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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1109.83807" ID-GBIF-Dataset="178820d5-d720-47fd-a85e-1ac28fca6c90" ID-Pensoft-Pub="1313-2970-1109-67" ID-Pensoft-UUID="534F47F344D858A49331F110D3D21111" ID-ZooBank="9C8BF86D194A46EF9D49072D09BF9E48" ModsDocID="1313-2970-1109-67" checkinTime="1656747431702" checkinUser="pensoft" docAuthor="Suzuki, Yuya, Hiramatsu, Takehisa &amp; Tatsuta, Haruki" docDate="2022" docId="8B939A91DEA757C6A4947D81CBB7F986" docLanguage="en" docName="ZooKeys 1109: 67-101" docOrigin="ZooKeys 1109" docPubDate="2022-07-01" docSource="http://dx.doi.org/10.3897/zookeys.1109.83807" docTitle="Theridiosoma O. Pickard-Cambridge 1879" docType="treatment" docVersion="2" id="534F47F344D858A49331F110D3D21111" lastPageNumber="67" masterDocId="534F47F344D858A49331F110D3D21111" masterDocTitle="Two new species and a new genus of ray spiders (Araneae, Theridiosomatidae) from the Ryukyu Islands, southwest Japan, with notes on their natural history" masterLastPageNumber="101" masterPageNumber="67" pageNumber="67" updateTime="1656748190795" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species and a new genus of ray spiders (Araneae, Theridiosomatidae) from the Ryukyu Islands, southwest Japan, with notes on their natural history</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Suzuki, Yuya</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6523-9272</mods:nameIdentifier>
<mods:affiliation>The United Graduate School of Agricultural Sciences, Kagoshima University, 1 - 21 - 24, Korimoto, Kagoshima-shi, Kagoshima, 890 - 0065, Japan &amp; Graduate School of Systems Life Sciences, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395 Japan</mods:affiliation>
<mods:nameIdentifier type="email">sasaganiya1206@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hiramatsu, Takehisa</mods:namePart>
<mods:affiliation>Fregrance-Uwado 203, Uwado, Kawagoe-shi, Saitama, 350 - 0816, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tatsuta, Haruki</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8847-8874</mods:nameIdentifier>
<mods:affiliation>Graduate School of Systems Life Sciences, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395 Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-07-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1109</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>67</mods:start>
<mods:end>101</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1109.83807</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1109.83807</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1109-67</mods:identifier>
<mods:identifier type="ZooBank">9C8BF86D194A46EF9D49072D09BF9E48</mods:identifier>
<mods:identifier type="Pensoft-UUID">534F47F344D858A49331F110D3D21111</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:8B939A91DEA757C6A4947D81CBB7F986" httpUri="http://treatment.plazi.org/id/8B939A91DEA757C6A4947D81CBB7F986" lastPageNumber="67" pageId="0" pageNumber="67">
<subSubSection pageId="0" pageNumber="67" type="nomenclature">
<paragraph pageId="0" pageNumber="67">
Genus
<taxonomicName LSID="8B939A91-DEA7-57C6-A494-7D81CBB7F986" authority="O. Pickard-Cambridge, 1879" authorityName="O. Pickard-Cambridge" authorityYear="1879" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theridiosoma" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">Theridiosoma O. Pickard-Cambridge, 1879</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="type species">
<paragraph pageId="0" pageNumber="67">Type species.</paragraph>
<paragraph pageId="0" pageNumber="67">
<taxonomicName baseAuthorityName="L. Koch" baseAuthorityYear="1877" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theridiosoma gemmosum" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="gemmosum">
<emphasis italics="true" pageId="0" pageNumber="67">Theridiosoma gemmosum</emphasis>
</taxonomicName>
(L. Koch, 1877), from Nuremberg, West Germany (not examined).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="remarks">
<paragraph pageId="0" pageNumber="67">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="67">
Males of
<taxonomicName authorityName="O. Pickard-Cambridge" authorityYear="1879" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theridiosoma" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="67">Theridiosoma</emphasis>
</taxonomicName>
species can be distinguished from other theridiosomatid genera by the morphology of the embolic division of the male palp: short and tubular embolus with embolic apophyses fragmented into several long bristle-like parts (
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.422" author="Coddington, JA" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="67" pagination="1 - 96" refId="B4" refString="Coddington, JA, 1986a. The genera of the spider family Theridiosomatidae. Smithsonian Contributions to Zoology 422 (422): 1 - 96, DOI: https://doi.org/10.5479/si.00810282.422" title="The genera of the spider family Theridiosomatidae." url="https://doi.org/10.5479/si.00810282.422" volume="422" year="1986 a">Coddington 1986a</bibRefCitation>
: figs 131, 133). Embolic apophysis varies in number and shape among species and is regarded as an important taxonomic character (e.g.,
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.255.3272" author="Zhao, QY" journalOrPublisher="ZooKeys" pageId="0" pageNumber="67" pagination="1 - 48" refId="B35" refString="Zhao, QY, Li, SQ, 2012. Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae). ZooKeys 255: 1 - 48, DOI: https://doi.org/10.3897/zookeys.255.3272" title="Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae)." url="https://doi.org/10.3897/zookeys.255.3272" volume="255" year="2012">Zhao and Li 2012</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.69.133" author="Suzuki, Y" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="67" pagination="133 - 150" refId="B28" refString="Suzuki, Y, Serita, R, Hiramatsu, T, 2020. Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species. Acta Arachnologica 69 (2): 133 - 150, DOI: https://doi.org/10.2476/asjaa.69.133" title="Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species." url="https://doi.org/10.2476/asjaa.69.133" volume="69" year="2020">Suzuki et al. 2020</bibRefCitation>
). Median apophysis is less sclerotized, curved and attenuates distally (
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.422" author="Coddington, JA" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="67" pagination="1 - 96" refId="B4" refString="Coddington, JA, 1986a. The genera of the spider family Theridiosomatidae. Smithsonian Contributions to Zoology 422 (422): 1 - 96, DOI: https://doi.org/10.5479/si.00810282.422" title="The genera of the spider family Theridiosomatidae." url="https://doi.org/10.5479/si.00810282.422" volume="422" year="1986 a">Coddington 1986a</bibRefCitation>
: figs 132, 133), which is less useful for distinguishing species. A sclerotized projection ('conductor
<normalizedToken originalValue="projection’">projection'</normalizedToken>
) is present on prolateral side of conductor in some species, while absent in others (
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.422" author="Coddington, JA" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="67" pagination="1 - 96" refId="B4" refString="Coddington, JA, 1986a. The genera of the spider family Theridiosomatidae. Smithsonian Contributions to Zoology 422 (422): 1 - 96, DOI: https://doi.org/10.5479/si.00810282.422" title="The genera of the spider family Theridiosomatidae." url="https://doi.org/10.5479/si.00810282.422" volume="422" year="1986 a">Coddington 1986a</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.69.133" author="Suzuki, Y" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="67" pagination="133 - 150" refId="B28" refString="Suzuki, Y, Serita, R, Hiramatsu, T, 2020. Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species. Acta Arachnologica 69 (2): 133 - 150, DOI: https://doi.org/10.2476/asjaa.69.133" title="Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species." url="https://doi.org/10.2476/asjaa.69.133" volume="69" year="2020">Suzuki et al. 2020</bibRefCitation>
). Distal margin of conductor beneath embolic apophyses ('posterior margin of embolic
<normalizedToken originalValue="division’">division'</normalizedToken>
in
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.69.133" author="Suzuki, Y" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="67" pagination="133 - 150" refId="B28" refString="Suzuki, Y, Serita, R, Hiramatsu, T, 2020. Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species. Acta Arachnologica 69 (2): 133 - 150, DOI: https://doi.org/10.2476/asjaa.69.133" title="Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species." url="https://doi.org/10.2476/asjaa.69.133" volume="69" year="2020">Suzuki et al. 2020</bibRefCitation>
) is generally sclerotized and the shape is useful as a taxonomic character (e.g.,
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.69.133" author="Suzuki, Y" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="67" pagination="133 - 150" refId="B28" refString="Suzuki, Y, Serita, R, Hiramatsu, T, 2020. Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species. Acta Arachnologica 69 (2): 133 - 150, DOI: https://doi.org/10.2476/asjaa.69.133" title="Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species." url="https://doi.org/10.2476/asjaa.69.133" volume="69" year="2020">Suzuki et al. 2020</bibRefCitation>
: figs 7E, 8E, 10E, 11E). Tegular surface beneath conductor is generally sclerotized with many folds (referred as 'ventral side of tegulum beneath posterior edge of embolic
<normalizedToken originalValue="division’">division'</normalizedToken>
in
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.69.133" author="Suzuki, Y" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="67" pagination="133 - 150" refId="B28" refString="Suzuki, Y, Serita, R, Hiramatsu, T, 2020. Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species. Acta Arachnologica 69 (2): 133 - 150, DOI: https://doi.org/10.2476/asjaa.69.133" title="Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species." url="https://doi.org/10.2476/asjaa.69.133" volume="69" year="2020">Suzuki et al. 2020</bibRefCitation>
), of which shape and surface texture vary among species (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.255.3272" author="Zhao, QY" journalOrPublisher="ZooKeys" pageId="0" pageNumber="67" pagination="1 - 48" refId="B35" refString="Zhao, QY, Li, SQ, 2012. Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae). ZooKeys 255: 1 - 48, DOI: https://doi.org/10.3897/zookeys.255.3272" title="Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae)." url="https://doi.org/10.3897/zookeys.255.3272" volume="255" year="2012">Zhao and Li 2012</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.69.133" author="Suzuki, Y" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="67" pagination="133 - 150" refId="B28" refString="Suzuki, Y, Serita, R, Hiramatsu, T, 2020. Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species. Acta Arachnologica 69 (2): 133 - 150, DOI: https://doi.org/10.2476/asjaa.69.133" title="Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species." url="https://doi.org/10.2476/asjaa.69.133" volume="69" year="2020">Suzuki et al. 2020</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="67">
Females of the genus can be distinguished from related genera (
<taxonomicName authorityName="Coddington" authorityYear="1986" class="Arachnida" family="Theridiosomatidae" genus="Baalzebub" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baalzebub" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="67">Baalzebub</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Coddington" authorityYear="1986" class="Arachnida" family="Theridiosomatidae" genus="Epilineutes" kingdom="Animalia" lsidName="Epilineutes" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="67">Epilineutes</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Suzuki &amp; Hiramatsu &amp; Tatsuta" authorityYear="2022" baseAuthorityName="Suzuki &amp; Hiramatsu &amp; Tatsuta" baseAuthorityYear="2022" class="Arachnida" family="Theridiosomatidae" genus="Wendilgarda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wendilgarda" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="67">Wendilgarda</emphasis>
</taxonomicName>
) by having relatively sclerotized, robust copulatory ducts running from the bursa to the spermathecae (
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.422" author="Coddington, JA" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="67" pagination="1 - 96" refId="B4" refString="Coddington, JA, 1986a. The genera of the spider family Theridiosomatidae. Smithsonian Contributions to Zoology 422 (422): 1 - 96, DOI: https://doi.org/10.5479/si.00810282.422" title="The genera of the spider family Theridiosomatidae." url="https://doi.org/10.5479/si.00810282.422" volume="422" year="1986 a">Coddington 1986a</bibRefCitation>
: figs 145, 152). Surface of epigynal plate is smooth and its posterior margin generally lacks scape-like structures. Shape of posterior margin of epigynal plate varies among
<taxonomicName authorityName="O. Pickard-Cambridge" authorityYear="1879" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theridiosoma" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="67">Theridiosoma</emphasis>
</taxonomicName>
species: rounded or almost straight in some species, while having a pair of small, sclerotized processes (named as
<normalizedToken originalValue="‘spurs’">'spurs'</normalizedToken>
in
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.422" author="Coddington, JA" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="67" pagination="1 - 96" refId="B4" refString="Coddington, JA, 1986a. The genera of the spider family Theridiosomatidae. Smithsonian Contributions to Zoology 422 (422): 1 - 96, DOI: https://doi.org/10.5479/si.00810282.422" title="The genera of the spider family Theridiosomatidae." url="https://doi.org/10.5479/si.00810282.422" volume="422" year="1986 a">Coddington 1986a</bibRefCitation>
) or a small slit-like invagination in others (
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.422" author="Coddington, JA" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="67" pagination="1 - 96" refId="B4" refString="Coddington, JA, 1986a. The genera of the spider family Theridiosomatidae. Smithsonian Contributions to Zoology 422 (422): 1 - 96, DOI: https://doi.org/10.5479/si.00810282.422" title="The genera of the spider family Theridiosomatidae." url="https://doi.org/10.5479/si.00810282.422" volume="422" year="1986 a">Coddington 1986a</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="67" pagination="9 - 195" refId="B13" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160" volume="11" year="2009">Miller et al. 2009</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.69.133" author="Suzuki, Y" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="67" pagination="133 - 150" refId="B28" refString="Suzuki, Y, Serita, R, Hiramatsu, T, 2020. Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species. Acta Arachnologica 69 (2): 133 - 150, DOI: https://doi.org/10.2476/asjaa.69.133" title="Japanese spiders of the genus Theridiosoma (Araneae: Theridiosomatidae) with the description of four new species." url="https://doi.org/10.2476/asjaa.69.133" volume="69" year="2020">Suzuki et al. 2020</bibRefCitation>
).
<taxonomicName authorityName="Saaristo" authorityYear="1996" class="Arachnida" family="Theridiosomatidae" genus="Zoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoma" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="67">Zoma</emphasis>
</taxonomicName>
females possess a similar genitalia except in having a sclerotized median pit on the surface of the epigynal plate and lacking any processes nor invaginations on the posterior margin of the epigynal plate in known species (
<bibRefCitation author="Saaristo, MI" journalOrPublisher="Phelsuma" pageId="0" pageNumber="67" pagination="48 - 52" refId="B21" refString="Saaristo, MI, 1996. Theridiosomatid spiders of the granitic islands of Seychelles (Araneae, Theridiosomatidae). Phelsuma 4: 48 - 52" title="Theridiosomatid spiders of the granitic islands of Seychelles (Araneae, Theridiosomatidae)." volume="4" year="1996">Saaristo 1996</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="67" pagination="9 - 195" refId="B13" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160" volume="11" year="2009">Miller et al. 2009</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.255.3272" author="Zhao, QY" journalOrPublisher="ZooKeys" pageId="0" pageNumber="67" pagination="1 - 48" refId="B35" refString="Zhao, QY, Li, SQ, 2012. Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae). ZooKeys 255: 1 - 48, DOI: https://doi.org/10.3897/zookeys.255.3272" title="Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae)." url="https://doi.org/10.3897/zookeys.255.3272" volume="255" year="2012">Zhao and Li 2012</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4927.2.2" author="Ballarin, F" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="197 - 208" refId="B2" refString="Ballarin, F, Yamasaki, T, Su, YC, 2021. A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae). Zootaxa 4927 (2): 197 - 208, DOI: https://doi.org/10.11646/zootaxa.4927.2.2" title="A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae)." url="https://doi.org/10.11646/zootaxa.4927.2.2" volume="4927" year="2021">Ballarin et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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