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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.20.4344" ID-GBIF-Dataset="5aeaed8f-855a-47aa-a72b-588b29d98ef8" ID-PMC="PMC3652410" ID-Pensoft-Pub="1314-2003-20-17" ID-Pensoft-UUID="FFFDFF9BFFDFDE546166FFE4D472FFD1" ID-PubMed="23717192" ID-Zenodo-Dep="576158" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1314-2003-20-17" ModsDocOrigin="PhytoKeys 20" ModsDocTitle="A revised circumscription for the Blakeeae (Melastomataceae) with associated nomenclatural adjustments" checkinTime="1555332075685" checkinUser="pensoft" docAuthor="Penneys, Darin S. &amp; Judd, Walter S." docDate="2013" docId="FAAA538068FD5A3EF474AAFF7B9B19C2" docLanguage="en" docName="PhytoKeys 20: 17-32" docOrigin="PhytoKeys 20" docSource="http://dx.doi.org/10.3897/phytokeys.20.4344" docTitle="Chalybea Naudin, Ann. Sci. Nat., Bot. ser. 3 16: 99. 1851" docType="treatment" docVersion="6" id="FFFDFF9BFFDFDE546166FFE4D472FFD1" lastPageNumber="20" masterDocId="FFFDFF9BFFDFDE546166FFE4D472FFD1" masterDocTitle="A revised circumscription for the Blakeeae (Melastomataceae) with associated nomenclatural adjustments" masterLastPageNumber="32" masterPageNumber="17" pageNumber="20" updateTime="1668140735209" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revised circumscription for the Blakeeae (Melastomataceae) with associated nomenclatural adjustments</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Penneys, Darin S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Judd, Walter S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>20</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>17</mods:start>
<mods:end>32</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.20.4344</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.20.4344</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-20-17</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFFDFF9BFFDFDE546166FFE4D472FFD1</mods:identifier>
<mods:identifier type="Zenodo-Dep">576158</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="156201567" LSID="urn:lsid:plazi:treatment:FAAA538068FD5A3EF474AAFF7B9B19C2" httpUri="http://treatment.plazi.org/id/FAAA538068FD5A3EF474AAFF7B9B19C2" lastPageNumber="20" pageId="3" pageNumber="20">
<subSubSection pageId="3" pageNumber="20" type="nomenclature">
<paragraph pageId="3" pageNumber="20">
<taxonomicName LSID="F154FE85-2D66-5A52-8248-A3CE122988AE" authority="Naudin, Ann. Sci. Nat., Bot. ser. 3 16: 99. 1851." class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Chalybea" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
Chalybea Naudin, Ann. Sci. Nat., Bot.
<normalizedToken originalValue="sér">ser</normalizedToken>
. 3 16: 99. 1851.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="20" type="reference_group">
<paragraph pageId="3" pageNumber="20">
<taxonomicName authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">Huilaea</taxonomicName>
Wurdack, Brittonia 9: 106. 1957. Type:
<taxonomicName authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="species" species="penduliflora">
<emphasis italics="true" pageId="3" pageNumber="20">Huilaea penduliflora</emphasis>
</taxonomicName>
Wurdack, Brittonia 9: 106. 1957.
<emphasis bold="true" pageId="3" pageNumber="20">TYPE. COLOMBIA.</emphasis>
<emphasis bold="true" pageId="3" pageNumber="20">Huila:</emphasis>
Cordillera Oriental, wet temperate forest of deep moist canyon near camp on north side of
<normalizedToken originalValue="Río">Rio</normalizedToken>
Venadito, 25 km SE of La Bodega, 2450 m, 1 Dec 1944, E.Little 9101 (holotype: US!; isotypes: COL!, NY!).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="20" type="type">
<paragraph pageId="3" pageNumber="20">Type.</paragraph>
<paragraph pageId="3" pageNumber="20">
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="species" species="corymbifera">
<emphasis italics="true" pageId="3" pageNumber="20">Chalybea corymbifera</emphasis>
</taxonomicName>
Naudin, Ann. Sci. Nat., Bot.
<normalizedToken originalValue="sér">ser</normalizedToken>
. 3 16: 100. 1851. TYPE. COLOMBIA. Santander: Pamplona, La Baja, ca. 3000 m, Dec 1846, Funck &amp; Schlim 1312 (holotype: P!, photos at NY!, US!; isotypes: BM!, BR! (2 sheets), G!, photos of G sheet at NY!, US!).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="20" type="remarks">
<paragraph pageId="3" pageNumber="20">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="20">
<emphasis bold="true" pageId="3" pageNumber="20">Phylogenetic analyses</emphasis>
(
<bibRefCitation author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="12" pageNumber="29" refId="B12" refString="Morales-P, ME, 2010. Analisis filogenetico de Huilaea Wurdack (Melastomataceae) basado en datos morfologicos y moleculares. PhD Thesis, Universidad Nacional de Colombia, Colombia." title="Analisis filogenetico de Huilaea Wurdack (Melastomataceae) basado en datos morfologicos y moleculares. PhD Thesis, Universidad Nacional de Colombia, Colombia." year="2010">Morales-P 2010</bibRefCitation>
,
<bibRefCitation author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="13" pageNumber="30" publicationUrl="10.1007/s12228-009-9092-x" refId="B15" refString="Morales-P, ME, Penneys, DS, Judd, W, Gonzalez, F, submitted. Revision taxonomica del clado Chalybea/Huilaea (Melastomataceae). Caldasia., 10.1007/s12228-009-9092-x" title="Revision taxonomica del clado Chalybea / Huilaea (Melastomataceae). Caldasia" url="10.1007/s12228-009-9092-x" year="submitted">Morales-P et al. submitted</bibRefCitation>
,
<bibRefCitation author="Penneys, DS" journalOrPublisher="Caldasia" pageId="13" pageNumber="30" refId="B19" refString="Penneys, DS, Judd, WS, 2010. Phylogeny and character evolution in the Blakeeae (Melastomataceae). Abstract ID no. 699. Botany 2010. July 31 -August 4. Providence, Rhode Island, USA. http://2010.botanyconference.org/engine/search/index.php?func=detail&amp;aid=699." title="Phylogeny and character evolution in the Blakeeae (Melastomataceae). Abstract ID no. 699. Botany 2010. July 31 - August 4. Providence, Rhode Island, USA. http: // 2010. botanyconference. org / engine / search / index. php? func = detail &amp; aid = 699" year="2010">Penneys and Judd 2010</bibRefCitation>
) indicate that
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Huilaea</emphasis>
</taxonomicName>
form a strongly-supported clade. However, as
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
is nested within
<taxonomicName authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Huilaea</emphasis>
</taxonomicName>
, species belonging to the latter genus must be transferred because
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
has nomenclatural priority.
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
has been distinguished from
<taxonomicName authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Huilaea</emphasis>
</taxonomicName>
by having inflorescences with 21-39 flowers (vs. 3-17 in
<taxonomicName authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Huilaea</emphasis>
</taxonomicName>
), the flowers 13-20 mm long (vs. 45-55 mm long), the hypanthium 7-10 mm long and 8-10 mm wide (vs. 13-31 mm long x 11-19 mm wide), the petals 12-14 mm long and 5-7 mm wide (vs. 26-44 mm long x 10-22 mm wide), and white to cream or green, sometimes
<normalizedToken originalValue="pink–tinged">pink-tinged</normalizedToken>
apically (vs. pink to red, paler inside), the anthers 3-4 mm long (vs. 7-10 mm long), and the style 10-12 mm long (vs. 22-33 mm long).
</paragraph>
<paragraph pageId="3" pageNumber="20">
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
includes ten described species of small, South American trees: seven endemic to Colombia, two to Ecuador, and one to Peru. All are found in the Andes, except one species that is restricted to the Sierra Nevada de Santa Marta, Colombia.
</paragraph>
<paragraph pageId="3" pageNumber="20">
For more information on the systematics of
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
(and
<taxonomicName authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="20">Huilaea</emphasis>
</taxonomicName>
), see
<bibRefCitation author="Lozano-Contreras, G" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="12" pageNumber="29" refId="B10" refString="Lozano-Contreras, G, Ruiz-R, N, 1996. El genero Huilaea Wurdack (Melastomataceae). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 20: 237-242. http://www.accefyn.org.co/revista/Vol_20/77/237-242.pdf." title="El genero Huilaea Wurdack (Melastomataceae). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 20: 237 - 242. http: // www. accefyn. org. co / revista / Vol _ 20 / 77 / 237 - 242. pdf" year="1996">Lozano-Contreras and Ruiz-R (1996)</bibRefCitation>
,
<bibRefCitation author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="13" pageNumber="30" refId="B13" refString="Morales-P, ME, Gonzalez, F, 2005. Redescubrimiento de Chalybea Naudin e implicaciones en la delimitacion generica de Huilaea Wurdack (Melastomataceae). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 29: 171-178. http://www.accefyn.org.co/revista/Vol_29/111/03_171_178.pdf." title="Redescubrimiento de Chalybea Naudin e implicaciones en la delimitacion generica de Huilaea Wurdack (Melastomataceae). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 29: 171 - 178. http: // www. accefyn. org. co / revista / Vol _ 29 / 111 / 03 _ 171 _ 178. pdf" year="2005">
Morales-P and
<normalizedToken originalValue="González">Gonzalez</normalizedToken>
(2005)
</bibRefCitation>
,
<bibRefCitation author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="12" pageNumber="29" refId="B12" refString="Morales-P, ME, 2010. Analisis filogenetico de Huilaea Wurdack (Melastomataceae) basado en datos morfologicos y moleculares. PhD Thesis, Universidad Nacional de Colombia, Colombia." title="Analisis filogenetico de Huilaea Wurdack (Melastomataceae) basado en datos morfologicos y moleculares. PhD Thesis, Universidad Nacional de Colombia, Colombia." year="2010">Morales-P (2010)</bibRefCitation>
,
<bibRefCitation author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="13" pageNumber="30" refId="B14" refString="Morales-P, ME, Penneys, DS, 2010. New species of Chalybea Naudin and Huilaea Wurdack. Brittonia 62: 26-34. http://www.springerlink.com/content/47873303k2k84836/." title="New species of Chalybea Naudin and Huilaea Wurdack. Brittonia 62: 26 - 34. http: // www. springerlink. com / content / 47873303 k 2 k 84836 /" year="2010">Morales-P and Penneys (2010)</bibRefCitation>
, and
<bibRefCitation author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="13" pageNumber="30" publicationUrl="10.1007/s12228-009-9092-x" refId="B15" refString="Morales-P, ME, Penneys, DS, Judd, W, Gonzalez, F, submitted. Revision taxonomica del clado Chalybea/Huilaea (Melastomataceae). Caldasia., 10.1007/s12228-009-9092-x" title="Revision taxonomica del clado Chalybea / Huilaea (Melastomataceae). Caldasia" url="10.1007/s12228-009-9092-x" year="submitted">Morales-P et al. (submitted)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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