</materialsCitation>
</date>
, A.
<collectorName id="DE605331F23FAC48FED9F8E3FE34AC89" box="[297,411,1904,1930]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FE5AF8E2FE51AC88" box="[426,510,1905,1931]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
);
<specimenCount id="6593FD6EF23FAC48FDE1F8E2FD99AC88" box="[529,566,1905,1931]" pageId="79" pageNumber="80" type="male">1♂</specimenCount>
, same locality,
<date id="072B1027F23FAC48FD13F8E2FCDCAC88" box="[739,883,1905,1931]" pageId="79" pageNumber="80" value="2010-06-17">
17.
<collectionCode id="1584AE22F23FAC48FCFCF8E2FC82AC88" box="[780,813,1905,1931]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="79" pageNumber="80">VI</collectionCode>
<materialsCitation id="C3FD3CBAF23FAC48FCDDF8E2FCDCAC88" ID-GBIF-Occurrence="3866537348" box="[813,883,1905,1931]" collectionCode="NSMT" collectorName="Shinohara" country="South Korea" pageId="79" pageNumber="80" specimenCount="1" stateProvince="Gangwon-do" typeStatus="paratype">. 2010</materialsCitation>
</date>
, A.
<collectorName id="DE605331F23FAC48FC6FF8E3FBBEAC89" box="[927,1041,1904,1930]" pageId="79" pageNumber="80">Shinohara</collectorName>
(
<collectionCode id="1584AE22F23FAC48FBEFF8E2FBDCAC88" box="[1055,1139,1905,1931]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="79" pageNumber="80">NSMT</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF23FAC48FF37F807FCB2ACAC" box="[199,797,1940,1967]" pageId="79" pageNumber="80" type="distribution">
<paragraph id="732A36E7F23FAC48FF37F807FCB2ACAC" blockId="79.[151,1437,356,2039]" box="[199,797,1940,1967]" pageId="79" pageNumber="80">
<emphasis id="41E1EAF5F23FAC48FF37F807FEF6ACAD" bold="true" box="[199,345,1940,1966]" pageId="79" pageNumber="80">Distribution</emphasis>
.
<collectingCountry id="0B827677F23FAC48FE97F807FE1DACAD" box="[359,434,1940,1966]" name="Russia" pageId="79" pageNumber="80">Russia</collectingCountry>
(
<collectingRegion id="B151F805F23FAC48FE30F807FDD9ACAD" box="[448,630,1940,1966]" country="Russia" name="Primorskiy" pageId="79" pageNumber="80">Primorskij Kraj</collectingRegion>
),
<collectingCountry id="0B827677F23FAC48FD79F807FCB6ACAC" box="[649,793,1940,1967]" name="South Korea" pageId="79" pageNumber="80">South Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF23FAC48FF37F82AFF4FACF4" pageId="79" pageNumber="80" type="biology_ecology">
<paragraph id="732A36E7F23FAC48FF37F82AFF4FACF4" blockId="79.[151,1437,356,2039]" pageId="79" pageNumber="80">
<emphasis id="41E1EAF5F23FAC48FF37F82AFEE8ACD0" bold="true" box="[199,327,1977,2003]" pageId="79" pageNumber="80">Host plant</emphasis>
. Unknown. The closely related Japanese species
<taxonomicName id="B4954D64F23FAC48FC72F82AFC58ACD1" authorityName=": Shinohara" authorityYear="2001" box="[898,1015,1976,2002]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F23FAC48FC72F82AFC58ACD1" box="[898,1015,1976,2002]" italics="true" pageId="79" pageNumber="80">P. gracilis</emphasis>
</taxonomicName>
feeds on
<taxonomicName id="B4954D64F23FAC48FB9AF82BFF73ACF4" authority="(Shinohara &amp; Hara 1992)" baseAuthorityName="Shinohara &amp; Hara" baseAuthorityYear="1992" class="Magnoliopsida" family="Rosaceae" genus="Sorbus" kingdom="Plantae" order="Rosales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="genus">
<emphasis id="41E1EAF5F23FAC48FB9AF82BFB17ACD1" box="[1130,1208,1976,2002]" italics="true" pageId="79" pageNumber="80">Sorbus</emphasis>
(
<bibRefCitation id="17044B16F23FAC48FB39F82BFF7CACF4" author="Shinohara, A. &amp; Hara, H." pageId="79" pageNumber="80" pagination="432" refId="ref77480" refString="Shinohara, A. &amp; Hara, H. (1992) A new host record for Pamphilius gracilis (Hymenoptera, Pamphiliidae). Japanese Journal of Entomology, Tokyo, 60, 432." type="journal article" year="1992">Shinohara &amp; Hara 1992</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF220AC57FF37FF04FA37ABB2" box="[199,1432,151,177]" pageId="80" pageNumber="81" type="etymology">
<paragraph id="732A36E7F220AC57FF37FF04FA37ABB2" blockId="80.[151,1437,151,1111]" box="[199,1432,151,177]" pageId="80" pageNumber="81">
<emphasis id="41E1EAF5F220AC57FF37FF04FEE1ABB2" bold="true" box="[199,334,151,177]" pageId="80" pageNumber="81">Etymology.</emphasis>
The new specific epithet is
<taxonomicName id="B4954D64F220AC57FD77FF04FD72ABB2" authorityName=": Shinohara" authorityYear="2001" box="[647,733,151,177]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FD77FF04FD72ABB2" box="[647,733,151,177]" italics="true" pageId="80" pageNumber="81">gracilis</emphasis>
</taxonomicName>
plus a suffix, -
<emphasis id="41E1EAF5F220AC57FC75FF04FC6FABB2" box="[901,960,151,177]" italics="true" pageId="80" pageNumber="81">oides</emphasis>
, suggesting a species resembling
<taxonomicName id="B4954D64F220AC57FACCFF04FA3DABB2" authorityName=": Shinohara" authorityYear="2001" box="[1340,1426,151,177]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FACCFF04FA3DABB2" box="[1340,1426,151,177]" italics="true" pageId="80" pageNumber="81">gracilis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF220AC57FF37FF2FFD1CAF54" pageId="80" pageNumber="81" type="discussion">
<paragraph id="732A36E7F220AC57FF37FF2FFBE6AA44" blockId="80.[151,1437,151,1111]" pageId="80" pageNumber="81">
<emphasis id="41E1EAF5F220AC57FF37FF2FFE9BABD5" bold="true" box="[199,308,188,214]" pageId="80" pageNumber="81">Remarks</emphasis>
.
<taxonomicName id="B4954D64F220AC57FEB5FF2FFDE3ABD5" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[325,588,188,214]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FEB5FF2FFDE3ABD5" box="[325,588,188,214]" italics="true" pageId="80" pageNumber="81">Pamphilius graciloides</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="B4954D64F220AC57FCD8FF2EFBCDABD4" authority="Shinohara, 1985" authorityName="Shinohara" authorityYear="1985" box="[808,1122,188,215]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FCD8FF2EFC33ABD5" box="[808,924,188,214]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
Shinohara, 1985
</taxonomicName>
, from
<collectingCountry id="0B827677F220AC57FB5FFF2EFB40ABD4" box="[1199,1263,189,215]" name="Japan" pageId="80" pageNumber="81">Japan</collectingCountry>
and
<bibRefCitation id="17044B16F220AC57FADBFF2FFF4DABF8" author="Shinohara, A." pageId="80" pageNumber="81" pagination="99 - 116" refId="ref76721" refString="Shinohara, A. (2001) The group of Pamphilius sylvaticus (Hymenoptera, Pamphiliidae): Two new species, new collection records, and a key to Palearctic species. Japanese Journal of Systematic Entomology, 7, 99 - 116." type="journal article" year="2001">Shinohara (2001)</bibRefCitation>
recorded it as “
<taxonomicName id="B4954D64F220AC57FE61FF71FDADABF8" authorityName="Shinohara" authorityYear="1985" box="[401,514,225,251]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FE61FF71FDADABF8" box="[401,514,225,251]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
” from
<collectingCountry id="0B827677F220AC57FDA1FF72FD4EABFF" box="[593,737,225,252]" name="South Korea" pageId="80" pageNumber="81">South Korea</collectingCountry>
. Because of the existence of recognizable, though small differences in colour pattern in both sexes and quite a large difference in COI sequences, we here treat the continental and Japanese populations as representing different species,
<taxonomicName id="B4954D64F220AC57FCF1FEBEFC35AA45" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[769,922,300,326]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FCF1FEBEFC35AA45" box="[769,922,300,326]" italics="true" pageId="80" pageNumber="81">P. graciloides</emphasis>
</taxonomicName>
and
<taxonomicName id="B4954D64F220AC57FC21FEBEFBEDAA45" authorityName="Shinohara" authorityYear="1985" box="[977,1090,300,326]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FC21FEBEFBEDAA45" box="[977,1090,300,326]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="732A36E7F220AC57FF37FEC2FB40A902" blockId="80.[151,1437,151,1111]" pageId="80" pageNumber="81">
<bibRefCitation id="17044B16F220AC57FF37FEC2FE23AA68" author="Shinohara, A." box="[199,396,337,363]" pageId="80" pageNumber="81" pagination="99 - 116" refId="ref76721" refString="Shinohara, A. (2001) The group of Pamphilius sylvaticus (Hymenoptera, Pamphiliidae): Two new species, new collection records, and a key to Palearctic species. Japanese Journal of Systematic Entomology, 7, 99 - 116." type="journal article" year="2001">Shinohara (2001)</bibRefCitation>
discussed the variation of “
<taxonomicName id="B4954D64F220AC57FD3AFEC1FC93AA68" authorityName="Shinohara" authorityYear="1985" box="[714,828,337,363]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FD3AFEC1FC93AA68" box="[714,828,337,363]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
” in detail and pointed out that the three Korean male specimens then available (now identified as
<taxonomicName id="B4954D64F220AC57FD75FEE4FCB0AA92" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[645,799,375,401]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FD75FEE4FCB0AA92" box="[645,799,375,401]" italics="true" pageId="80" pageNumber="81">P. graciloides</emphasis>
</taxonomicName>
) were darker in colour pattern than the 17 Japanese male specimens examined (
<taxonomicName id="B4954D64F220AC57FE60FE0EFDADAAB5" authorityName="Shinohara" authorityYear="1985" box="[400,514,412,438]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FE60FE0EFDADAAB5" box="[400,514,412,438]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
). Examination of the additional
<specimenCount id="6593FD6EF220AC57FC9EFE0FFC7EAAB5" box="[878,977,412,438]" pageId="80" pageNumber="81" type="male">11 males</specimenCount>
from
<collectingCountry id="0B827677F220AC57FBE6FE0EFBF5AAB4" box="[1046,1114,413,439]" name="South Korea" pageId="80" pageNumber="81">Korea</collectingCountry>
listed above (
<taxonomicName id="B4954D64F220AC57FB09FE0EFA3CAAB5" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[1273,1427,412,438]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FB09FE0EFA3CAAB5" box="[1273,1427,412,438]" italics="true" pageId="80" pageNumber="81">P. graciloides</emphasis>
</taxonomicName>
) has shown that the differences in color pattern are fairly stable. The female of
<taxonomicName id="B4954D64F220AC57FBD7FE51FB6CAAD8" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[1063,1219,449,475]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FBD7FE51FB6CAAD8" box="[1063,1219,449,475]" italics="true" pageId="80" pageNumber="81">P. graciloides</emphasis>
</taxonomicName>
(only one Russian specimen available) is slightly different in colour pattern from those of
<taxonomicName id="B4954D64F220AC57FC47FE74FB87A902" authorityName="Shinohara" authorityYear="1985" box="[951,1064,487,513]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FC47FE74FB87A902" box="[951,1064,487,513]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
, as shown below.
</paragraph>
<paragraph id="732A36E7F220AC57FF37FD9EFB94A804" blockId="80.[151,1437,151,1111]" pageId="80" pageNumber="81">
In the analysis of COI sequences (
<figureCitation id="EBAE2A62F220AC57FDA7FD9FFD14A924" box="[599,699,524,551]" captionStart="FIGURE 145" captionStartId="232.[151,250,1669,1693]" captionTargetBox="[157,1428,190,1638]" captionTargetId="figure-26@232.[151,1436,181,1645]" captionTargetPageId="232" captionText="FIGURE 145. Condensed ML tree based on CO1 gene sequences, with details of Pamphilius sylvaticus group, part 1." figureDoi="http://doi.org/10.5281/zenodo.6876851" httpUri="https://zenodo.org/record/6876851/files/figure.png" pageId="80" pageNumber="81">Fig. 145</figureCitation>
), each of
<taxonomicName id="B4954D64F220AC57FCC1FD9EFC62A925" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[817,973,524,550]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FCC1FD9EFC62A925" box="[817,973,524,550]" italics="true" pageId="80" pageNumber="81">P. graciloides</emphasis>
</taxonomicName>
and
<taxonomicName id="B4954D64F220AC57FBF9FD9EFBD1A925" authorityName="Shinohara" authorityYear="1985" box="[1033,1150,524,550]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FBF9FD9EFBD1A925" box="[1033,1150,524,550]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
was retrieved as a clade with 100% UFBoot support and they formed a clade with 100% UFBoot support. The minimum distance (4.0%) between the two species was much larger than the interspecific distance between closely related congeners, such as
<taxonomicName id="B4954D64F220AC57FF67FDEEFE87A995" box="[151,296,636,662]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="albopictus">
<emphasis id="41E1EAF5F220AC57FF67FDEEFE87A995" box="[151,296,636,662]" italics="true" pageId="80" pageNumber="81">P. albopictus</emphasis>
</taxonomicName>
and
<taxonomicName id="B4954D64F220AC57FEAFFDEEFD7EA995" authority="Takeuchi, 1930" authorityName="Takeuchi" authorityYear="1930" box="[351,721,636,662]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="kamikochensis">
<emphasis id="41E1EAF5F220AC57FEAFFDEEFDB2A995" box="[351,541,636,662]" italics="true" pageId="80" pageNumber="81">P. kamikochensis</emphasis>
<bibRefCitation id="17044B16F220AC57FDD4FDEFFD7EA995" author="Takeuchi, K." box="[548,721,636,662]" pageId="80" pageNumber="81" pagination="3 - 16" refId="ref80368" refString="Takeuchi, K. (1930) A revisional list of the Japanese Pamphiliidae, with description of nine new species. Transactions of the Kansai Entomological Society, 1, 3 - 16." type="journal article" year="1930">Takeuchi, 1930</bibRefCitation>
</taxonomicName>
(2.0%) or
<taxonomicName id="B4954D64F220AC57FCBBFDEEFC00A995" authorityName="Shinohara" authorityYear="1985" box="[843,943,636,662]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="benesi">
<emphasis id="41E1EAF5F220AC57FCBBFDEEFC00A995" box="[843,943,636,662]" italics="true" pageId="80" pageNumber="81">P. benesi</emphasis>
</taxonomicName>
and
<taxonomicName id="B4954D64F220AC57FC15FDEEFBF4A995" authorityName="Shinohara" authorityYear="1988" box="[997,1115,636,662]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="41E1EAF5F220AC57FC15FDEEFBF4A995" box="[997,1115,636,662]" italics="true" pageId="80" pageNumber="81">P. pallidus</emphasis>
</taxonomicName>
(2.9%). In the NaK analysis (
<figureCitation id="EBAE2A62F220AC57FF6FFD32FEABA9BF" box="[159,260,673,700]" captionStart="FIGURE 159" captionStartId="244.[151,250,1756,1780]" captionTargetBox="[157,1427,187,1724]" captionTargetId="figure-26@244.[151,1436,181,1733]" captionTargetPageId="244" captionText="FIGURE 159. Condensed ML tree based on NaK gene sequences, with details of Pamphilius sylvaticus group." figureDoi="http://doi.org/10.5281/zenodo.6876882" httpUri="https://zenodo.org/record/6876882/files/figure.png" pageId="80" pageNumber="81">Fig. 159</figureCitation>
), the available specimens of
<taxonomicName id="B4954D64F220AC57FDA6FD31FD5DA9B8" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[598,754,673,699]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FDA6FD31FD5DA9B8" box="[598,754,673,699]" italics="true" pageId="80" pageNumber="81">P. graciloides</emphasis>
</taxonomicName>
and
<taxonomicName id="B4954D64F220AC57FCDEFD31FC0CA9B8" authorityName="Shinohara" authorityYear="1985" box="[814,931,673,699]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FCDEFD31FC0CA9B8" box="[814,931,673,699]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
were not separable but altogether formed a monophyletic group with 100% UFBoot support.
<taxonomicName id="B4954D64F220AC57FD4DFD54FC6FA9E2" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[701,960,711,737]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FD4DFD54FC6FA9E2" box="[701,960,711,737]" italics="true" pageId="80" pageNumber="81">Pamphilius graciloides</emphasis>
</taxonomicName>
and
<taxonomicName id="B4954D64F220AC57FC04FD54FBCBA9E2" authorityName="Shinohara" authorityYear="1985" box="[1012,1124,711,737]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FC04FD54FBCBA9E2" box="[1012,1124,711,737]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
clearly show a large genetic differentiation in COI, though they are not clearly differentiated in NaK sequences.
</paragraph>
<paragraph id="732A36E7F220AC57FF37FC82FD1CAF54" blockId="80.[151,1437,151,1111]" pageId="80" pageNumber="81">
The female of
<taxonomicName id="B4954D64F220AC57FE9FFC81FDA5A828" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[367,522,785,811]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FE9FFC81FDA5A828" box="[367,522,785,811]" italics="true" pageId="80" pageNumber="81">P. graciloides</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="B4954D64F220AC57FD52FC81FCBAA828" authorityName="Shinohara" authorityYear="1985" box="[674,789,785,811]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FD52FC81FCBAA828" box="[674,789,785,811]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
, but the base of the fore femur, including the trochantellus, is almost all black, and the mesepisternum has only a small pale yellow spot on the anterodorsal corner. In
<taxonomicName id="B4954D64F220AC57FA78FCA4FF42A875" authorityName="Shinohara" authorityYear="1985" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FA78FCA4FF42A875" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
, the base of the fore femur is also blackish but the trochantellus is mostly pale yellow and the entire anterior margin of the mesepisternum is pale yellow. The male of
<taxonomicName id="B4954D64F220AC57FCD6FC11FC6EA898" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[806,961,897,923]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FCD6FC11FC6EA898" box="[806,961,897,923]" italics="true" pageId="80" pageNumber="81">P. graciloides</emphasis>
</taxonomicName>
has the mesepisternum mostly or entirely black with at most small pale-yellow marks and the malar space medially marked with black, whereas
<taxonomicName id="B4954D64F220AC57FADAFC34FA32A8C2" authorityName="Shinohara" authorityYear="1985" box="[1322,1437,935,961]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FADAFC34FA32A8C2" box="[1322,1437,935,961]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
has the mesepisternum largely marked with pale yellow and the malar space entirely pale yellow. One specimen of
<taxonomicName id="B4954D64F220AC57FF67FC61FEA7AF08" authorityName="Shinohara" authorityYear="1985" box="[151,264,1009,1035]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="41E1EAF5F220AC57FF67FC61FEA7AF08" box="[151,264,1009,1035]" italics="true" pageId="80" pageNumber="81">P. gracilis</emphasis>
</taxonomicName>
from Shiga-kogen, Honshu, which was referred to by
<bibRefCitation id="17044B16F220AC57FC93FC62FB87AF08" author="Shinohara, A." box="[867,1064,1009,1035]" pageId="80" pageNumber="81" pagination="99 - 116" refId="ref76721" refString="Shinohara, A. (2001) The group of Pamphilius sylvaticus (Hymenoptera, Pamphiliidae): Two new species, new collection records, and a key to Palearctic species. Japanese Journal of Systematic Entomology, 7, 99 - 116." type="journal article" year="2001">Shinohara (2001)</bibRefCitation>
, has an exceptionally dark colour pattern and is almost indistinguishable from
<taxonomicName id="B4954D64F220AC57FD7AFB84FC8CAF32" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[650,803,1047,1073]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FD7AFB84FC8CAF32" box="[650,803,1047,1073]" italics="true" pageId="80" pageNumber="81">P. graciloides</emphasis>
</taxonomicName>
. The maximum intraspecific
<emphasis id="41E1EAF5F220AC57FB9CFB84FBD5AF33" box="[1132,1146,1047,1072]" italics="true" pageId="80" pageNumber="81">p</emphasis>
-distance in
<taxonomicName id="B4954D64F220AC57FAF3FB84FA33AF32" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[1283,1436,1047,1073]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F220AC57FAF3FB84FA33AF32" box="[1283,1436,1047,1073]" italics="true" pageId="80" pageNumber="81">P. graciloides</emphasis>
</taxonomicName>
was 0.8% in COI (n=7) and 0.2% in NaK (n=5).
</paragraph>
</subSubSection>
</treatment>
</document>