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<document id="9C18E7E961EF9F80EBADF1A444AAD84C" ID-DOI="10.5281/zenodo.275382" ID-GBIF-Dataset="fea16b1e-514a-45dc-9264-74ac141d7b40" ID-ISSN="1175-5326" ID-Zenodo-Dep="275382" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461133638375" checkinUser="plazi" docAuthor="Zhang, Feng, Fu, Jianying &amp; Zhu, Mingsheng" docDate="2009" docId="B50B87B11E71015598F3FF3C3DF92F74" docLanguage="en" docName="zt02305p060.pdf" docOrigin="Zootaxa 2305" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Systaria mengla Song &amp; Zhu 1994, comb. nov." docType="treatment" docVersion="8" lastPageNumber="59" masterDocId="4932FFC91E77015D9864FFAB3E3D295C" masterDocTitle="First report of the sac spider genus Systaria Simon (Araneae: Clubionidae) from China: Systaria hainanensis sp. nov. and Systaria mengla (Song &amp; Zhu, 1994) comb. nov." masterLastPageNumber="60" masterPageNumber="51" pageNumber="57" updateTime="1698598193725" updateUser="plazi">
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<mods:title id="A4BE3A3F55440390F16F6D480C05FAFC">First report of the sac spider genus Systaria Simon (Araneae: Clubionidae) from China: Systaria hainanensis sp. nov. and Systaria mengla (Song &amp; Zhu, 1994) comb. nov.</mods:title>
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<mods:namePart id="3BBA20A4111CFD6714C0583382C89F61">Zhang, Feng</mods:namePart>
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<mods:namePart id="DD1EC3066BCDDD7B15BBA2B955D9FB7E">Fu, Jianying</mods:namePart>
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<mods:namePart id="70E475BC97D354C0ED933D8D0220A9B4">Zhu, Mingsheng</mods:namePart>
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<mods:date id="42C359A770B7CE012E126B8FF3035782">2009</mods:date>
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<mods:number id="B172315FDC52832410394669CEC2593C">2305</mods:number>
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<treatment id="B50B87B11E71015598F3FF3C3DF92F74" ID-DOI="http://doi.org/10.5281/zenodo.6222087" ID-GBIF-Taxon="119627421" ID-Zenodo-Dep="6222087" LSID="urn:lsid:plazi:treatment:B50B87B11E71015598F3FF3C3DF92F74" httpUri="http://treatment.plazi.org/id/B50B87B11E71015598F3FF3C3DF92F74" lastPageId="8" lastPageNumber="59" pageId="6" pageNumber="57">
<subSubSection id="75B8652C1E71015B98F3FF3C3F132988" pageId="6" pageNumber="57" type="nomenclature">
<paragraph id="3D1D36A71E71015B98F3FF3C3CDE29ED" blockId="6.[151,739,151,212]" box="[151,739,151,177]" pageId="6" pageNumber="57">
<heading id="665581CB1E71015B98F3FF3C3CDE29ED" bold="true" box="[151,739,151,177]" fontSize="11" level="1" pageId="6" pageNumber="57" reason="1">
<emphasis id="0FD6EAB51E71015B98F3FF3C3CDE29ED" bold="true" box="[151,739,151,177]" pageId="6" pageNumber="57">
<taxonomicName id="FAA24D241E71015B98F3FF3C3C6829ED" ID-CoL="7BBVN" authority="Song &amp; Zhu, 1994" authorityName="Song &amp; Zhu" authorityYear="1994" box="[151,597,151,177]" class="Arachnida" family="Miturgidae" genus="Systaria" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="mengla" status="comb. nov.">
<emphasis id="0FD6EAB51E71015B98F3FF3C3F6429ED" bold="true" box="[151,345,151,177]" italics="true" pageId="6" pageNumber="57">Systaria mengla</emphasis>
(
<bibRefCitation id="59334B561E71015B990CFF3C3C7029ED" author="Song" box="[360,589,151,177]" pageId="6" pageNumber="60" refString="Song, D. X. &amp; Zhu, M. S. (1994) On some species of cave arachnids of China. In Collected Papers on Zoology in Celebration of the Sixtieth Anniversary of the Founding of the Zoological Society of China and in Honor of the Centenary of the Birth of Professor Zhen Chen. China Zoological Society, Beijing, pp. 35 - 46." type="book chapter" year="1994">Song &amp; Zhu, 1994</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="14E557CE1E71015B9A39FF3C3CDE29ED" box="[605,739,151,177]" pageId="6" pageNumber="57">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3D1D36A71E71015B98F3FF113F132988" blockId="6.[151,739,151,212]" box="[151,302,186,212]" pageId="6" pageNumber="57">
(
<figureCitation id="A5992A221E71015B98FBFF113F182988" box="[159,293,186,212]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,269,1897,1921]" captionTargetBox="[189,1412,199,1861]" captionTargetId="figure@7.[177,1421,194,1866]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Systaria mengla (Song &amp; Zhu, 1994) comb. nov., female holotype. 13. Female body, dorsal view. 14. Eye region of female, dorsal view. 15. Endites, labium and sternum of female, ventral view. 16. Left chelicera of female, posterior view. 17. Spinnerets, ventral view. 18. Epigynum, ventral view. 19. Vulva, dorsal view. B—bursa; COcopulatory opening; S—spermatheca. Scale bars: 1 mm (13 – 17); 0.5 mm (18 – 19)." httpUri="https://zenodo.org/record/275385/files/figure.png" pageId="6" pageNumber="57">Figs 13–19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="75B8652C1E71015B98F3FEA33A072843" box="[151,1082,264,287]" pageId="6" pageNumber="57" type="reference_group">
<paragraph id="3D1D36A71E71015B98F3FEA33A072843" blockId="6.[151,1082,264,287]" box="[151,1082,264,287]" pageId="6" pageNumber="57">
<treatmentCitationGroup id="1DB211891E71015B98F3FEA33A072843" box="[151,1082,264,287]" pageId="6" pageNumber="57">
<taxonomicName id="FAA24D241E71015B98F3FEA33C122843" authority="Song &amp; Zhu 1994: 42" authorityName="Song &amp; Zhu" authorityPageNumber="42" authorityYear="1994" box="[151,559,264,287]" class="Arachnida" family="Liocranidae" genus="Itatsina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="mengla">
<emphasis id="0FD6EAB51E71015B98F3FEA33F002843" box="[151,317,264,287]" italics="true" pageId="6" pageNumber="57">Itatsina mengla</emphasis>
<treatmentCitation id="BC0310B61E71015B9920FEA33C122843" author="Song" box="[324,559,264,287]" page="42" pageId="6" pageNumber="57" year="1994">
<bibRefCitation id="59334B561E71015B9920FEA33C3A2843" author="Song" box="[324,519,264,287]" pageId="6" pageNumber="60" refString="Song, D. X. &amp; Zhu, M. S. (1994) On some species of cave arachnids of China. In Collected Papers on Zoology in Celebration of the Sixtieth Anniversary of the Founding of the Zoological Society of China and in Honor of the Centenary of the Birth of Professor Zhen Chen. China Zoological Society, Beijing, pp. 35 - 46." type="book chapter" year="1994">Song &amp; Zhu 1994</bibRefCitation>
: 42
</treatmentCitation>
</taxonomicName>
, figs 7A–B;
<treatmentCitation id="BC0310B61E71015B9AD2FEA33DA02843" author="Song" box="[694,925,264,287]" page="402" pageId="6" pageNumber="57" year="1999">
<bibRefCitation id="59334B561E71015B9AD2FEA33D542843" author="Song" box="[694,873,264,287]" pageId="6" pageNumber="60" refString="Song, D. X., Zhu, M. S. &amp; Chen, J. (1999) The Spiders of China. Hebei Science &amp; Technology Publishing House, Shijiazhuang, 640 pp." type="book" year="1999">
Song
<emphasis id="0FD6EAB51E71015B9A90FEA33D142843" box="[756,809,264,287]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
1999
</bibRefCitation>
: 402
</treatmentCitation>
, figs 238G–H.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="75B8652C1E71015B98F3FEE53CFE28D3" pageId="6" pageNumber="57" type="materials_examined">
<paragraph id="3D1D36A71E71015B98F3FEE53CFE28D3" blockId="6.[151,1437,334,1056]" pageId="6" pageNumber="57">
<emphasis id="0FD6EAB51E71015B98F3FEE53F652834" bold="true" box="[151,344,334,360]" pageId="6" pageNumber="57">
<typeStatus id="E21988051E71015B98F3FEE53EEB2834" box="[151,214,334,360]" pageId="6" pageNumber="57">Type</typeStatus>
material.
</emphasis>
<typeStatus id="E21988051E71015B9907FEE53FEA2834" box="[355,471,334,360]" pageId="6" pageNumber="57" type="holotype">Holotype</typeStatus>
female,
<collectingCountry id="45B576371E71015B9A22FEE53C902834" box="[582,685,334,360]" name="China" pageId="6" pageNumber="57">CHINA</collectingCountry>
: Yunnan Province,
<taxonomicName id="FAA24D241E71015B9BC6FEE53A3C2834" box="[930,1025,334,360]" class="Arachnida" family="Liocranidae" genus="Itatsina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="mengla">Mengla</taxonomicName>
County, Caiyun Cave [
<geoCoordinate id="589650601E71015B9D4CFEE53BA52834" box="[1320,1432,334,360]" direction="north" orientation="latitude" pageId="6" pageNumber="57" precision="555" value="21.48">21.48°N</geoCoordinate>
,
<geoCoordinate id="589650601E71015B98F3FEDE3F3528D3" box="[151,264,373,399]" direction="east" orientation="longitude" pageId="6" pageNumber="57" precision="555" value="101.56">101.56°E</geoCoordinate>
],
<date id="491C10671E71015B9979FEDE3FF928D3" box="[285,452,373,399]" pageId="6" pageNumber="57" value="1992-04-19">19 April 1992</date>
, deposited in
<collectionCode id="5BB3AE621E71015B9A02FEDE3C8228D3" box="[614,703,373,399]" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="6" pageNumber="57">IZCAS</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="75B8652C1E71015B98A1FE303C402E7E" pageId="6" pageNumber="57" type="diagnosis">
<paragraph id="3D1D36A71E71015B98A1FE303DC52B5F" blockId="6.[151,1437,334,1056]" pageId="6" pageNumber="57">
<emphasis id="0FD6EAB51E71015B98A1FE303F7B28E9" bold="true" box="[197,326,411,437]" pageId="6" pageNumber="57">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="FAA24D241E71015B9A0DFE373D0428E9" box="[617,825,412,437]" class="Arachnida" family="Miturgidae" genus="Systaria" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="insulana">
<emphasis id="0FD6EAB51E71015B9A0DFE373D0428E9" box="[617,825,412,437]" italics="true" pageId="6" pageNumber="57">Systaria insulana</emphasis>
</taxonomicName>
, but can easily be distinguished from the latter by: (1) the central epigynal atrium that is distinctly larger than that of the latter; (2) chelicerae with two retromarginal teeth, while three retromarginal teeth are found in the latter.
</paragraph>
<paragraph id="3D1D36A71E71015B98A1FDA43D642D7C" blockId="6.[151,1437,334,1056]" pageId="6" pageNumber="57">
<emphasis id="0FD6EAB51E71015B98A1FDA43F462B75" bold="true" box="[197,379,527,553]" pageId="6" pageNumber="57">Redescription.</emphasis>
Female (
<typeStatus id="E21988051E71015B998DFDBB3C6B2B76" box="[489,598,528,554]" pageId="6" pageNumber="57" type="holotype">holotype</typeStatus>
). Total length 7.87; cephalothorax 3.51 long, 2.34 wide; abdomen 4.36 long, 2.52 wide. Carapace (
<figureCitation id="A5992A221E71015B9989FD9D3C7A2B0C" box="[493,583,566,592]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,269,1897,1921]" captionTargetBox="[189,1412,199,1861]" captionTargetId="figure@7.[177,1421,194,1866]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Systaria mengla (Song &amp; Zhu, 1994) comb. nov., female holotype. 13. Female body, dorsal view. 14. Eye region of female, dorsal view. 15. Endites, labium and sternum of female, ventral view. 16. Left chelicera of female, posterior view. 17. Spinnerets, ventral view. 18. Epigynum, ventral view. 19. Vulva, dorsal view. B—bursa; COcopulatory opening; S—spermatheca. Scale bars: 1 mm (13 – 17); 0.5 mm (18 – 19)." httpUri="https://zenodo.org/record/275385/files/figure.png" pageId="6" pageNumber="57">Fig. 13</figureCitation>
) yellow-brown, darker in eye area; longitudinal ovoid, cephalic area relatively broad; fovea longitudinal and distinct. Eye diameters: AME 0.11, ALE 0.09, PME 0.08, PLE 0.08; eye interdistances: AME–AME 0.15, AME–ALE 0.12, PME–PME 0.18, PME–PLE 0.15, ALE–PLE 0.12; MOA 0.25 long, anterior width 0.23, posterior width 0.26. AER slightly recurved (
<figureCitation id="A5992A221E71015B9CEAFD013AD42B98" box="[1166,1257,682,708]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,269,1897,1921]" captionTargetBox="[189,1412,199,1861]" captionTargetId="figure@7.[177,1421,194,1866]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Systaria mengla (Song &amp; Zhu, 1994) comb. nov., female holotype. 13. Female body, dorsal view. 14. Eye region of female, dorsal view. 15. Endites, labium and sternum of female, ventral view. 16. Left chelicera of female, posterior view. 17. Spinnerets, ventral view. 18. Epigynum, ventral view. 19. Vulva, dorsal view. B—bursa; COcopulatory opening; S—spermatheca. Scale bars: 1 mm (13 – 17); 0.5 mm (18 – 19)." httpUri="https://zenodo.org/record/275385/files/figure.png" pageId="6" pageNumber="57">Fig. 14</figureCitation>
), PER slightly procurved and longer than AER. Clypeal height 0.11, equal to the diameter of AME. Chelicerae (
<figureCitation id="A5992A221E71015B9D5EFD7A3BAE2BB7" box="[1338,1427,721,747]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,269,1897,1921]" captionTargetBox="[189,1412,199,1861]" captionTargetId="figure@7.[177,1421,194,1866]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Systaria mengla (Song &amp; Zhu, 1994) comb. nov., female holotype. 13. Female body, dorsal view. 14. Eye region of female, dorsal view. 15. Endites, labium and sternum of female, ventral view. 16. Left chelicera of female, posterior view. 17. Spinnerets, ventral view. 18. Epigynum, ventral view. 19. Vulva, dorsal view. B—bursa; COcopulatory opening; S—spermatheca. Scale bars: 1 mm (13 – 17); 0.5 mm (18 – 19)." httpUri="https://zenodo.org/record/275385/files/figure.png" pageId="6" pageNumber="57">Fig. 16</figureCitation>
) moderately long, brown, with three promarginal and two retromarginal teeth. Endites (
<figureCitation id="A5992A221E71015B9CC9FD533B392A4E" box="[1197,1284,760,786]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,269,1897,1921]" captionTargetBox="[189,1412,199,1861]" captionTargetId="figure@7.[177,1421,194,1866]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Systaria mengla (Song &amp; Zhu, 1994) comb. nov., female holotype. 13. Female body, dorsal view. 14. Eye region of female, dorsal view. 15. Endites, labium and sternum of female, ventral view. 16. Left chelicera of female, posterior view. 17. Spinnerets, ventral view. 18. Epigynum, ventral view. 19. Vulva, dorsal view. B—bursa; COcopulatory opening; S—spermatheca. Scale bars: 1 mm (13 – 17); 0.5 mm (18 – 19)." httpUri="https://zenodo.org/record/275385/files/figure.png" pageId="6" pageNumber="57">Fig. 15</figureCitation>
) and labium yellowish-brown. Endites widest anteriorly, with a tuft of setae prolaterally. Labium long as wide, tip rounded and with several setae. Sternum (
<figureCitation id="A5992A221E71015B9A45FCEE3C4A2A03" box="[545,631,837,863]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,269,1897,1921]" captionTargetBox="[189,1412,199,1861]" captionTargetId="figure@7.[177,1421,194,1866]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Systaria mengla (Song &amp; Zhu, 1994) comb. nov., female holotype. 13. Female body, dorsal view. 14. Eye region of female, dorsal view. 15. Endites, labium and sternum of female, ventral view. 16. Left chelicera of female, posterior view. 17. Spinnerets, ventral view. 18. Epigynum, ventral view. 19. Vulva, dorsal view. B—bursa; COcopulatory opening; S—spermatheca. Scale bars: 1 mm (13 – 17); 0.5 mm (18 – 19)." httpUri="https://zenodo.org/record/275385/files/figure.png" pageId="6" pageNumber="57">Fig. 15</figureCitation>
) light brown, shield-shaped, not rebordered, with single sclerotised extensions to each coxa; anterior margin truncate and posterior margin not extending between coxae IV, only slightly narrower in front than in the middle. Legs light brown, leg spination: femora: I d1-1-1, r0-0-2; II d1- 1-1, r1-1-1, v0-0-1; III d1-1-0, p0-1-1, r1-1-1; IV d1-1-1, p0-0-1, r1-1-1; tibiae: I
<date id="491C10671E71015B9C2AFC123A532A8F" box="[1102,1134,953,979]" pageId="6" pageNumber="57">v2</date>
-0-0; II
<date id="491C10671E71015B9CA1FC123ADF2A8F" box="[1221,1250,953,979]" pageId="6" pageNumber="57">v1</date>
-0-0; III d1-0-0, p0-1-1, r0-1-1,
<date id="491C10671E71015B992FFC4B3FB92AA6" box="[331,388,992,1018]" pageId="6" pageNumber="57">v2-2</date>
-2; IV d1-0-0, p0-1-1, r0-1-1,
<date id="491C10671E71015B9ABBFC4B3D252AA6" box="[735,792,992,1018]" pageId="6" pageNumber="57">v2-2</date>
-0; metatarsi: I
<date id="491C10671E71015B9BA3FC4B3DD52AA6" box="[967,1000,992,1018]" pageId="6" pageNumber="57">v2</date>
-0-0; II
<date id="491C10671E71015B9C5BFC4B3A622AA6" box="[1087,1119,992,1018]" pageId="6" pageNumber="57">v2</date>
-0-0; III p1-2-2, r1-2-2,
<date id="491C10671E71015B9D11FC4B3E9A2D7C" pageId="6" pageNumber="57">v2- 2</date>
-2; IV p1-2-2, r1-2-2,
<date id="491C10671E71015B99CCFBAD3FDC2D7C" box="[424,481,1030,1056]" pageId="6" pageNumber="57">v2-2</date>
-2. Leg formula: 4123 (Table 3).
</paragraph>
<paragraph id="3D1D36A71E71015B98F3FBF83B732C67" pageId="6" pageNumber="57">
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<emphasis id="0FD6EAB51E71015B98F3FBF83ED22D37" bold="true" box="[151,239,1107,1131]" pageId="6" pageNumber="57">TABLE</emphasis>
</th>
<th id="C0435D991E71FEA29892FBF83F352D37" box="[246,264,1107,1131]" gridcol="1" gridrow="0" pageId="6" pageNumber="57">
<emphasis id="0FD6EAB51E71015B9892FBF83F352D37" bold="true" box="[246,264,1107,1131]" pageId="6" pageNumber="57">3.</emphasis>
</th>
<th id="C0435D991E71FEA29974FBF83FE52D37" box="[272,472,1107,1131]" gridcol="2" gridrow="0" pageId="6" pageNumber="57">Leg measurements</th>
<th id="C0435D991E71FEA299BBFBF83C6E2D37" box="[479,595,1107,1131]" gridcol="3" gridrow="0" pageId="6" pageNumber="57">
of
<taxonomicName id="FAA24D241E71015B9998FBFF3C6E2D37" box="[508,595,1108,1131]" class="Arachnida" family="Miturgidae" genus="Systaria" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="0FD6EAB51E71015B9998FBFF3C6E2D37" box="[508,595,1108,1131]" italics="true" pageId="6" pageNumber="57">Systaria</emphasis>
</taxonomicName>
</th>
<th id="C0435D991E71FEA29A3FFBF83CD22D37" box="[603,751,1107,1131]" gridcol="4" gridrow="0" pageId="6" pageNumber="57">
<taxonomicName id="FAA24D241E71015B9A3FFBFF3CD22D37" authority="Song" authorityName="Song" box="[603,751,1108,1131]" class="Arachnida" family="Liocranidae" genus="Itatsina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="mengla">
<emphasis id="0FD6EAB51E71015B9A3FFBFF3C972D37" box="[603,682,1108,1131]" italics="true" pageId="6" pageNumber="57">mengla</emphasis>
(Song
</taxonomicName>
</th>
<th id="C0435D991E71FEA29A92FBF83D7E2D37" box="[758,835,1107,1131]" gridcol="5" gridrow="0" pageId="6" pageNumber="57">&amp; Zhu,</th>
<th id="C0435D991E71FEA29B2EFBF83AB72D37" box="[842,1162,1107,1131]" gridcol="6" gridrow="0" pageId="6" pageNumber="57">
1994)
<emphasis id="0FD6EAB51E71015B9BEAFBF83A342D37" bold="true" box="[910,1033,1107,1131]" pageId="6" pageNumber="57">comb. nov.</emphasis>
, female.
</th>
</tr>
<tr id="839234E51E71FEA298F3FB2D3B6D2DC7" box="[151,1360,1158,1179]" gridrow="1" pageId="6" pageNumber="57" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="C0435D991E71FEA29974FB2D3FE52DC7" box="[272,472,1158,1179]" gridcol="2" gridrow="1" pageId="6" pageNumber="57">Femur</td>
<td id="C0435D991E71FEA299BBFB2D3C6E2DC7" box="[479,595,1158,1179]" gridcol="3" gridrow="1" pageId="6" pageNumber="57">Patella</td>
<td id="C0435D991E71FEA29A3FFB2D3CD22DC7" box="[603,751,1158,1179]" gridcol="4" gridrow="1" pageId="6" pageNumber="57">Tibia</td>
<td id="C0435D991E71FEA29B2EFB2D3AB72DC7" box="[842,1162,1158,1179]" gridcol="6" gridrow="1" pageId="6" pageNumber="57">Metatarsus Tarsus</td>
<td id="C0435D991E71FEA29D7EFB2D3B6D2DC7" box="[1306,1360,1158,1179]" gridcol="7" gridrow="1" pageId="6" pageNumber="57">Total</td>
</tr>
<tr id="839234E51E71FEA298F3FB053B6D2D9F" box="[151,1360,1198,1219]" gridrow="2" pageId="6" pageNumber="57" rowspan-1="1" rowspan-5="1">
<th id="C0435D991E71FEA298F3FB053ED22D9F" box="[151,239,1198,1219]" gridcol="0" gridrow="2" pageId="6" pageNumber="57">I</th>
<td id="C0435D991E71FEA29974FB053FE52D9F" box="[272,472,1198,1219]" gridcol="2" gridrow="2" pageId="6" pageNumber="57">2.84</td>
<td id="C0435D991E71FEA299BBFB053C6E2D9F" box="[479,595,1198,1219]" gridcol="3" gridrow="2" pageId="6" pageNumber="57">1.35</td>
<td id="C0435D991E71FEA29A3FFB053CD22D9F" box="[603,751,1198,1219]" gridcol="4" gridrow="2" pageId="6" pageNumber="57">2.70</td>
<td id="C0435D991E71FEA29B2EFB053AB72D9F" box="[842,1162,1198,1219]" gridcol="6" gridrow="2" pageId="6" pageNumber="57">2.02 1.44</td>
<td id="C0435D991E71FEA29D7EFB053B6D2D9F" box="[1306,1360,1198,1219]" gridcol="7" gridrow="2" pageId="6" pageNumber="57">10.35</td>
</tr>
<tr id="839234E51E71FEA298F3FB7D3B6D2DB7" box="[151,1360,1238,1259]" gridrow="3" pageId="6" pageNumber="57" rowspan-1="1" rowspan-5="1">
<th id="C0435D991E71FEA298F3FB7D3ED22DB7" box="[151,239,1238,1259]" gridcol="0" gridrow="3" pageId="6" pageNumber="57">II</th>
<td id="C0435D991E71FEA29974FB7D3FE52DB7" box="[272,472,1238,1259]" gridcol="2" gridrow="3" pageId="6" pageNumber="57">2.61</td>
<td id="C0435D991E71FEA299BBFB7D3C6E2DB7" box="[479,595,1238,1259]" gridcol="3" gridrow="3" pageId="6" pageNumber="57">0.90</td>
<td id="C0435D991E71FEA29A3FFB7D3CD22DB7" box="[603,751,1238,1259]" gridcol="4" gridrow="3" pageId="6" pageNumber="57">2.25</td>
<td id="C0435D991E71FEA29B2EFB7D3AB72DB7" box="[842,1162,1238,1259]" gridcol="6" gridrow="3" pageId="6" pageNumber="57">2.11 1.30</td>
<td id="C0435D991E71FEA29D7EFB7D3B6D2DB7" box="[1306,1360,1238,1259]" gridcol="7" gridrow="3" pageId="6" pageNumber="57">9.17</td>
</tr>
<tr id="839234E51E71FEA298F3FB553B6D2C4F" box="[151,1360,1278,1299]" gridrow="4" pageId="6" pageNumber="57" rowspan-1="1" rowspan-5="1">
<th id="C0435D991E71FEA298F3FB553ED22C4F" box="[151,239,1278,1299]" gridcol="0" gridrow="4" pageId="6" pageNumber="57">III</th>
<td id="C0435D991E71FEA29974FB553FE52C4F" box="[272,472,1278,1299]" gridcol="2" gridrow="4" pageId="6" pageNumber="57">1.89</td>
<td id="C0435D991E71FEA299BBFB553C6E2C4F" box="[479,595,1278,1299]" gridcol="3" gridrow="4" pageId="6" pageNumber="57">1.08</td>
<td id="C0435D991E71FEA29A3FFB553CD22C4F" box="[603,751,1278,1299]" gridcol="4" gridrow="4" pageId="6" pageNumber="57">1.80</td>
<td id="C0435D991E71FEA29B2EFB553AB72C4F" box="[842,1162,1278,1299]" gridcol="6" gridrow="4" pageId="6" pageNumber="57">2.29 1.39</td>
<td id="C0435D991E71FEA29D7EFB553B6D2C4F" box="[1306,1360,1278,1299]" gridcol="7" gridrow="4" pageId="6" pageNumber="57">8.45</td>
</tr>
<tr id="839234E51E71FEA298F3FA8D3B6D2C67" box="[151,1360,1318,1339]" gridrow="5" pageId="6" pageNumber="57" rowspan-1="1" rowspan-5="1">
<th id="C0435D991E71FEA298F3FA8D3ED22C67" box="[151,239,1318,1339]" gridcol="0" gridrow="5" pageId="6" pageNumber="57">IV</th>
<td id="C0435D991E71FEA29974FA8D3FE52C67" box="[272,472,1318,1339]" gridcol="2" gridrow="5" pageId="6" pageNumber="57">3.19</td>
<td id="C0435D991E71FEA299BBFA8D3C6E2C67" box="[479,595,1318,1339]" gridcol="3" gridrow="5" pageId="6" pageNumber="57">1.35</td>
<td id="C0435D991E71FEA29A3FFA8D3CD22C67" box="[603,751,1318,1339]" gridcol="4" gridrow="5" pageId="6" pageNumber="57">2.65</td>
<td id="C0435D991E71FEA29B2EFA8D3AB72C67" box="[842,1162,1318,1339]" gridcol="6" gridrow="5" pageId="6" pageNumber="57">2.75 1.40</td>
<td id="C0435D991E71FEA29D7EFA8D3B6D2C67" box="[1306,1360,1318,1339]" gridcol="7" gridrow="5" pageId="6" pageNumber="57">11.34</td>
</tr>
</table>
</paragraph>
<paragraph id="3D1D36A71E71015B98A1FA2E3AC42F66" blockId="6.[151,1437,1413,1980]" pageId="6" pageNumber="57">
Abdomen (
<figureCitation id="A5992A221E71015B9929FA2E3F9B2CC3" box="[333,422,1413,1439]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,269,1897,1921]" captionTargetBox="[189,1412,199,1861]" captionTargetId="figure@7.[177,1421,194,1866]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Systaria mengla (Song &amp; Zhu, 1994) comb. nov., female holotype. 13. Female body, dorsal view. 14. Eye region of female, dorsal view. 15. Endites, labium and sternum of female, ventral view. 16. Left chelicera of female, posterior view. 17. Spinnerets, ventral view. 18. Epigynum, ventral view. 19. Vulva, dorsal view. B—bursa; COcopulatory opening; S—spermatheca. Scale bars: 1 mm (13 – 17); 0.5 mm (18 – 19)." httpUri="https://zenodo.org/record/275385/files/figure.png" pageId="6" pageNumber="57">Fig. 13</figureCitation>
) ovoid, yellow-brown, slightly darker centrally; covered with long and dark setae, without muscular impressions centrally. Venter of abdomen light brown. ALS (
<figureCitation id="A5992A221E71015B9C2FFA073A9F2C9A" box="[1099,1186,1452,1478]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,269,1897,1921]" captionTargetBox="[189,1412,199,1861]" captionTargetId="figure@7.[177,1421,194,1866]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Systaria mengla (Song &amp; Zhu, 1994) comb. nov., female holotype. 13. Female body, dorsal view. 14. Eye region of female, dorsal view. 15. Endites, labium and sternum of female, ventral view. 16. Left chelicera of female, posterior view. 17. Spinnerets, ventral view. 18. Epigynum, ventral view. 19. Vulva, dorsal view. B—bursa; COcopulatory opening; S—spermatheca. Scale bars: 1 mm (13 – 17); 0.5 mm (18 – 19)." httpUri="https://zenodo.org/record/275385/files/figure.png" pageId="6" pageNumber="57">Fig. 17</figureCitation>
) cylindrical, slightly tapering, separated almost by more than half of their diameter. PMS thin and conical, as long as anterior spinnerets. PLS with distal segment shorter than basal segment, tapering; and basal segment of PLS with similar length as basal segment of ALS. Tracheal spiracle located just anterior to the spinnerets.
</paragraph>
<paragraph id="3D1D36A71E71015B98A1F9ED3C402E7E" blockId="6.[151,1437,1413,1980]" pageId="6" pageNumber="57">
Epigynum (
<figureCitation id="A5992A221E71015B9936F9ED3FE72F3C" box="[338,474,1606,1632]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,269,1897,1921]" captionTargetBox="[189,1412,199,1861]" captionTargetId="figure@7.[177,1421,194,1866]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Systaria mengla (Song &amp; Zhu, 1994) comb. nov., female holotype. 13. Female body, dorsal view. 14. Eye region of female, dorsal view. 15. Endites, labium and sternum of female, ventral view. 16. Left chelicera of female, posterior view. 17. Spinnerets, ventral view. 18. Epigynum, ventral view. 19. Vulva, dorsal view. B—bursa; COcopulatory opening; S—spermatheca. Scale bars: 1 mm (13 – 17); 0.5 mm (18 – 19)." httpUri="https://zenodo.org/record/275385/files/figure.png" pageId="6" pageNumber="57">Figs 18–19</figureCitation>
) with weakly sclerotised epigynal plate, epigynal atrium large and central, with arched anterior margin, two copulatory openings concave, located postero-medially near the posterior margin of atrium. Bursae situated anterior of copulatory openings, large sac-shaped and semi-transparent. Spermathecae globular, distinctly smaller than bursae, situated posteriorly and near the genital furrow; connecting with the copulatory opening through a wide duct; fertilization ducts short, arising from the postero-central ends of the spermathecae.
</paragraph>
</subSubSection>
<subSubSection id="75B8652C1E71015B98A1F8853CC22E14" box="[197,767,1838,1864]" pageId="6" pageNumber="57" type="distribution">
<paragraph id="3D1D36A71E71015B98A1F8853CC22E14" blockId="6.[151,1437,1413,1980]" box="[197,767,1838,1864]" pageId="6" pageNumber="57">
<emphasis id="0FD6EAB51E71015B98A1F8853F582E14" bold="true" box="[197,357,1838,1864]" pageId="6" pageNumber="57">Distribution.</emphasis>
Yunnan Province,
<collectingCountry id="45B576371E71015B9A20F8853CB72E14" box="[580,650,1838,1864]" name="China" pageId="6" pageNumber="57">China</collectingCountry>
(
<figureCitation id="A5992A221E71015B9AFEF8853CCF2E14" box="[666,754,1838,1864]" captionStart="FIGURE 20" captionStartId="8.[151,255,1132,1156]" captionTargetBox="[151,1436,189,1108]" captionTargetId="figure@8.[151,1436,176,1108]" captionTargetPageId="8" captionText="FIGURE 20. Distribution records of Systaria in China." httpUri="https://zenodo.org/record/275386/files/figure.png" pageId="6" pageNumber="57">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="75B8652C1E71015598A1F8FE3DF92F74" lastPageId="8" lastPageNumber="59" pageId="6" pageNumber="57" type="discussion">
<paragraph id="3D1D36A71E71015B98A1F8FE3D992EE0" blockId="6.[151,1437,1413,1980]" pageId="6" pageNumber="57">
<emphasis id="0FD6EAB51E71015B98A1F8FE3F7E2E33" bold="true" box="[197,323,1877,1903]" pageId="6" pageNumber="57">Remarks.</emphasis>
<taxonomicName id="FAA24D241E71015B9928F8FE3C322E32" box="[332,527,1877,1902]" class="Arachnida" family="Liocranidae" genus="Itatsina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="mengla">
<emphasis id="0FD6EAB51E71015B9928F8FE3C322E32" box="[332,527,1877,1902]" italics="true" pageId="6" pageNumber="57">Itatsina mengla</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="59334B561E71015B9A93F8FE3DD62E33" author="Song" box="[759,1003,1877,1903]" pageId="6" pageNumber="60" refString="Song, D. X. &amp; Zhu, M. S. (1994) On some species of cave arachnids of China. In Collected Papers on Zoology in Celebration of the Sixtieth Anniversary of the Founding of the Zoological Society of China and in Honor of the Centenary of the Birth of Professor Zhen Chen. China Zoological Society, Beijing, pp. 35 - 46." type="book chapter" year="1994">Song &amp; Zhu (1994)</bibRefCitation>
based on a single female
<typeStatus id="E21988051E71015B9D54F8FE3BA12E33" box="[1328,1436,1877,1903]" pageId="6" pageNumber="57" type="holotype">holotype</typeStatus>
collected from Cave Caiyun,
<taxonomicName id="FAA24D241E71015B9996F8D73C712ECA" box="[498,588,1916,1942]" class="Arachnida" family="Liocranidae" genus="Itatsina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="mengla">Mengla</taxonomicName>
County, Yunnan Province,
<collectingCountry id="45B576371E71015B9BF5F8D73DE62ECA" box="[913,987,1916,1942]" name="China" pageId="6" pageNumber="57">China</collectingCountry>
. No male has so far been found.
<bibRefCitation id="59334B561E71015B9D04F8D73F1B2EE0" pageId="6" pageNumber="60" refString="Song, D. X., Zhu, M. S. &amp; Chen, J. (1999) The Spiders of China. Hebei Science &amp; Technology Publishing House, Shijiazhuang, 640 pp." type="book">
Song
<emphasis id="0FD6EAB51E71015B98F3F8083EEC2EE0" box="[151,209,1955,1980]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
(1999)
</bibRefCitation>
reproduced the illustrations from
<bibRefCitation id="59334B561E71015B9AD2F8093D9D2EE0" author="Song" box="[694,928,1954,1980]" pageId="6" pageNumber="60" refString="Song, D. X. &amp; Zhu, M. S. (1994) On some species of cave arachnids of China. In Collected Papers on Zoology in Celebration of the Sixtieth Anniversary of the Founding of the Zoological Society of China and in Honor of the Centenary of the Birth of Professor Zhen Chen. China Zoological Society, Beijing, pp. 35 - 46." type="book chapter" year="1994">Song &amp; Zhu (1994)</bibRefCitation>
.
</paragraph>
<caption id="69DD662F1E70015A98F3F8C23A382EB5" httpUri="https://zenodo.org/record/275385/files/figure.png" pageId="7" pageNumber="58" targetBox="[189,1412,199,1861]" targetPageId="7">
<paragraph id="3D1D36A71E70015A98F3F8C23A382EB5" blockId="7.[151,1436,1897,2025]" pageId="7" pageNumber="58">
<emphasis id="0FD6EAB51E70015A98F3F8C23F602EDD" bold="true" box="[151,349,1897,1921]" pageId="7" pageNumber="58">FIGURES 13–19.</emphasis>
<taxonomicName id="FAA24D241E70015A9900F8C13CD22EDD" authority="Song &amp; Zhu, 1994" authorityName="Song &amp; Zhu" authorityYear="1994" box="[356,751,1898,1921]" class="Arachnida" family="Miturgidae" genus="Systaria" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="mengla" status="comb. nov.">
<emphasis id="0FD6EAB51E70015A9900F8C13C2C2EDD" box="[356,529,1898,1921]" italics="true" pageId="7" pageNumber="58">Systaria mengla</emphasis>
(Song &amp; Zhu, 1994)
</taxonomicName>
<taxonomicNameLabel id="14E557CE1E70015A9A91F8C23D522EDD" box="[757,879,1897,1921]" pageId="7" pageNumber="58">
<emphasis id="0FD6EAB51E70015A9A91F8C23D522EDD" bold="true" box="[757,879,1897,1921]" pageId="7" pageNumber="58">comb. nov.</emphasis>
</taxonomicNameLabel>
, female holotype. 13. Female body, dorsal view. 14. Eye region of female, dorsal view. 15. Endites, labium and sternum of female, ventral view. 16. Left chelicera of female, posterior view. 17. Spinnerets, ventral view. 18. Epigynum, ventral view. 19. Vulva, dorsal view. B—bursa; COcopulatory opening; S—spermatheca. Scale bars: 1 mm (13–17); 0.5 mm (18–19).
</paragraph>
</caption>
<caption id="69DD662F1E7F015598F3FBC73CDA2DD8" box="[151,743,1132,1156]" httpUri="https://zenodo.org/record/275386/files/figure.png" pageId="8" pageNumber="59" targetBox="[151,1436,189,1108]" targetPageId="8">
<paragraph id="3D1D36A71E7F015598F3FBC73CDA2DD8" blockId="8.[151,743,1132,1156]" box="[151,743,1132,1156]" pageId="8" pageNumber="59">
<emphasis id="0FD6EAB51E7F015598F3FBC73F1A2DD8" bold="true" box="[151,295,1132,1156]" pageId="8" pageNumber="59">FIGURE 20.</emphasis>
Distribution records of
<taxonomicName id="FAA24D241E7F01559A42FBC63C432DD8" box="[550,638,1133,1156]" class="Arachnida" family="Miturgidae" genus="Systaria" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="0FD6EAB51E7F01559A42FBC63C432DD8" box="[550,638,1133,1156]" italics="true" pageId="8" pageNumber="59">Systaria</emphasis>
</taxonomicName>
in China.
</paragraph>
</caption>
<paragraph id="3D1D36A71E7F015598A1FB193DF92F74" blockId="8.[151,1437,1202,1576]" pageId="8" pageNumber="59">
When we borrowed and examined the female
<typeStatus id="E21988051E7F01559B6CFB193D482D90" box="[776,885,1202,1228]" pageId="8" pageNumber="59" type="holotype">holotype</typeStatus>
(No. IZCAS-Ar 9221) from the Institute of Zoology, Chinese Academy of Sciences (IZCAS), Beijing, we found the general appearance of this specimen to conform to the diagnosis of the Systariinae. The
<typeStatus id="E21988051E7F01559B6AFAAB3D452C46" box="[782,888,1280,1306]" pageId="8" pageNumber="59" type="holotype">holotype</typeStatus>
has enlarged AME, slightly procurved PER, widely separated and slightly tapering ALS, an elongated distal segment of PLS, and the genital organs were very similar to those of
<emphasis id="0FD6EAB51E7F015599C9FAE63C852C3A" box="[429,696,1357,1382]" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="FAA24D241E7F015599C9FAE63C892C3A" box="[429,692,1357,1382]" class="Arachnida" family="Miturgidae" genus="Systaria" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="bohorokensis">Systaria bohorokensis</taxonomicName>
.
</emphasis>
Furthermore,
<taxonomicName id="FAA24D241E7F01559B3BFAE63A2B2C3A" box="[863,1046,1357,1382]" class="Arachnida" family="Liocranidae" genus="Itatsina" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="mengla">
<emphasis id="0FD6EAB51E7F01559B3BFAE63A2B2C3A" box="[863,1046,1357,1382]" italics="true" pageId="8" pageNumber="59">Itatsina mengla</emphasis>
</taxonomicName>
was collected from Cave Caiyun (Yunnan Province). The cave habitat of this species is consistent with the preferred habitat of many species of
<taxonomicName id="FAA24D241E7F015598F3FA313EC52CEF" box="[151,248,1434,1459]" class="Arachnida" family="Miturgidae" genus="Systaria" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="0FD6EAB51E7F015598F3FA313EC52CEF" box="[151,248,1434,1459]" italics="true" pageId="8" pageNumber="59">Systaria</emphasis>
</taxonomicName>
as described by
<bibRefCitation id="59334B561E7F015599DAFA313D622CE8" author="Deeleman-Reinhold" box="[446,863,1434,1460]" pageId="8" pageNumber="60" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, 591 pp." type="book" year="2001" yearSuffix="p">Deeleman-Reinhold (2001, p. 204)</bibRefCitation>
, “Most specimens were found in primary forest, leaf litter or under bark, often in caves, those found in caves fail to show morphological adaptation to lightless conditions”. We therefore transfer the species
<taxonomicName id="FAA24D241E7F01559AD3FA433A6F2F5E" authority="Song &amp; Zhu, 1994" authorityName="Song &amp; Zhu" authorityYear="1994" box="[695,1106,1512,1538]" class="Arachnida" family="Liocranidae" genus="Itatsina" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="mengla">
<emphasis id="0FD6EAB51E7F01559AD3FA433D532F5D" box="[695,878,1512,1537]" italics="true" pageId="8" pageNumber="59">Itatsina mengla</emphasis>
<bibRefCitation id="59334B561E7F01559B12FA433A6F2F5E" author="Song" box="[886,1106,1512,1538]" pageId="8" pageNumber="60" refString="Song, D. X. &amp; Zhu, M. S. (1994) On some species of cave arachnids of China. In Collected Papers on Zoology in Celebration of the Sixtieth Anniversary of the Founding of the Zoological Society of China and in Honor of the Centenary of the Birth of Professor Zhen Chen. China Zoological Society, Beijing, pp. 35 - 46." type="book chapter" year="1994">Song &amp; Zhu, 1994</bibRefCitation>
</taxonomicName>
from
<taxonomicName id="FAA24D241E7F01559CFFFA433BA12F5E" box="[1179,1436,1512,1538]" class="Arachnida" family="Liocranidae" genus="Itatsina" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="0FD6EAB51E7F01559CFFFA433AC82F5D" box="[1179,1269,1512,1537]" italics="true" pageId="8" pageNumber="59">Itatsina</emphasis>
(Liocranidae)
</taxonomicName>
to
<taxonomicName id="FAA24D241E7F015598D1F9A53FFF2F74" box="[181,450,1550,1576]" class="Arachnida" family="Miturgidae" genus="Systaria" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="0FD6EAB51E7F015598D1F9A53F282F7B" box="[181,277,1550,1575]" italics="true" pageId="8" pageNumber="59">Systaria</emphasis>
(Clubionidae)
</taxonomicName>
,
<taxonomicName id="FAA24D241E7F015599AAF9A53D0B2F74" authority="Song &amp; Zhu, 1994" authorityName="Song &amp; Zhu" authorityYear="1994" box="[462,822,1550,1576]" class="Arachnida" family="Liocranidae" genus="Itatsina" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="mengla" status="comb. nov.">
<emphasis id="0FD6EAB51E7F015599AAF9A53C7C2F7B" box="[462,577,1550,1575]" italics="true" pageId="8" pageNumber="59">S. mengla</emphasis>
(
<bibRefCitation id="59334B561E7F01559A35F9A53D132F74" author="Song" box="[593,814,1550,1576]" pageId="8" pageNumber="60" refString="Song, D. X. &amp; Zhu, M. S. (1994) On some species of cave arachnids of China. In Collected Papers on Zoology in Celebration of the Sixtieth Anniversary of the Founding of the Zoological Society of China and in Honor of the Centenary of the Birth of Professor Zhen Chen. China Zoological Society, Beijing, pp. 35 - 46." type="book chapter" year="1994">Song &amp; Zhu, 1994</bibRefCitation>
)
</taxonomicName>
<emphasis id="0FD6EAB51E7F01559B59F9A53DF92F74" bold="true" box="[829,964,1550,1576]" pageId="8" pageNumber="59">
<taxonomicNameLabel id="14E557CE1E7F01559B59F9A53DF92F74" box="[829,964,1550,1576]" pageId="8" pageNumber="59">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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