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<document id="026D69498EDC84CBE5978799C40099C2" ID-DOI="10.5852/ejt.2023.908.2339" ID-ISSN="2118-9773" ID-Zenodo-Dep="10158922" ID-ZooBank="0F6C37D2-DD6A-49E8-9447-5666859BE75E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700470817120" checkinUser="plazi" docAuthor="Akbar, Shahid Ali, Bharti, Himender, Schifani, Enrico &amp; Wachkoo, Aijaz Ahmad" docDate="2023" docId="8B339A0CB3529D0730DCBA14A7A54799" docLanguage="en" docName="EJT.2023.908.77-107.pdf" docOrigin="European Journal of Taxonomy 908" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2339/10187" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.33:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="33" docTitle="Vollenhovia keralensis Kripakaran &amp; Sadasivan 2022" docType="treatment" docVersion="4" lastPageNumber="87" masterDocId="770AE274B3559D0D3105BE4AA54A462F" masterDocTitle="Overview of the ant genus Vollenhovia (Hymenoptera, Formicidae) in India and Sri Lanka, with an illustrated key and the description of a new species" masterLastPageNumber="107" masterPageNumber="77" pageNumber="84" updateTime="1704722585636" updateUser="felipe" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8898744FD09D5F350A3F2EC8DE59481A">Overview of the ant genus Vollenhovia (Hymenoptera, Formicidae) in India and Sri Lanka, with an illustrated key and the description of a new species</mods:title>
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<mods:name id="F4DA03D1196E281231E9A85F046418B2" type="personal">
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<mods:namePart id="BCA488A52FAC06A2B870642654E3FD1F">Akbar, Shahid Ali</mods:namePart>
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<mods:affiliation id="E37962214FA7C136C0F625A1D88F33B4">Central Institute of Temperate Horticulture, Srinagar, Jammu and Kashmir - 191132, India.</mods:affiliation>
<mods:nameIdentifier id="5D0B42B6C03B19A69C38D07C84FA7F52" type="email">kingakbarali@gmail.com</mods:nameIdentifier>
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<mods:name id="1876A34515B7C4B7A2E66A37292FB516" type="personal">
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<mods:namePart id="B5DF05CD62AE00ABA5DD8137FD84BBCF">Bharti, Himender</mods:namePart>
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<mods:affiliation id="C4C17514EFF9FA0B06051726B9777D4D">Department of Zoology and Environmental Sciences, Punjabi University Patiala, Punjab - 147002, India.</mods:affiliation>
<mods:nameIdentifier id="FFF9CC14771A476E3578C753EC4B7874" type="email">himenderbharti@gmail.com</mods:nameIdentifier>
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<mods:name id="1B719BFFC48B78EE2DFBAA9552833CE3" type="personal">
<mods:role id="C319EBF8A652E935E51C2F4EC83DE29D">
<mods:roleTerm id="5958236D2667375CECAB0EACF3AFE0EE">Author</mods:roleTerm>
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<mods:namePart id="0EF4E41DCD3916225498E31A5A36F240">Schifani, Enrico</mods:namePart>
<mods:nameIdentifier id="4A9A405E400507571221966C1A3412F5" type="LSID">18D1CCD1-4A50-452E-8CD8-225596E5304B</mods:nameIdentifier>
<mods:affiliation id="E469507BB496597023BCAF5C7B8A16E7">Department of Chemistry, Life Sciences and Environmental Sustainability, University of Parma - 43124 Parma, Italy.</mods:affiliation>
<mods:nameIdentifier id="3206DD5E8650C285656E2D1E3EA36A60" type="email">enrico.schifani@unipr.it</mods:nameIdentifier>
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<mods:namePart id="2323666C363AC3F391B13FCA8C0E0821">Wachkoo, Aijaz Ahmad</mods:namePart>
<mods:nameIdentifier id="01784AD03C5301F5A570EE1D9CA9831D" type="LSID">6F19EB1F-5DDC-4722-BBD3-F75C29F901D9</mods:nameIdentifier>
<mods:affiliation id="90983D24BA8E81F4E7F67A4A9720CDE0">Department of Zoology, Imtiyaz Memorial Government Degree College, Shopian, Jammu and Kashmir - 192303, India.</mods:affiliation>
<mods:nameIdentifier id="69BACD8DDD4B17B4E66B69AACF8AEBD0" type="email">aijaz_shoorida@yahoo.co.in</mods:nameIdentifier>
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<mods:title id="559718BE618EE0C10871C62052FCFA83">European Journal of Taxonomy</mods:title>
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<mods:date id="0E986C03D73E4945AFAF753D01BBC6E9">2023</mods:date>
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<mods:identifier id="189C72E6DCA140D7209DAF007DA3831F" type="DOI">10.5852/ejt.2023.908.2339</mods:identifier>
<mods:identifier id="1A3CA3459940BE4A384D509E00BD8BD7" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="7B57A8A373216B5350CB1909C1EF3F5D" type="Zenodo-Dep">10158922</mods:identifier>
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<treatment id="8B339A0CB3529D0730DCBA14A7A54799" LSID="urn:lsid:plazi:treatment:8B339A0CB3529D0730DCBA14A7A54799" httpUri="http://treatment.plazi.org/id/8B339A0CB3529D0730DCBA14A7A54799" lastPageId="10" lastPageNumber="87" pageId="7" pageNumber="84">
<subSubSection id="4B807891B3529D0A30DCBA14A1114256" box="[473,1115,1118,1145]" pageId="7" pageNumber="84" type="nomenclature">
<paragraph id="03252B1AB3529D0A30DCBA14A1114256" blockId="7.[473,1115,1118,1181]" box="[473,1115,1118,1145]" pageId="7" pageNumber="84">
<heading id="586D9C76B3529D0A30DCBA14A1114256" box="[473,1115,1118,1145]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="84" reason="2">
<taxonomicName id="C49A5099B3529D0A30DCBA14A1114256" authority="Kripakaran &amp; Sadasivan, 2022" authorityName="Kripakaran &amp; Sadasivan" authorityYear="2022" box="[473,1115,1118,1145]" class="Insecta" family="Formicidae" genus="Vollenhovia" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="keralensis">
<emphasis id="31EEF708B3529D0A30DCBA14A7A34256" bold="true" box="[473,745,1118,1145]" italics="true" pageId="7" pageNumber="84">Vollenhovia keralensis</emphasis>
Kripakaran &amp;
<bibRefCitation id="670B56EBB3529D0A329FBA15A1114256" author="Sadasivan" box="[922,1115,1119,1145]" firstAuthor="Sadasivan" pageId="7" pageNumber="84" pagination="21368 - 21387" refId="ref11849" refString="Sadasivan K. &amp; Kripakaran M. 2022. First record of Proceratium Roger, 1863, Zasphinctus Wheeler, 1918, and Vollenhovia Mayr, 1865 (Hymenoptera: Formicidae) from the Western Ghats of peninsular India, description of three new species, and implications for Indian biogeography. Journal of Threatened Taxa 14 (7): 21368 - 21387. https: // doi. org / 10.11609 / jott. 7682.14.7.21368 - 21387" type="journal article" year="2022">Sadasivan, 2022</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4B807891B3529D0A33F2BAC8A67642B2" box="[759,828,1154,1181]" pageId="7" pageNumber="84" type="description">
<paragraph id="03252B1AB3529D0A33F2BAC8A67642B2" blockId="7.[473,1115,1118,1181]" box="[759,828,1154,1181]" pageId="7" pageNumber="84">
<figureCitation id="9BA1379FB3529D0A33F2BAC8A67642B2" box="[759,828,1154,1181]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[221,1366,265,1884]" captionTargetId="figure-12@9.[221,1366,265,1884]" captionTargetPageId="9" captionText="Fig. 5. Vollenhovia keralensis Kripakaran &amp; Sadasivan, 2022, worker from India (PUAC0026, photographer: Shahid Ali Akbar). A. Head, full face view. B. Habitus, lateral view. C. Habitus, dorsal view. D. Type locality." figureDoi="http://doi.org/10.5281/zenodo.10158936" httpUri="https://zenodo.org/record/10158936/files/figure.png" pageId="7" pageNumber="84">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4B807891B3529D0A31B8BA89A19A42F1" box="[189,1232,1219,1246]" pageId="7" pageNumber="84" type="reference_group">
<paragraph id="03252B1AB3529D0A31B8BA89A19A42F1" blockId="7.[189,1398,1219,1317]" box="[189,1232,1219,1246]" pageId="7" pageNumber="84">
<treatmentCitationGroup id="238A0C34B3529D0A31B8BA89A19A42F1" box="[189,1232,1219,1246]" pageId="7" pageNumber="84">
<taxonomicName id="C49A5099B3529D0A31B8BA89A7AD42F2" authority="Kripakaran &amp; Sadasivan" authorityName="Kripakaran &amp; Sadasivan" authorityYear="2022" box="[189,743,1219,1246]" class="Insecta" family="Formicidae" genus="Vollenhovia" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="keralensis">
<emphasis id="31EEF708B3529D0A31B8BA89A48F42F2" box="[189,453,1219,1245]" italics="true" pageId="7" pageNumber="84">Vollenhovia keralensis</emphasis>
Kripakaran &amp; Sadasivan
</taxonomicName>
in
<treatmentCitation id="823B0D0BB3529D0A320FBA89A18642F1" author="Sadasivan K. &amp; Kripakaran M." box="[778,1228,1219,1246]" page="21381" pageId="7" pageNumber="84" year="2022">
<bibRefCitation id="670B56EBB3529D0A320FBA89A13F42F1" author="Sadasivan &amp; Kripakaran" box="[778,1141,1219,1246]" firstAuthor="Sadasivan" pageId="7" pageNumber="84" pagination="21368 - 21387" refId="ref11849" refString="Sadasivan K. &amp; Kripakaran M. 2022. First record of Proceratium Roger, 1863, Zasphinctus Wheeler, 1918, and Vollenhovia Mayr, 1865 (Hymenoptera: Formicidae) from the Western Ghats of peninsular India, description of three new species, and implications for Indian biogeography. Journal of Threatened Taxa 14 (7): 21368 - 21387. https: // doi. org / 10.11609 / jott. 7682.14.7.21368 - 21387" type="journal article" year="2022">Sadasivan &amp; Kripakaran, 2022</bibRefCitation>
: 21381
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4B807891B3529D0A35D0BA8EA02543F4" pageId="7" pageNumber="84" type="materials_examined">
<paragraph id="03252B1AB3529D0A35D0BA8EA6F4430A" blockId="7.[189,1398,1219,1317]" pageId="7" pageNumber="84">
<materialsCitation id="B3F22147B3529D0A35D0BA8EA6F4430A" collectionCode="TNHS" country="India" location="Peppara Wildlife Sanctuary" municipality="Trivandrum Bonaccord" pageId="7" pageNumber="84" specimenCount="1" specimenCount-worker="1" stateProvince="Kerala" typeStatus="holotype">
Type locality:
<collectingCountry id="7B8D6B8AB3529D0A31ECBAADA462432E" box="[233,296,1255,1281]" name="India" pageId="7" pageNumber="84">India</collectingCountry>
,
<collectingRegion id="C15EE5F8B3529D0A303CBAADA4C0432E" box="[313,394,1255,1281]" country="India" name="Kerala" pageId="7" pageNumber="84">Kerala</collectingRegion>
,
<collectingMunicipality id="E341B160B3529D0A309FBAADA7F9432E" box="[410,691,1255,1281]" pageId="7" pageNumber="84">Trivandrum Bonaccord</collectingMunicipality>
,
<location id="06457DC1B3529D0A33C6BAADA144432E" LSID="urn:lsid:plazi:treatment:8B339A0CB3529D0730DCBA14A7A54799:06457DC1B3529D0A33C6BAADA144432E" box="[707,1038,1255,1281]" country="India" municipality="Trivandrum Bonaccord" name="Peppara Wildlife Sanctuary" pageId="7" pageNumber="84" stateProvince="Kerala">Peppara Wildlife Sanctuary</location>
[
<typeStatus id="DC2195B8B3529D0A3524BAADA1DB432E" box="[1057,1169,1255,1281]" pageId="7" pageNumber="84" type="holotype">Holotype</typeStatus>
<collectionCode id="658BB3DFB3529D0A3598BAADA1A7432E" box="[1181,1261,1255,1281]" collectionName="TNHS" pageId="7" pageNumber="84">TNHS</collectionCode>
; images of holotype
<specimenCount id="159CE093B3529D0A305DBB40A4E1430B" box="[344,427,1290,1316]" pageId="7" pageNumber="84" type="worker">worker</specimenCount>
examined in
<bibRefCitation id="670B56EBB3529D0A334FBB40A6E7430B" author="Sadasivan &amp; Kripakaran" box="[586,941,1290,1316]" firstAuthor="Sadasivan" pageId="7" pageNumber="84" pagination="21368 - 21387" refId="ref11849" refString="Sadasivan K. &amp; Kripakaran M. 2022. First record of Proceratium Roger, 1863, Zasphinctus Wheeler, 1918, and Vollenhovia Mayr, 1865 (Hymenoptera: Formicidae) from the Western Ghats of peninsular India, description of three new species, and implications for Indian biogeography. Journal of Threatened Taxa 14 (7): 21368 - 21387. https: // doi. org / 10.11609 / jott. 7682.14.7.21368 - 21387" type="journal article" year="2022">Sadasivan &amp; Kripakaran 2022</bibRefCitation>
].
</materialsCitation>
</paragraph>
<paragraph id="03252B1AB3529D0A31B8BB01A4E0434A" blockId="7.[189,426,1355,1381]" box="[189,426,1355,1381]" pageId="7" pageNumber="84">
<heading id="586D9C76B3529D0A31B8BB01A4E0434A" bold="true" box="[189,426,1355,1381]" fontSize="11" level="3" pageId="7" pageNumber="84" reason="3">
<emphasis id="31EEF708B3529D0A31B8BB01A4E0434A" bold="true" box="[189,426,1355,1381]" pageId="7" pageNumber="84">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="03252B1AB3529D0A31B8BB30A02543F4" blockId="7.[189,1399,1400,1499]" pageId="7" pageNumber="84">
<materialsCitation id="B3F22147B3529D0A31B8BB30A6694398" collectingDate="2011-10-17" collectingMethod="hand picking" collectionCode="PUAC" collectorName="S. A. Akbar" country="India" elevation="1005" latitude="9.8" location="Periyar Tiger Reserve" longLatPrecision="1299" longitude="77.4" municipality="Periyar Tiger Reserve" pageId="7" pageNumber="84" specimenCode="PUAC0026, PUAC0027, PUAC0028, PUAC0029, PUAC0030, PUAC0031, PUAC0032, PUAC0033" specimenCount="8" specimenCount-worker="8" stateProvince="Kerala">
<collectingCountry id="7B8D6B8AB3529D0A31B8BB30A44443BB" box="[189,270,1402,1428]" name="India" pageId="7" pageNumber="84">INDIA</collectingCountry>
•
<specimenCount id="159CE093B3529D0A302CBB30A4EB43BB" box="[297,417,1402,1428]" count="8" pageId="7" pageNumber="84" type="worker">8 workers</specimenCount>
;
<collectingRegion id="C15EE5F8B3529D0A30ABBB30A4B543BB" box="[430,511,1402,1428]" country="India" name="Kerala" pageId="7" pageNumber="84">Kerala</collectingRegion>
,
<collectingMunicipality id="E341B160B3529D0A3309BB30A65A43BB" box="[524,784,1402,1428]" pageId="7" pageNumber="84">Periyar Tiger Reserve</collectingMunicipality>
;
<geoCoordinate id="66AE4DDDB3529D0A3218BB32A63143BB" box="[797,891,1400,1428]" degrees="9" direction="north" minutes="48" orientation="latitude" pageId="7" pageNumber="84" precision="925" value="9.8">9°48′ N</geoCoordinate>
,
<geoCoordinate id="66AE4DDDB3529D0A3282BB32A6BB43BB" box="[903,1009,1400,1428]" degrees="77" direction="east" minutes="24" orientation="longitude" pageId="7" pageNumber="84" precision="925" value="77.4">77°24′ E</geoCoordinate>
; alt.
<quantity id="C46286FFB3529D0A352EBB30A1C243BB" box="[1067,1160,1402,1429]" metricMagnitude="3" metricUnit="m" metricValue="1.005" pageId="7" pageNumber="84" unit="m" value="1005.0">
<elevation id="88B7CC29B3529D0A352EBB30A1C243BB" box="[1067,1160,1402,1429]" metricMagnitude="3" metricUnit="m" metricValue="1.005" pageId="7" pageNumber="84" unit="m" value="1005.0">1005 m</elevation>
</quantity>
;
<date id="77240DDAB3529D0A3590BB30A07B43BB" box="[1173,1329,1402,1428]" pageId="7" pageNumber="84" value="2011-10-17">
<collectingDate id="6760F432B3529D0A3590BB30A07B43BB" box="[1173,1329,1402,1428]" pageId="7" pageNumber="84" value="2011-10-17">17 Oct. 2011</collectingDate>
</date>
;
<collectingMethod id="DADB530DB3529D0A3438BB30A4534398" pageId="7" pageNumber="84">hand picking</collectingMethod>
;
<collectorName id="AE6F4ECCB3529D0A3020BBD7A4E24398" box="[293,424,1437,1463]" pageId="7" pageNumber="84">S.A. Akbar</collectorName>
leg.;
<specimenCode id="533C8361B3529D0A30EEBBD7A73E4398" box="[491,628,1437,1463]" collectionCode="PUAC" pageId="7" pageNumber="84">PUAC0026</specimenCode>
to
<specimenCode id="533C8361B3529D0A339FBBD7A6694398" box="[666,803,1437,1463]" collectionCode="PUAC" pageId="7" pageNumber="84">PUAC0033</specimenCode>
</materialsCitation>
;
<materialsCitation id="B3F22147B3529D0A3236BBD7A02543F4" collectingDate="2011-10-27" collectingMethod="hand picking" collectionCode="PUAC" collectorName="S. A. Akbar" elevation="1630" latitude="9.583333" location="Manalar" longLatPrecision="1299" longitude="77.3" municipality="Periyar Tiger Reserve" pageId="7" pageNumber="84" specimenCode="PUAC0034, PUAC0035, PUAC0036, PUAC0037" specimenCount="4" specimenCount-worker="4">
<specimenCount id="159CE093B3529D0A3236BBD7A6E14398" box="[819,939,1437,1463]" count="4" pageId="7" pageNumber="84" type="worker">4 workers</specimenCount>
;
<collectingMunicipality id="E341B160B3529D0A32B2BBD7A1F04397" box="[951,1210,1437,1464]" pageId="7" pageNumber="84">Periyar Tiger Reserve</collectingMunicipality>
,
<location id="06457DC1B3529D0A35C2BBD7A0674398" LSID="urn:lsid:plazi:treatment:8B339A0CB3529D0730DCBA14A7A54799:06457DC1B3529D0A35C2BBD7A0674398" box="[1223,1325,1437,1463]" latitude="9.583333" longLatPrecision="1299" longitude="77.3" municipality="Periyar Tiger Reserve" name="Manalar" pageId="7" pageNumber="84">Manalar</location>
;
<geoCoordinate id="66AE4DDDB3529D0A343DBBD6A59F43F4" degrees="9" direction="north" minutes="35" orientation="latitude" pageId="7" pageNumber="84" precision="925" value="9.583333">9°35′ N</geoCoordinate>
,
<geoCoordinate id="66AE4DDDB3529D0A31E4BBF5A40043F4" box="[225,330,1471,1499]" degrees="77" direction="east" minutes="18" orientation="longitude" pageId="7" pageNumber="84" precision="925" value="77.3">77°18′ E</geoCoordinate>
; alt.
<quantity id="C46286FFB3529D0A3087BB8AA49443F4" box="[386,478,1472,1499]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="7" pageNumber="84" unit="m" value="1630.0">
<elevation id="88B7CC29B3529D0A3087BB8AA49443F4" box="[386,478,1472,1499]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="7" pageNumber="84" unit="m" value="1630.0">1630 m</elevation>
</quantity>
;
<date id="77240DDAB3529D0A30EEBB8AA7CF43F5" box="[491,645,1472,1498]" pageId="7" pageNumber="84" value="2011-10-27">
<collectingDate id="6760F432B3529D0A30EEBB8AA7CF43F5" box="[491,645,1472,1498]" pageId="7" pageNumber="84" value="2011-10-27">27 Oct. 2011</collectingDate>
</date>
;
<collectingMethod id="DADB530DB3529D0A3394BB8AA66443F5" box="[657,814,1472,1498]" pageId="7" pageNumber="84">hand picking</collectingMethod>
; leg.
<collectorName id="AE6F4ECCB3529D0A3269BB8AA6A543F5" box="[876,1007,1472,1498]" pageId="7" pageNumber="84">S.A. Akbar</collectorName>
leg.;
<specimenCode id="533C8361B3529D0A3534BB8AA1F043F5" box="[1073,1210,1472,1498]" collectionCode="PUAC" pageId="7" pageNumber="84">PUAC0034</specimenCode>
to
<specimenCode id="533C8361B3529D0A35DABB8AA02343F5" box="[1247,1385,1472,1498]" collectionCode="PUAC" pageId="7" pageNumber="84">PUAC0037</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4B807891B3529D0A31B8B848A1D540D5" pageId="7" pageNumber="84" type="description">
<paragraph id="03252B1AB3529D0A31B8B848A16C4033" blockId="7.[189,1062,1538,1564]" box="[189,1062,1538,1564]" pageId="7" pageNumber="84">
<emphasis id="31EEF708B3529D0A31B8B848A71D4033" bold="true" box="[189,599,1538,1564]" pageId="7" pageNumber="84">Worker measurements &amp; indices</emphasis>
(
<specimenCount id="159CE093B3529D0A336DB848A6404033" box="[616,778,1538,1564]" count="12" pageId="7" pageNumber="84" type="generic">12 specimens</specimenCount>
, 2 colonies, 2 localities)
</paragraph>
<paragraph id="03252B1AB3529D0A31B8B87AA1D540D5" blockId="7.[188,1399,1584,1787]" pageId="7" pageNumber="84">
CL 0.65–0.70 (0.67), CW 0.56–0.58 (0.57),
<collectionCode id="658BB3DFB3529D0A33CEB87AA7A54065" box="[715,751,1584,1610]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
0.60–0.63 (0.62), EL 0.13–0.15 (0.14),
<collectionCode id="658BB3DFB3529D0A35C8B87BA1B24064" box="[1229,1272,1585,1611]" country="France" name="Musee de Lectoure" pageId="7" pageNumber="84">ML</collectionCode>
0.80–0.92 (0.85), PEH 0.35–0.37 (0.36), PEL 0.22–0.26 (0.24),
<collectionCode id="658BB3DFB3529D0A322DB81EA62F4041" box="[808,869,1620,1646]" pageId="7" pageNumber="84">PEW</collectionCode>
0.23–0.24 (0.24), PPH 0.21–0.22 (0.21), PPL 0.23–0.27 (0.24), PPW 0.24–0.25 (0.24),
<collectionCode id="658BB3DFB3529D0A33AFB83DA79C40BE" box="[682,726,1655,1681]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="84">PW</collectionCode>
0.42–0.51 (0.47), SL 0.38–0.40 (0.39) mm. Indices: CL/ CW 1.12–1.25 (1.17), EL/
<collectionCode id="658BB3DFB3529D0A30F9B8D0A754409B" box="[508,542,1690,1716]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
0.21–0.24 (0.22),
<collectionCode id="658BB3DFB3529D0A3205B8D0A667409B" box="[768,813,1690,1716]" country="France" name="Musee de Lectoure" pageId="7" pageNumber="84">ML</collectionCode>
/
<collectionCode id="658BB3DFB3529D0A3233B8D0A612409B" box="[822,856,1690,1716]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
1.30–1.47 (1.38), PEH/
<collectionCode id="658BB3DFB3529D0A3578B8D0A1D5409B" box="[1149,1183,1690,1716]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
0.56–0.60 (0.58), PEL/
<collectionCode id="658BB3DFB3529D0A31FEB8F7A45740F8" box="[251,285,1725,1751]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
0.36–0.43 (0.39),
<collectionCode id="658BB3DFB3529D0A3307B8F7A70840F8" box="[514,578,1725,1751]" pageId="7" pageNumber="84">PEW</collectionCode>
/
<collectionCode id="658BB3DFB3529D0A334FB8F7A72640F8" box="[586,620,1725,1751]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
0.37–0.40 (0.38), PPH/
<collectionCode id="658BB3DFB3529D0A3294B8F7A6F940F8" box="[913,947,1725,1751]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
0.34–0.37 (0.35), PPL/
<collectionCode id="658BB3DFB3529D0A35D1B8F7A1BF40F8" box="[1236,1269,1725,1751]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
0.37–0.44 (0.39), PPW/
<collectionCode id="658BB3DFB3529D0A305CB8AAA43040D5" box="[345,378,1760,1786]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
0.38–0.42 (0.39),
<collectionCode id="658BB3DFB3529D0A3351B8AAA7CB40D5" box="[596,641,1760,1786]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="84">PW</collectionCode>
/
<collectionCode id="658BB3DFB3529D0A338EB8AAA7E740D5" box="[651,685,1760,1786]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
0.68–0.82 (0.76), SL/
<collectionCode id="658BB3DFB3529D0A32B4B8AAA69E40D5" box="[945,980,1760,1786]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="7" pageNumber="84">CS</collectionCode>
0.60–065 (0.63).
</paragraph>
</subSubSection>
<subSubSection id="4B807891B3529D0731B8B968A7384768" lastPageId="10" lastPageNumber="87" pageId="7" pageNumber="84" type="discussion">
<paragraph id="03252B1AB3529D0A31B8B968A4654113" blockId="7.[189,303,1826,1852]" box="[189,303,1826,1852]" pageId="7" pageNumber="84">
<heading id="586D9C76B3529D0A31B8B968A4654113" bold="true" box="[189,303,1826,1852]" fontSize="11" level="3" pageId="7" pageNumber="84" reason="3">
<emphasis id="31EEF708B3529D0A31B8B968A4654113" bold="true" box="[189,303,1826,1852]" pageId="7" pageNumber="84">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="03252B1AB3529D0731B8B91AA7384768" blockId="7.[189,1400,1872,2004]" lastBlockId="10.[189,1399,265,327]" lastPageId="10" lastPageNumber="87" pageId="7" pageNumber="84">
This species from the Western Ghats closely resembles
<taxonomicName id="C49A5099B3529D0A3254B91BA15E4145" authorityName="Dhadwal, Rilta &amp; Bharti" authorityYear="2023" box="[849,1044,1872,1898]" class="Insecta" family="Formicidae" genus="Vollenhovia" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="karimalaensis">
<emphasis id="31EEF708B3529D0A3254B91BA15E4145" box="[849,1044,1872,1898]" italics="true" pageId="7" pageNumber="84">V. karimalaensis</emphasis>
</taxonomicName>
from the same region but can be separated based on the number of mandible teeth, the sculpture, and the petiole shape (see Remarks under
<taxonomicName id="C49A5099B3529D0A300CB9DDA487419E" authorityName="Dhadwal, Rilta &amp; Bharti" authorityYear="2023" box="[265,461,1943,1969]" class="Insecta" family="Formicidae" genus="Vollenhovia" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="karimalaensis">
<emphasis id="31EEF708B3529D0A300CB9DDA487419E" box="[265,461,1943,1969]" italics="true" pageId="7" pageNumber="84">V. karimalaensis</emphasis>
</taxonomicName>
). It is one of the few species of the genus of which all three castes have been described (
<bibRefCitation id="670B56EBB3529D0A3045B9F0A7E041FB" author="Sadasivan &amp; Kripakaran" box="[320,682,1978,2004]" firstAuthor="Sadasivan" pageId="7" pageNumber="84" pagination="21368 - 21387" refId="ref11849" refString="Sadasivan K. &amp; Kripakaran M. 2022. First record of Proceratium Roger, 1863, Zasphinctus Wheeler, 1918, and Vollenhovia Mayr, 1865 (Hymenoptera: Formicidae) from the Western Ghats of peninsular India, description of three new species, and implications for Indian biogeography. Journal of Threatened Taxa 14 (7): 21368 - 21387. https: // doi. org / 10.11609 / jott. 7682.14.7.21368 - 21387" type="journal article" year="2022">Sadasivan &amp; Kripakaran 2022</bibRefCitation>
). We were able to collect specimens from two new localities in the Periyar Tiger Reserve under the bark of a log; the general vegetation and overview of one of the collecting sites are shown in
<figureCitation id="9BA1379FB35F9D073314BF66A7244768" box="[529,622,300,327]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[221,1366,265,1884]" captionTargetId="figure-12@9.[221,1366,265,1884]" captionTargetPageId="9" captionText="Fig. 5. Vollenhovia keralensis Kripakaran &amp; Sadasivan, 2022, worker from India (PUAC0026, photographer: Shahid Ali Akbar). A. Head, full face view. B. Habitus, lateral view. C. Habitus, dorsal view. D. Type locality." figureDoi="http://doi.org/10.5281/zenodo.10158936" httpUri="https://zenodo.org/record/10158936/files/figure.png" pageId="10" pageNumber="87">Fig. 5D</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="57E57B92B35D9D0531B8B9CFA75F41C9" ID-DOI="http://doi.org/10.5281/zenodo.10158934" ID-Zenodo-Dep="10158934" httpUri="https://zenodo.org/record/10158934/files/figure.png" pageId="8" pageNumber="85" startId="8.[189,232,1925,1951]" targetBox="[221,1367,265,1884]" targetPageId="8">
<paragraph id="03252B1AB35D9D0531B8B9CFA75F41C9" blockId="8.[189,1398,1925,2023]" pageId="8" pageNumber="85">
<emphasis id="31EEF708B35D9D0531B8B9CFA44641B0" bold="true" box="[189,268,1925,1951]" pageId="8" pageNumber="85">Fig. 4.</emphasis>
<taxonomicName id="C49A5099B35D9D053010B9CCA688418F" authority="Dhadwal, Rilta &amp; Bharti, 2023" authorityName="Dhadwal, Rilta &amp; Bharti" authorityYear="2023" box="[277,962,1926,1952]" class="Insecta" family="Formicidae" genus="Vollenhovia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="karimalaensis">
<emphasis id="31EEF708B35D9D053010B9CCA707418F" box="[277,589,1926,1952]" italics="true" pageId="8" pageNumber="85">Vollenhovia karimalaensis</emphasis>
<bibRefCitation id="670B56EBB35D9D053351B9CCA688418F" author="Dhadwal T. &amp; Rilta J. S. &amp; Bharti H." box="[596,962,1926,1952]" journalOrPublisher="Sociobiology" pageId="8" pageNumber="85" pagination="9103" part="70" refId="ref10102" refString="Dhadwal T., Rilta J. S. &amp; Bharti H. 2023. Taxonomic studies on the genus Vollenhovia Mayr, 1865 (Hymenoptera: Formicidae): five new species from India. Sociobiology 70: e 9103. https: // doi. org / 10.13102 / sociobiology. v 70 i 3.9103" title="Taxonomic studies on the genus Vollenhovia Mayr, 1865 (Hymenoptera: Formicidae): five new species from India" type="journal article" year="2023">Dhadwal, Rilta &amp; Bharti, 2023</bibRefCitation>
</taxonomicName>
, holotype worker from India (PUAC T701, photographer: Himender Bharti).
<emphasis id="31EEF708B35D9D05339BB9E3A7FB41EC" bold="true" box="[670,689,1961,1987]" pageId="8" pageNumber="85">A</emphasis>
. Head, full face view.
<emphasis id="31EEF708B35D9D0532C3B9E3A69241EC" bold="true" box="[966,984,1961,1987]" pageId="8" pageNumber="85">B</emphasis>
. Habitus, lateral view.
<emphasis id="31EEF708B35D9D0535F5B9E3A04941EC" bold="true" box="[1264,1283,1961,1987]" pageId="8" pageNumber="85">C</emphasis>
. Habitus, dorsal view.
<emphasis id="31EEF708B35D9D053057B986A42D41C9" bold="true" box="[338,359,1996,2022]" pageId="8" pageNumber="85">D</emphasis>
. Type locality.
</paragraph>
</caption>
<caption id="57E57B92B35C9D0431B8B9C2A48C41C7" ID-DOI="http://doi.org/10.5281/zenodo.10158936" ID-Zenodo-Dep="10158936" httpUri="https://zenodo.org/record/10158936/files/figure.png" pageId="9" pageNumber="86" startId="9.[189,232,1928,1954]" targetBox="[221,1366,265,1884]" targetPageId="9">
<paragraph id="03252B1AB35C9D0431B8B9C2A48C41C7" blockId="9.[189,1399,1928,2025]" pageId="9" pageNumber="86">
<emphasis id="31EEF708B35C9D0431B8B9C2A45E418D" bold="true" box="[189,276,1928,1954]" pageId="9" pageNumber="86">Fig. 5.</emphasis>
<taxonomicName id="C49A5099B35C9D043020B9C2A685418D" authority="Kripakaran &amp; Sadasivan, 2022" authorityName="Kripakaran &amp; Sadasivan" authorityYear="2022" box="[293,975,1928,1954]" class="Insecta" family="Formicidae" genus="Vollenhovia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="keralensis">
<emphasis id="31EEF708B35C9D043020B9C2A77D418D" box="[293,567,1928,1954]" italics="true" pageId="9" pageNumber="86">Vollenhovia keralensis</emphasis>
Kripakaran &amp;
<bibRefCitation id="670B56EBB35C9D043207B9C2A685418D" author="Sadasivan" box="[770,975,1928,1954]" firstAuthor="Sadasivan" pageId="9" pageNumber="86" pagination="21368 - 21387" refId="ref11849" refString="Sadasivan K. &amp; Kripakaran M. 2022. First record of Proceratium Roger, 1863, Zasphinctus Wheeler, 1918, and Vollenhovia Mayr, 1865 (Hymenoptera: Formicidae) from the Western Ghats of peninsular India, description of three new species, and implications for Indian biogeography. Journal of Threatened Taxa 14 (7): 21368 - 21387. https: // doi. org / 10.11609 / jott. 7682.14.7.21368 - 21387" type="journal article" year="2022">Sadasivan, 2022</bibRefCitation>
</taxonomicName>
, worker from India (PUAC0026, photographer: Shahid Ali Akbar).
<emphasis id="31EEF708B35C9D043357B9E1A72D41EA" bold="true" box="[594,615,1963,1989]" pageId="9" pageNumber="86">A</emphasis>
. Head, full face view.
<emphasis id="31EEF708B35C9D04327DB9E1A6C041EA" bold="true" box="[888,906,1963,1989]" pageId="9" pageNumber="86">B</emphasis>
. Habitus, lateral view.
<emphasis id="31EEF708B35C9D0435A5B9E1A1FE41EA" bold="true" box="[1184,1204,1963,1989]" pageId="9" pageNumber="86">C</emphasis>
. Habitus, dorsal view.
<emphasis id="31EEF708B35C9D043006B984A45241C7" bold="true" box="[259,280,1998,2024]" pageId="9" pageNumber="86">D</emphasis>
. Type locality.
</paragraph>
</caption>
<subSubSection id="4B807891B35F9D0731B8BF27A7A54799" box="[189,751,365,438]" pageId="10" pageNumber="87" type="distribution">
<paragraph id="03252B1AB35F9D0731B8BF27A41C47A8" blockId="10.[189,342,365,391]" box="[189,342,365,391]" pageId="10" pageNumber="87">
<heading id="586D9C76B35F9D0731B8BF27A41C47A8" bold="true" box="[189,342,365,391]" fontSize="11" level="3" pageId="10" pageNumber="87" reason="3">
<emphasis id="31EEF708B35F9D0731B8BF27A41C47A8" bold="true" box="[189,342,365,391]" pageId="10" pageNumber="87">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="03252B1AB35F9D0731B8BFD6A7A54799" blockId="10.[189,751,412,438]" box="[189,751,412,438]" pageId="10" pageNumber="87">
<collectingCountry id="7B8D6B8AB35F9D0731B8BFD6A5B34799" box="[189,249,412,438]" name="India" pageId="10" pageNumber="87">India</collectingCountry>
(
<collectingRegion id="C15EE5F8B35F9D07300FBFD6A4124799" box="[266,344,412,438]" country="India" name="Kerala" pageId="10" pageNumber="87">Kerala</collectingRegion>
) (
<bibRefCitation id="670B56EBB35F9D073077BFD6A7A84799" author="Sadasivan &amp; Kripakaran" box="[370,738,412,438]" firstAuthor="Sadasivan" pageId="10" pageNumber="87" pagination="21368 - 21387" refId="ref11849" refString="Sadasivan K. &amp; Kripakaran M. 2022. First record of Proceratium Roger, 1863, Zasphinctus Wheeler, 1918, and Vollenhovia Mayr, 1865 (Hymenoptera: Formicidae) from the Western Ghats of peninsular India, description of three new species, and implications for Indian biogeography. Journal of Threatened Taxa 14 (7): 21368 - 21387. https: // doi. org / 10.11609 / jott. 7682.14.7.21368 - 21387" type="journal article" year="2022">Sadasivan &amp; Kripakaran 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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